319 lines
39 KiB
XML
319 lines
39 KiB
XML
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<mods:title id="D9D274E2FF71E9E14CCD85E9589A8B84">First record of the genus Passiena (Araneae, Lycosidae) from China, with the first description of the male of P. spinicrus Thorell, 1890 from Malaysia</mods:title>
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<mods:namePart id="5D871B5399B69DA5212AD28E7229D6E6">Tan, Bing</mods:namePart>
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<mods:affiliation id="BE48CD3228F6BEE08B876532B6DAB4E1">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Science, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="CCB047A99B230D74ADD31DB538FE7BAA">Irfan, Muhammad</mods:namePart>
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<mods:nameIdentifier id="7EE989E28566CF5AE532FB9ED0D11196" type="ORCID">https://orcid.org/0000-0003-0445-9612</mods:nameIdentifier>
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<mods:affiliation id="97F12554F68EE08B946C5434EB395308">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Science, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="263E6C21137A444DD4FF92B33B2EDBB0">Zhang, Zhi-Sheng</mods:namePart>
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<mods:nameIdentifier id="55D24B0079743850D335D86691375611" type="ORCID">https://orcid.org/0000-0002-9304-1789</mods:nameIdentifier>
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<mods:affiliation id="0286251931CC2CCB0EBB194E33476860">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Science, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="2D34CE3100DA4E6D14BED9D72DA86045">Wang, Lu-Yu</mods:namePart>
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<mods:affiliation id="A5F4D94EB109941D14935B3D676B4C14">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Science, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:nameIdentifier id="96041E73F5705E4535E924331A45D64F" type="email">wangluyu1989@163.com</mods:nameIdentifier>
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<mods:title id="7F7BE537ECF7DB4AE4D4062591E0EAEE">ZooKeys</mods:title>
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<mods:date id="85EE27867ADB97A0819B70366976E3B8">2023</mods:date>
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<mods:url id="B6E1A3C5EFFF4BEBA41FA6A8D49E484B">http://dx.doi.org/10.3897/zookeys.1182.109532</mods:url>
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<mods:classification id="F739527C0BA765C713C478FE6F68B114">journal article</mods:classification>
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<mods:identifier id="CC3594A7E10E147F6B7A26C58A66EBF2" type="DOI">http://dx.doi.org/10.3897/zookeys.1182.109532</mods:identifier>
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<treatment id="44FD99A2F8E650F9A741F6A7BB7A6D98" LSID="urn:lsid:zoobank.org:act:6CE69B00-418B-4697-B35E-C2347209460E" httpUri="http://treatment.plazi.org/id/44FD99A2F8E650F9A741F6A7BB7A6D98" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="C666BF2016B80D17C5ABE804A3D310DB" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="021345380AF19C7A48C8AA8288810F2A" pageId="0" pageNumber="1">
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<taxonomicName id="4B68D8E5369833DCBF2D7E2678A36DB8" LSID="https://zoobank.org/6CE69B00-418B-4697-B35E-C2347209460E" authority="sp. nov. (段氏帕狼蛛)" authorityName="sp. nov. (段氏帕狼蛛)" class="Arachnida" family="Lycosidae" genus="Passiena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passiena duani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="duani">Passiena duani sp. nov. (段氏帕狼蛛)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="84D3F8D5877AC0BDECD43639A8036435" pageId="0" pageNumber="1" type="description">
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<paragraph id="223EC63C6DA0248AFF9058068D8FBCCD" pageId="0" pageNumber="1">
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<figureCitation id="4C505BDA19A65512E72BD1841D758CB4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live photo of Passiena duani sp. nov. (A, B) and P. spinicrus Thorell, 1890 (C, D) A, C male B, D female. Live photos taken by Qian-Le Lu (A, B) and Lu-Yu Wang (C, D)." figureDoi="10.3897/zookeys.1182.109532.figure1" httpUri="https://binary.pensoft.net/fig/919164" pageId="0" pageNumber="1">Figs 1A, B</figureCitation>
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<figureCitation id="0FEFB5942C1523DC40D88FF31470E454" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">, 2A-D</figureCitation>
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<figureCitation id="468FDA9F65E59182D7F07D981EA14995" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">, 3A-I</figureCitation>
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<figureCitation id="174EEE7062CF5CFECA51A6564D7843A3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map showing distribution records of Passiena duani sp. nov. and P. spinicrus Thorell, 1890." figureDoi="10.3897/zookeys.1182.109532.figure5" httpUri="https://binary.pensoft.net/fig/919168" pageId="0" pageNumber="1">, 5</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="040C1948077E2DA089764D863F7DF251" pageId="0" pageNumber="1" type="type material">
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<paragraph id="3951F041D5A99A441E131BF53C285997" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="C89C89F44E69D6B7F60333C08B8EFF75" pageId="0" pageNumber="1">
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<emphasis id="F750566AF8A74A3F9072B9EDF22E0D60" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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(male): China, Guangxi Zhuang Autonomous Region, Chongzuo City, Ningming County, Chengzhong Town, Panlong,
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<geoCoordinate id="FDA674912A8D56F73D43E6FF21A29627" degrees="22.2347" direction="north" orientation="latitude" precision="5" value="22.2347">22.2347°N</geoCoordinate>
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,
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<geoCoordinate id="E731A2DF0716024AC850E81592A23326" degrees="107.0538" direction="east" orientation="longitude" precision="5" value="107.0538">107.0538°E</geoCoordinate>
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, elev. 138 m, 25 April 2023, L.Y. Wang and Q.L. Lu leg. (SWUC-T-LY-13-01);
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<emphasis id="4F193D11234964D1C97C077EF693D86F" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(3 males and 4 females): 2 males and 3 females, same data as holotype; 1 male and 1 female (SWUC-T-LY-13-07~08), Ningming County, Tuolong Township, Nongna Village,
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<geoCoordinate id="3F38C5CC08558F333D9D7321124A50D4" degrees="22.2325" direction="north" orientation="latitude" precision="5" value="22.2325">22.2325°N</geoCoordinate>
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,
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<geoCoordinate id="9EF60D23A600FB25E143B7F73423228E" degrees="107.0558" direction="east" orientation="longitude" precision="5" value="107.0558">107.0558°E</geoCoordinate>
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, elev. 152 m, 19 June 2017, L.Y. Wang and R.B. Wu leg. (SWUC-T-LY-13-02~06).
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</paragraph>
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</subSubSection>
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<subSubSection id="388C19F0F0F23437B932014FD82D1BB0" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="65FB50EDE95605DB5205414449749922" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="D0F19540D99D62BBA07F203C8FD7B3D0" pageId="0" pageNumber="1">The specific name comes from the family name of Dr Meichun Duan, who gave much support to our research on spiders; noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection id="DE35CE40A743084FE45DBE9DDF1DAAFC" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="8D46183B06312CA36A3825B609DB6BA7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="5284F1E52B6003DFFF9303EF0FFB5F10" pageId="0" pageNumber="1">
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The new species resembles
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<taxonomicName id="9ADF1360B6A6E18F91411E8648F85A27" lsidName="P. bayi" pageId="0" pageNumber="1" rank="species" species="bayi">
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<emphasis id="BA3638E13E7D7AAC04D4D009A3E529D8" italics="true" pageId="0" pageNumber="1">P. bayi</emphasis>
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</taxonomicName>
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Omelko & Marusik, 2020,
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<taxonomicName id="5B9272BFF685626800AC5F3DCD91504D" lsidName="P. torbjoerni" pageId="0" pageNumber="1" rank="species" species="torbjoerni">
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<emphasis id="D0441B511476848E11D2DA9F0DBFF8A1" italics="true" pageId="0" pageNumber="1">P. torbjoerni</emphasis>
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</taxonomicName>
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Lehtinen, 2005 (Figs
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<figureCitation id="DAB40018ACF8DE9FC81E2096464DBE60" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">2A-D</figureCitation>
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,
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<figureCitation id="FDA0F950A69295526B44AEB8D618E05F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3C-I</figureCitation>
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;
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<bibRefCitation id="920015A1B55FA598E7F975207DBE1011" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
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, figs 19-29) and
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<taxonomicName id="E5B4514DAB89508643B46C470D9AF403" lsidName="P. spinicrus" pageId="0" pageNumber="1" rank="species" species="spinicrus">
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<emphasis id="CAF4449423174108058AF88F782A8608" italics="true" pageId="0" pageNumber="1">P. spinicrus</emphasis>
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</taxonomicName>
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Thorell, 1890 (4C-I) in having similar median apophysis and terminal apophysis of the male pedipalp and variable sclerotization of the lateral plates at the base of the epigyne (Figs
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<figureCitation id="CCB025CF8DCA92404DCB17AD6BD822E6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">2A-D</figureCitation>
|
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,
|
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<figureCitation id="337C8C89D4D42E990B097BDCE081C28B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3C-I</figureCitation>
|
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,
|
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<figureCitation id="852B6F8BDC796CAC59B9777798C4FB3B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Passiena spinicrus Thorell, 1890, male (A, C-G) and female (B, H, I). A male habitus, dorsal view B female habitus, dorsal view C bulb, ventral view D same, retrolateral view E pedipalp, ventral view F same, retrolateral view G embolic division, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure4" httpUri="https://binary.pensoft.net/fig/919167" pageId="0" pageNumber="1">4C-I</figureCitation>
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;
|
||
<bibRefCitation id="C492F76002060079A33F9C1B5F22A324" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
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, figs 30-35). However, it can be distinguished by the combination of the following characters: 1) apical edge of anterior apophysis of palea as long as the stalk of posterior apophysis of palea (Fig.
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<figureCitation id="6FD84331CF206BABC650741C22BF552D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3G</figureCitation>
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) vs. about half the length of the stalk of posterior apophysis of palea in
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<taxonomicName id="E88CC566080DDD05D219924AA61EA4FB" lsidName="P. bayi" pageId="0" pageNumber="1" rank="species" species="bayi">
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<emphasis id="72CB10D82AAE461BF31F7C608B523176" italics="true" pageId="0" pageNumber="1">P. bayi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="2587C43902D5E545FE60269F16A0D0D9" lsidName="P. torbjoerni" pageId="0" pageNumber="1" rank="species" species="torbjoerni">
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<emphasis id="BA5DCDAAD522128318A075D7597D2E1B" italics="true" pageId="0" pageNumber="1">P. torbjoerni</emphasis>
|
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</taxonomicName>
|
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and
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<taxonomicName id="D20CE56F6D9630E909861EFA7ECD49A4" lsidName="P. spinicrus" pageId="0" pageNumber="1" rank="species" species="spinicrus">
|
||
<emphasis id="C96C52C8587BBBB76DFC284B6240EC0F" italics="true" pageId="0" pageNumber="1">P. spinicrus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="0D23C144984F5D555E7FD9DE724C1E85" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
|
||
, figs 25, 26; Fig.
|
||
<figureCitation id="0524B21F6427235BE6719C27268A590F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Passiena spinicrus Thorell, 1890, male (A, C-G) and female (B, H, I). A male habitus, dorsal view B female habitus, dorsal view C bulb, ventral view D same, retrolateral view E pedipalp, ventral view F same, retrolateral view G embolic division, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure4" httpUri="https://binary.pensoft.net/fig/919167" pageId="0" pageNumber="1">4G</figureCitation>
|
||
); 2) cymbium apex with two claws (Fig.
|
||
<figureCitation id="C685B2F25A62CC61FDB35D8F43DFF979" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3G</figureCitation>
|
||
) vs. with one claw in
|
||
<taxonomicName id="EB1B1D6B278716F0066A95E112738AB2" lsidName="P. bayi" pageId="0" pageNumber="1" rank="species" species="bayi">
|
||
<emphasis id="7CF459DEDDD65E309BF1D35FB09E7059" italics="true" pageId="0" pageNumber="1">P. bayi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D0D781B66569655580125CE02B4F732F" lsidName="P. torbjoerni" pageId="0" pageNumber="1" rank="species" species="torbjoerni">
|
||
<emphasis id="4DD36F8D2931B531BB425DBEFA33C644" italics="true" pageId="0" pageNumber="1">P. torbjoerni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8839B2C0E7A544CBC5F5C5B7C5ED5C58" lsidName="P. spinicrus" pageId="0" pageNumber="1" rank="species" species="spinicrus">
|
||
<emphasis id="81F45A27D40347D322074A3DBD305849" italics="true" pageId="0" pageNumber="1">P. spinicrus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="9BE5D8FF84A13A90C051BAD01DCAD6B2" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
|
||
, figs 27, 28; Fig.
|
||
<figureCitation id="51009432B15C9C57A9D78FA46979A9B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Passiena spinicrus Thorell, 1890, male (A, C-G) and female (B, H, I). A male habitus, dorsal view B female habitus, dorsal view C bulb, ventral view D same, retrolateral view E pedipalp, ventral view F same, retrolateral view G embolic division, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure4" httpUri="https://binary.pensoft.net/fig/919167" pageId="0" pageNumber="1">4G</figureCitation>
|
||
); and 3) epigynal septum stem length/septum base width ratio 1.2 in
|
||
<taxonomicName id="2C959190B88875B3B4CEA2305398DB4F" lsidName="P. duani" pageId="0" pageNumber="1" rank="species" species="duani">
|
||
<emphasis id="835129333950C363F291F265FDA212C5" italics="true" pageId="0" pageNumber="1">P. duani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="07A7E5B7DB0226F6FDB9DFE4B9394ACD" lsidName="P. torbjoerni" pageId="0" pageNumber="1" rank="species" species="torbjoerni">
|
||
<emphasis id="9C51761936921017AF8DC78A1EF3C988" italics="true" pageId="0" pageNumber="1">P. torbjoerni</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="EBF9CF39E58D11001FAF59EF7FF692D8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">2C, D</figureCitation>
|
||
,
|
||
<figureCitation id="4D6C4E96C41CCCEB6773D62D5E4BF5FB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3H, I</figureCitation>
|
||
;
|
||
<bibRefCitation id="513F08D4D62FE47B1BD278E0FCC30FF6" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
|
||
, figs 33, 34) vs. septum stem length/septum base width ratio 1.5 in
|
||
<taxonomicName id="A161B0E363BF9F3EEB23AB411DFF4A3A" lsidName="P. bayi" pageId="0" pageNumber="1" rank="species" species="bayi">
|
||
<emphasis id="1C104587342D0323694F61840FFE8669" italics="true" pageId="0" pageNumber="1">P. bayi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="70BE6FB8265F39F828C800441F6FB328" DOI="https://doi.org/10.26107/RBZ-2020-0066" author="Omelko, MM" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="479 - 487" refId="B4" refString="Omelko, MM, Marusik, YM, 2020. A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos. The Raffles Bulletin of Zoology 68: 479 - 487, DOI: https://doi.org/10.26107/RBZ-2020-0066" title="A new species and new records of wolf spiders (Araneae: Lycosidae) from Laos." url="https://doi.org/10.26107/RBZ-2020-0066" volume="68" year="2020">Omelko and Marusik 2020</bibRefCitation>
|
||
, figs 30, 31) and
|
||
<taxonomicName id="EFB787578968FC76A5BB69D998AC67AD" lsidName="P. spinicrus" pageId="0" pageNumber="1" rank="species" species="spinicrus">
|
||
<emphasis id="768EB4A5700B7BF058D44A04E51D6B27" italics="true" pageId="0" pageNumber="1">P. spinicrus</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="382A777C5A02AEB157C028E209FF95A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Passiena spinicrus Thorell, 1890, male (A, C-G) and female (B, H, I). A male habitus, dorsal view B female habitus, dorsal view C bulb, ventral view D same, retrolateral view E pedipalp, ventral view F same, retrolateral view G embolic division, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure4" httpUri="https://binary.pensoft.net/fig/919167" pageId="0" pageNumber="1">4H, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B47BCC536ABF4D817CD519A66A4BFF15" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="EE8EC5208E7489A0DAE4405684C0B7BA" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="0113998CF9E4E847F186AF272743D686" pageId="0" pageNumber="1">
|
||
<emphasis id="B23B81095E2E4D1F3D5786FFA0E00BED" bold="true" pageId="0" pageNumber="1">Male</emphasis>
|
||
(holotype, Fig.
|
||
<figureCitation id="C364A622C71184AF040D3703AB50FABA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live photo of Passiena duani sp. nov. (A, B) and P. spinicrus Thorell, 1890 (C, D) A, C male B, D female. Live photos taken by Qian-Le Lu (A, B) and Lu-Yu Wang (C, D)." figureDoi="10.3897/zookeys.1182.109532.figure1" httpUri="https://binary.pensoft.net/fig/919164" pageId="0" pageNumber="1">1A</figureCitation>
|
||
,
|
||
<figureCitation id="101FE08CCDEB12A6EED52F3DCDB8ABF1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3A</figureCitation>
|
||
). Total length 4.08. Prosoma 2.05 long, 1.59 wide; opisthosoma 1.98 long, 1.30 wide. Carapace greyish brown. Eye sizes and interdistances: AME 0.09, ALE 0.07, PME 0.30, PLE 0.25; AME-AME 0.09, AME-ALE 0.04, PME-PME 0.33, PME-PLE 0.32. Clypeus height 0.18. Chelicerae dark brown, with three promarginal and three retromarginal teeth. Endites and labium dark brown, longer than wide. Sternum yellow brown, with sparse brown setae. Legs yellow brown. Tibia I with six pairs of ventral spines and metatarsus I with four pairs of ventral spines; tibia II with five pairs of ventral spines, metatarsus II with three pairs of ventral spines. Leg measurements: I 6.14 (1.63, 2.21, 1.48, 0.82); II 5.47 (1.54, 1.79, 1.36, 0.78); III 5.23 (1.46, 1.57, 1.43, 0.77); IV 8.02 (2.07, 2.38, 2.49, 1.08). Leg formula: 4123. Opisthosoma oval. Dorsum greyish brown, with black markings. Venter yellow brown.
|
||
</paragraph>
|
||
<caption id="81E0ABBEEF5B2B5DB8D1694EF16FA913" doi="10.3897/zookeys.1182.109532.figure1" httpUri="https://binary.pensoft.net/fig/919164" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
|
||
<paragraph id="54BC62B7152D8D0354CE0C2C5F00B4ED" pageId="0" pageNumber="1">
|
||
<emphasis id="F7E37B01ABE617D486BA78F7BC7A7ACA" bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
|
||
Live photo of
|
||
<taxonomicName id="2DDC337E03AE4577101C945ADECBF518" class="Arachnida" family="Lycosidae" genus="Passiena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passiena duani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="duani">
|
||
<emphasis id="E1E636BF43587D5C8DF9380A8CEB8186" italics="true" pageId="0" pageNumber="1">Passiena duani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis id="B7D39EF8C9C47F4DD9712C080900BAEB" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
) and
|
||
<taxonomicName id="D93B64BCC5F2326B16587A290225D733" lsidName="P. spinicrus" pageId="0" pageNumber="1" rank="species" species="spinicrus">
|
||
<emphasis id="E4601AB1EF4EA0E5FF32F486C4B8DA27" italics="true" pageId="0" pageNumber="1">P. spinicrus</emphasis>
|
||
</taxonomicName>
|
||
Thorell, 1890 (
|
||
<emphasis id="97D22EA6A0C7F8E2FF1D7D9C109C4B34" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
)
|
||
<emphasis id="9634FDD189DBAEA18BF1C1769C33642C" bold="true" pageId="0" pageNumber="1">A, C</emphasis>
|
||
male
|
||
<emphasis id="606AA441953537F9461B605B495D1BCD" bold="true" pageId="0" pageNumber="1">B, D</emphasis>
|
||
female. Live photos taken by Qian-Le Lu (
|
||
<emphasis id="CF8A609AF85BB6D3C8EAD4FE547416FF" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
) and Lu-Yu Wang (
|
||
<emphasis id="BFD9AD2985B2FF93CC4D11ADC0B6C3B4" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F9B92508E557D1771835A81876944C47" pageId="0" pageNumber="1">
|
||
<emphasis id="77FD4008F2747ABDE72CBD8F28C5A633" bold="true" italics="true" pageId="0" pageNumber="1">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation id="81ACFC70839596F3F5AF02058B5C1CDF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="4F4C10E60CD45A3DAC93FECA9BF9653E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3C-G</figureCitation>
|
||
): Cymbium proximal part brown, distal part yellowish with two large claws on the tip. Subtegulum distinct in ventral view, located baso-prolaterally. Conductor somewhat membranous, somewhat tongue-shaped in ventral view and triangular in retrolateral view. Terminal apophysis terminates at approx. 1
|
||
<normalizedToken id="3D0CAA4A85C105D975BFC4BF4FD11A61" originalValue="o’clock">o'clock</normalizedToken>
|
||
position in ventral view. Embolus originating on the dorsal side of the bulb, long, prolaterally accompanied with a membrane, terminating at approx. 2
|
||
<normalizedToken id="7029FA71E3F6F33B5AB2884591BE5A7E" originalValue="o’clock">o'clock</normalizedToken>
|
||
position; palea with two apophyses, anterior apophysis with smooth apical edge sharply pointed, and posterior one claw-like.
|
||
</paragraph>
|
||
<caption id="A598A2BAF65FC3D9C96157BCBC2CF8EE" doi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
|
||
<paragraph id="F924D82875E573D9100CF231D35FF0CA" pageId="0" pageNumber="1">
|
||
<emphasis id="4A930D0C17E8711564C85E3CB5A2B00B" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
|
||
<taxonomicName id="CBAC7068CC0FAB51402118F6BE2EC0B0" class="Arachnida" family="Lycosidae" genus="Passiena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passiena duani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="duani">
|
||
<emphasis id="66DD50EF93BF06D3A15E18FEC308D26F" italics="true" pageId="0" pageNumber="1">Passiena duani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male holotype (
|
||
<emphasis id="A991BCD730CDA32B1C293D32CD424A79" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
) and female paratype (
|
||
<emphasis id="808597BBF1DBBA884CF56D575D538D94" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
)
|
||
<emphasis id="2FC7870DA4165BE48881F8D2EA0D6484" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
pedipalp, ventral view
|
||
<emphasis id="90D8DAFDCCBF06E1C2B84189DA17A7B5" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
same, retrolateral view
|
||
<emphasis id="1370E4B96F4801C9AB22A63F02EBD5E2" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
epigyne, ventral view
|
||
<emphasis id="08BEECA7051E12F74EC66528333AB29D" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
same, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8554ECBE3652D774387694E94A2552F9" doi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||
<paragraph id="61AC8DE17A1CD3D0BE78F0A4FCECEA2F" pageId="0" pageNumber="1">
|
||
<emphasis id="92268244CD525D48956995AC8BFB86B9" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||
<taxonomicName id="A294B9F3AE9BCFA120C21DFC0305DCD9" class="Arachnida" family="Lycosidae" genus="Passiena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passiena duani" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="duani">
|
||
<emphasis id="717EC2BE22274DF9E225FB26BCBEB84B" italics="true" pageId="0" pageNumber="1">Passiena duani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male holotype (
|
||
<emphasis id="25026910CE4385883E90AA8ABA583756" bold="true" pageId="0" pageNumber="1">A, C-G</emphasis>
|
||
) and female paratype (
|
||
<emphasis id="082B2CBF6A3F6D935D86C6EDE2FC00DC" bold="true" pageId="0" pageNumber="1">B, H, I</emphasis>
|
||
)
|
||
<emphasis id="E7C03400712D9E3E8F0B448D0F1C2431" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
male habitus, dorsal view
|
||
<emphasis id="385483274FB125C70A411413170FED57" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
female habitus, dorsal view
|
||
<emphasis id="3090919A6F4A3600A20B9B9D5647D493" bold="true" pageId="0" pageNumber="1">C, E</emphasis>
|
||
pedipalp, ventral view
|
||
<emphasis id="487298B859253B20249F9869C44E588D" bold="true" pageId="0" pageNumber="1">D, F</emphasis>
|
||
same, retrolateral view
|
||
<emphasis id="89007D96A54AE5DC421036C78F4DF707" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
embolus and terminal apophysis, ventral view
|
||
<emphasis id="CDDA328C35373BF878FEDA645DA80992" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
||
epigyne, ventral view
|
||
<emphasis id="0F7659B8502B28DF91BA8149A7B9D5D6" bold="true" pageId="0" pageNumber="1">I</emphasis>
|
||
same, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="29F525F1F6ACF5FC2178E40F0DBBB360" pageId="0" pageNumber="1">
|
||
<emphasis id="59CEF3DE2A1888E26875762D295EC309" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||
(one paratype, SWUC-T-LY-13-02, Fig.
|
||
<figureCitation id="D4C8D918F44343719F1731661DA35666" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live photo of Passiena duani sp. nov. (A, B) and P. spinicrus Thorell, 1890 (C, D) A, C male B, D female. Live photos taken by Qian-Le Lu (A, B) and Lu-Yu Wang (C, D)." figureDoi="10.3897/zookeys.1182.109532.figure1" httpUri="https://binary.pensoft.net/fig/919164" pageId="0" pageNumber="1">1B</figureCitation>
|
||
,
|
||
<figureCitation id="7B057399B272E88B6F79A7A812F8CFD3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3B</figureCitation>
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). Total length 4.57. Prosoma 2.36 long, 1.83 wide; opisthosoma 2.28 long, 1.65 wide. Eye sizes and interdistances: AME 0.12, ALE 0.09, PME 0.35, PLE 0.27; AME-AME 0.10, AME-ALE 0.07, PME-PME 0.33, PME-PLE 0.38. Clypeus height 0.23. Leg measurements: I 6.87 (1.92, 2.43, 1.65, 0.87); II 6.12 (1.58, 2.18, 1.53, 0.83); III 5.96 (1.68, 1.77, 1.70, 0.81); IV 9.28 (2.28, 2.75, 2.98, 1.27). Leg formula: 4123. Tibia I with six pairs of ventral spines and metatarsus I with four pairs of ventral spines; tibia II with six pairs of ventral spines, metatarsus II with four pairs of ventral spines. Except genitalia, all other morphological characteristics same as in male.
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</paragraph>
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<paragraph id="990A97D6863DF61BE9E22EFAE0AF760A" pageId="0" pageNumber="1">
|
||
<emphasis id="19081849D6177B61BC1D1B3B6045705F" bold="true" italics="true" pageId="0" pageNumber="1">Epigyne</emphasis>
|
||
(Figs
|
||
<figureCitation id="BF86F0B36CD63D11E47BB39B0691ECFF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passiena duani sp. nov., male holotype (A, B) and female paratype (C, D) A pedipalp, ventral view B same, retrolateral view C epigyne, ventral view D same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure2" httpUri="https://binary.pensoft.net/fig/919165" pageId="0" pageNumber="1">2C-D</figureCitation>
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,
|
||
<figureCitation id="E20733DC3913C0B061F3919991AD1BA3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Passiena duani sp. nov., male holotype (A, C-G) and female paratype (B, H, I) A male habitus, dorsal view B female habitus, dorsal view C, E pedipalp, ventral view D, F same, retrolateral view G embolus and terminal apophysis, ventral view H epigyne, ventral view I same, dorsal view." figureDoi="10.3897/zookeys.1182.109532.figure3" httpUri="https://binary.pensoft.net/fig/919166" pageId="0" pageNumber="1">3H-I</figureCitation>
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||
). Anterior pocket with 2 hoods, septum reversed T-shaped with distinct stem becoming very thick in its anterior part and narrow posteriorly; stem 1.2 times longer than base width. Copulatory openings located posteriorly at the base of atrium transverse edges. Spermathecal heads sub-oval with the antero-lateral part angled, heads 2 times longer than septum base. Spermathecal stalks thick, short, slightly curved. Fertilization ducts teardrop-shaped.
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</paragraph>
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</subSubSection>
|
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<subSubSection id="DCE4F5284117F8D2CB6F215A6AB2C1C4" pageId="0" pageNumber="1" type="distribution">
|
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<paragraph id="B0D42B8D3FA6C6E4E60D8E008C5582F2" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="16392D2D96BE31E9A0E09357957F4D73" pageId="0" pageNumber="1">
|
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Currently known only from the type locality, Ningming County, Guangxi, China (Fig.
|
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<figureCitation id="907BE9320107443114DB687C9DB8EFC7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map showing distribution records of Passiena duani sp. nov. and P. spinicrus Thorell, 1890." figureDoi="10.3897/zookeys.1182.109532.figure5" httpUri="https://binary.pensoft.net/fig/919168" pageId="0" pageNumber="1">5</figureCitation>
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).
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</paragraph>
|
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</subSubSection>
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</treatment>
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</document> |