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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1087.64382" ID-GBIF-Dataset="9c9b7f37-a72e-4610-8ac5-908f601d1121" ID-PMC="PMC8888540" ID-Pensoft-Pub="1313-2970-1087-19" ID-Pensoft-UUID="ECD9B4DC2A3357AABC04DB88FB7D40B1" ID-PubMed="35437365" ID-ZooBank="94F2384E640E4A58B8B4D9D06675D2C2" ModsDocID="1313-2970-1087-19" checkinTime="1645627321867" checkinUser="pensoft" docAuthor="Nupponen 1, Kari &amp; Sihvonen, Pasi" docDate="2022" docId="403821CD76585DA2848913DDC6E8F2D3" docLanguage="en" docName="ZooKeys 1087: 19-104" docOrigin="ZooKeys 1087" docPubDate="2022-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1087.64382" docTitle="Scythris tibicina Meyrick 1916" docType="treatment" docVersion="4" id="ECD9B4DC2A3357AABC04DB88FB7D40B1" lastPageNumber="19" masterDocId="ECD9B4DC2A3357AABC04DB88FB7D40B1" masterDocTitle="Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)" masterLastPageNumber="104" masterPageNumber="19" pageNumber="19" updateTime="1668151506650" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nupponen 1, Kari</mods:namePart>
<mods:affiliation>Merenneidontie 19 D, FI- 02320 Espoo, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sihvonen, Pasi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2237-9325</mods:nameIdentifier>
<mods:affiliation>Finnish Museum of Natural History, P. O. Box 17, Pohjoinen Rautatiekatu 13, 00014 University of Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="email">pasi.sihvonen@helsinki.fi</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1087</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>104</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1087.64382</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1087.64382</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1087-19</mods:identifier>
<mods:identifier type="ZooBank">94F2384E640E4A58B8B4D9D06675D2C2</mods:identifier>
<mods:identifier type="Pensoft-UUID">ECD9B4DC2A3357AABC04DB88FB7D40B1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193585893" LSID="urn:lsid:plazi:treatment:403821CD76585DA2848913DDC6E8F2D3" httpUri="http://treatment.plazi.org/id/403821CD76585DA2848913DDC6E8F2D3" lastPageNumber="19" pageId="0" pageNumber="19">
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
<paragraph pageId="0" pageNumber="19">
<taxonomicName LSID="403821CD-7658-5DA2-8489-13DDC6E8F2D3" authority="Meyrick, 1916" authorityName="Meyrick" authorityYear="1916" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris tibicina" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="tibicina">Scythris tibicina Meyrick, 1916</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">
<figureCitation captionStart="Figures 1824A" captionStartId="F4" captionText="Figures 18 - 24 A. Scythrididae adults, genus Scythris 18 S. caimancitoensis Nupponen, sp. nov., male, holotype 19 S. ejiciens Meyrick, 1928, male, holotype 20 S. fluvialis Meyrick, 1916, male, lectotype 21 S. inanima Meyrick, 1916, male, holotype 22 A S. lequetepequensis Nupponen sp. nov., male, holotype 22 B S. lequetepequensis Nupponen sp. nov., male, paratype 23 S. plocogastra Meyrick, 1931, female, holotype 24 A S. tibicina Meyrick, 1916, male, lectotype." figureDoi="10.3897/zookeys.1087.64382.figures18-24a" httpUri="https://binary.pensoft.net/fig/649318" pageId="0" pageNumber="19">Figs 24</figureCitation>
<figureCitation captionStart="Figures 5354" captionStartId="F16" captionText="Figures 53 - 54. Male genitalia of Scythris 53 S. tibicina Meyrick, 1916, lectotype, slide JFGC No. 8053 (top right corner: tergum VIII and gnathos uncompressed, slide 5 / 11 Dec. 2019 KN 54 S. sanfranciscoensis Nupponen sp. nov., holotype, slide 2 / 10 Dec. 2019 KN." figureDoi="10.3897/zookeys.1087.64382.figures53-54" httpUri="https://binary.pensoft.net/fig/649330" pageId="0" pageNumber="19">, 53</figureCitation>
<figureCitation captionStart="Figure 69" captionStartId="F27" captionText="Figure 69. Female genitalia of Scythris tibicina Meyrick, 1916, slide 1 / 18 Dec. 2019 KN." figureDoi="10.3897/zookeys.1087.64382.figure69" httpUri="https://binary.pensoft.net/fig/649341" pageId="0" pageNumber="19">, 69</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="reference_group">
<paragraph pageId="0" pageNumber="19">
<taxonomicName authorityName="Meyrick" authorityYear="1916" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris tibicina" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="tibicina">Scythris tibicina</taxonomicName>
Meyrick, 1916. Exotic Microlepidoptera, vol. 2 (part 1): 12.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="19">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Parish, No." country="Peru" elevation="853" location="Peru" specimenCount="♂" typeStatus="Lectotype">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">
<typeStatus>Lectotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Peru">Peru</collectingCountry>
<specimenCount></specimenCount>
; Chosica;
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5344" unit="ft" value="2800.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5344" unit="ft" value="2800.0">2800 feet</quantity>
a.s.l.
</elevation>
; 7.14.;
<collectorName>Parish</collectorName>
leg.; [genitalia slide] JFGC
<collectorName>No.</collectorName>
8053; NHMUK ID 010922365; NHMUK slide ID 010316665; coll. NHMUK.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="19">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Parish &amp; No." country="Peru" elevation="853" location="Peru" specimenCount="1" typeStatus="Paralectotype">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">
<typeStatus>Paralectotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Peru">Peru</collectingCountry>
• 11 exx.; same data as for lectotype; coll. NHMUK
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
<paragraph pageId="0" pageNumber="19">Other material.</paragraph>
<paragraph pageId="0" pageNumber="19">
<materialsCitation collectingDate="2019-02-05" collectorName="K. Nupponen, R. Haverinen" country="Peru" elevation="1520" latitude="-10.506666" location="Huanchay village" longLatPrecision="129" longitude="-77.425" specimenCount="1" specimenCount-male="1" stateProvince="Ancash">
<collectingCountry name="Peru">Peru</collectingCountry>
<specimenCount type="male">1 ♂</specimenCount>
; prov.
<collectingRegion country="Peru" name="Ancash">Ancash</collectingRegion>
, near
<location LSID="urn:lsid:plazi:treatment:403821CD76585DA2848913DDC6E8F2D3:CA05BE8E798B70DCCEF371BF2E781E88" country="Peru" latitude="-10.506666" longLatPrecision="129" longitude="-77.425" name="Huanchay village" stateProvince="Ancash">Huanchay village</location>
;
<geoCoordinate degrees="10" direction="south" minutes="30.4" orientation="latitude" precision="92" value="-10.506666">10°30.4'S</geoCoordinate>
,
<geoCoordinate degrees="77" direction="west" minutes="25.5" orientation="longitude" precision="92" value="-77.425">77°25.5'W</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.52" unit="m" value="1520.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.52" unit="m" value="1520.0">1520 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2019-02-05">5 Feb. 2019</collectingDate>
;
<collectorName>K. Nupponen</collectorName>
&amp;
<collectorName>R. Haverinen</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="2019-12-11" country="Peru" location="Nupponen" specimenCount="1">
[BOLD sample ID] KN01075; [genitalia slide] K.
<location LSID="urn:lsid:plazi:treatment:403821CD76585DA2848913DDC6E8F2D3:7A061D32D87E17473B280124EF928511" country="Peru" name="Nupponen">Nupponen</location>
prep. no. 5/
<collectingDate value="2019-12-11">11 Dec. 2019</collectingDate>
</materialsCitation>
; coll. NUPP. •
<materialsCitation collectingDate="2019-01-31" collectorName="K. Nupponen, R. Haverinen" country="Peru" elevation="1180" latitude="-10.218333" location="Raquia village" longLatPrecision="129" longitude="-77.56" municipality="Fortaleza River" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Ancash">
<specimenCount type="male">2 ♂</specimenCount>
,
<specimenCount type="female">2 ♀</specimenCount>
; prov.
<collectingRegion country="Peru" name="Ancash">Ancash</collectingRegion>
,
<collectingMunicipality>Fortaleza River</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:403821CD76585DA2848913DDC6E8F2D3:C66EEF449D1E5A7D69EE400E23DDEE2F" country="Peru" latitude="-10.218333" longLatPrecision="129" longitude="-77.56" municipality="Fortaleza River" name="Raquia village" stateProvince="Ancash">Raquia village</location>
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
SW;
<geoCoordinate degrees="10" direction="south" minutes="13.1" orientation="latitude" precision="92" value="-10.218333">10°13.1'S</geoCoordinate>
,
<geoCoordinate degrees="77" direction="west" minutes="33.6" orientation="longitude" precision="92" value="-77.56">77°33.6'W</geoCoordinate>
;
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.18" unit="m" value="1180.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.18" unit="m" value="1180.0">1180 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2019-01-31">31 Jan. 2019</collectingDate>
;
<collectorName>K. Nupponen</collectorName>
&amp;
<collectorName>R. Haverinen</collectorName>
leg.
</materialsCitation>
<materialsCitation collectingDate="2019-12-18" country="Peru" location="Nupponen" specimenCount="1">
[BOLD sample IDs] KN01076, KN01077; [genitalia slides] K.
<location LSID="urn:lsid:plazi:treatment:403821CD76585DA2848913DDC6E8F2D3:42AD06CFD7BE1B22ACB1891B54C9EAD2" country="Peru" name="Nupponen">Nupponen</location>
prep. no. 1/
<collectingDate value="2019-12-18">18-XII-2019</collectingDate>
</materialsCitation>
<specimenCount></specimenCount>
, 4/
<date value="2019-12-17">17-XII-2019</date>
<specimenCount></specimenCount>
; coll. NUPP.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="19">Forewings with whitish streak on brownish background. Genitalia dissection is required for confident determination. The male genitalia are unmistakable, particularly the narrow, ventrally curved, hook-shaped gnathos; and phallus that bends at 90° angle; and densely bristled valvae. In the female genitalia, a crater-shaped margin of sterigma, adjoined by needle-like sclerotisation, are diagnostic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
<paragraph pageId="0" pageNumber="19">The original description is quoted: &quot;Wingspan 12-13 mm ♂, ♀. Head ochreous-grey more or less mixed with white. Palpi grey, suffused with white internally and at apex of second joint. Antennal ciliations of ♂ 1. Thorax ochreous-grey partially mixed with whitish. Abdomen light grey, anal tuft pale ochreous, ventral surface whitish. Forewings lanceolate; light grey: a double finely separated or united median whitish streak, from base, upper portion extending to about middle, lower to 0.33, both more or less enlarged into suffused spots posteriorly; an irregular elongate undefined spot of whitish suffusion in disc at 0.66; each of these whitish markings followed by a few indistinct dark fuscous scales, representing the stigmata: cilia grey, base mixed with whitish. Hindwings with 4 and 5 separate; grey; cilia grey.&quot;</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Male genitalia</emphasis>
.
</emphasis>
Uncus posterolaterally extended trapezoid plate, margin concave medially. Gnathos asymmetrical, basally channel-like, apex spoon-shaped. Distal arm of gnathos thin, curved ventrally, hook-shaped. Tegumen hood-shaped. Phallus 0.6
<normalizedToken originalValue="×">x</normalizedToken>
length of valva; basal 2/3 straight, then bent at 90° angle, distal 1/3 slender and straight, tip pointed. Valva long and narrow, bent at 0.4 length, distal portion straight and setose; ventrally at middle sub-oval bristled extension. Vinculum arched, short. Sternum VIII large trapezoid plate, medioposteriorly with small V-shaped indentation, laterally at 0.3 with anteriorly directed lobes. Tergum VIII small trapezoid plate.
</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Female genitalia</emphasis>
.
</emphasis>
Sterigma crater-shaped, twice as wide as high, adjoined by needle-like sclerotisation. Ostium situated at bottom of crater. Sternum VII semi-circular, medioposteriorly with small concave notch. Apophyses anteriores 0.25
<normalizedToken originalValue="×">x</normalizedToken>
length of apophyses posteriores.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="distribution">
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="19">Peru.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="habitat">
<paragraph pageId="0" pageNumber="19">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="19">Adults were collected in moist riverside meadows.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="genetic data">
<paragraph pageId="0" pageNumber="19">Genetic data.</paragraph>
<paragraph pageId="0" pageNumber="19">
BIN: BOLD:ADZ4797 (
<emphasis italics="true" pageId="0" pageNumber="19">n</emphasis>
= 3 from Peru). Genetically homogenous, variation 0%. Nearest neighbour: Unidentified
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" family="Scythrididae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="19" rank="family">Scythrididae</taxonomicName>
from Argentina (BIN: BOLD:ACY3332, 6.54%), see Suppl. material 2.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="remarks">
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="19">
Based on COI maximum likelihood phylogeny, taxa
<taxonomicName authorityName="Meyrick" authorityYear="1916" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris tibicina" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="tibicina">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris tibicina</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris sanfranciscoensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="sanfranciscoensis">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris sanfranciscoensis</emphasis>
</taxonomicName>
group together, associating with other Central and South American taxa, classified in apparently non-monophyletic
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris</emphasis>
</taxonomicName>
on BOLD (Suppl. material 2). With regard to
<taxonomicName authorityName="Meyrick" authorityYear="1916" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris tibicina" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="tibicina">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris tibicina</emphasis>
</taxonomicName>
, the male genitalia are similar to
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. mixaula" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="mixaula">
<emphasis italics="true" pageId="0" pageNumber="19">S. mixaula</emphasis>
</taxonomicName>
Meyrick, 1916 from California, Texas and Montana, sharing for instance narrow and setose valva, spear-shaped uncus (termed distal arm of gnathos in
<bibRefCitation DOI="https://doi.org/10.4039/entm123160fv" author="Landry, JF" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="19" pagination="1 - 341" refId="B22" refString="Landry, JF, 1991. Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species. Memoirs of the Entomological Society of Canada 160: 1 - 341, DOI: https://doi.org/10.4039/entm123160fv" title="Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species." url="https://doi.org/10.4039/entm123160fv" volume="160" year="1991">Landry (1991)</bibRefCitation>
) and mediodorsally convex vinculum. We have interpreted the mediodorsal structure as uncus (gnathos in
<bibRefCitation DOI="https://doi.org/10.4039/entm123160fv" author="Landry, JF" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="19" pagination="1 - 341" refId="B22" refString="Landry, JF, 1991. Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species. Memoirs of the Entomological Society of Canada 160: 1 - 341, DOI: https://doi.org/10.4039/entm123160fv" title="Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species." url="https://doi.org/10.4039/entm123160fv" volume="160" year="1991">Landry (1991)</bibRefCitation>
and the sclerotised structure on its ventral side as gnathos. We justify this interpretation by the origin of the narrow and ventrally curved process, which originates from the cup-shaped apex of gnathos. See Fig.
<figureCitation captionStart="Figures 5354" captionStartId="F16" captionText="Figures 53 - 54. Male genitalia of Scythris 53 S. tibicina Meyrick, 1916, lectotype, slide JFGC No. 8053 (top right corner: tergum VIII and gnathos uncompressed, slide 5 / 11 Dec. 2019 KN 54 S. sanfranciscoensis Nupponen sp. nov., holotype, slide 2 / 10 Dec. 2019 KN." figureDoi="10.3897/zookeys.1087.64382.figures53-54" httpUri="https://binary.pensoft.net/fig/649330" pageId="0" pageNumber="19">53</figureCitation>
, which shows the origin of the structure in lateral view. We classify
<taxonomicName authorityName="Meyrick" authorityYear="1916" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris tibicina" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="tibicina">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris tibicina</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Nupponen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris sanfranciscoensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="sanfranciscoensis">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris sanfranciscoensis</emphasis>
</taxonomicName>
in
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.1087.64382.figures18-24a" httpUri="https://binary.pensoft.net/fig/649318" pageId="0" pageNumber="19" start="Figures 1824A" startId="F4">
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">Figures 18-24A.</emphasis>
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" family="Scythrididae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="19" rank="family">Scythrididae</taxonomicName>
adults, genus
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Scythris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scythris" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Scythris</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="19">18</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. caimancitoensis" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="caimancitoensis">
<emphasis italics="true" pageId="0" pageNumber="19">S. caimancitoensis</emphasis>
</taxonomicName>
Nupponen, sp. nov., male, holotype
<emphasis bold="true" pageId="0" pageNumber="19">19</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. ejiciens" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="ejiciens">
<emphasis italics="true" pageId="0" pageNumber="19">S. ejiciens</emphasis>
</taxonomicName>
Meyrick, 1928, male, holotype
<emphasis bold="true" pageId="0" pageNumber="19">20</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. fluvialis" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="fluvialis">
<emphasis italics="true" pageId="0" pageNumber="19">S. fluvialis</emphasis>
</taxonomicName>
Meyrick, 1916, male, lectotype
<emphasis bold="true" pageId="0" pageNumber="19">21</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. inanima" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="inanima">
<emphasis italics="true" pageId="0" pageNumber="19">S. inanima</emphasis>
</taxonomicName>
Meyrick, 1916, male, holotype
<emphasis bold="true" pageId="0" pageNumber="19">22A</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. lequetepequensis" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="lequetepequensis">
<emphasis italics="true" pageId="0" pageNumber="19">S. lequetepequensis</emphasis>
</taxonomicName>
Nupponen sp. nov., male, holotype
<emphasis bold="true" pageId="0" pageNumber="19">22B</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. lequetepequensis" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="lequetepequensis">
<emphasis italics="true" pageId="0" pageNumber="19">S. lequetepequensis</emphasis>
</taxonomicName>
Nupponen sp. nov., male, paratype
<emphasis bold="true" pageId="0" pageNumber="19">23</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. plocogastra" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="plocogastra">
<emphasis italics="true" pageId="0" pageNumber="19">S. plocogastra</emphasis>
</taxonomicName>
Meyrick, 1931, female, holotype
<emphasis bold="true" pageId="0" pageNumber="19">24A</emphasis>
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" lsidName="S. tibicina" order="Caryophyllales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="species" species="tibicina">
<emphasis italics="true" pageId="0" pageNumber="19">S. tibicina</emphasis>
</taxonomicName>
Meyrick, 1916, male, lectotype.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>