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<document id="CC021C09E1DB8F3C413BE9378454F377" ID-DOI="10.5281/zenodo.5341416" ID-ISSN="0374-1036" ID-Zenodo-Dep="5341416" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190255773" checkinUser="felipe" docAuthor="Matocq, Armand" docDate="2008" docId="0E494614E344EE1839BCFA23FDD405C6" docLanguage="en" docName="ActaEntMusNatPra.2008.48.2.419-422.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Megalocoleus stysi Matocq 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="422" masterDocId="F2703E6CE344EE1B3855FFB5FFDE0724" masterDocTitle="A new species of Megalocoleus (Hemiptera: Heteroptera: Miridae: Phylinae) from Morocco" masterLastPageNumber="422" masterPageNumber="419" pageNumber="419" updateTime="1699181346709" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="068A9DE106BF104E48029FF8E05C0404" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85E9BAD9C59BC8BA4A36F5CA776B14A7">
<mods:title id="67C147514DA6545BBA8C6F4657418111">A new species of Megalocoleus (Hemiptera: Heteroptera: Miridae: Phylinae) from Morocco</mods:title>
</mods:titleInfo>
<mods:name id="53CA731F6D04A910ED2759DADF3AADD4" type="personal">
<mods:role id="323752E5E3A27927BEBD6DFE31B45200">
<mods:roleTerm id="C980354C5FBB0A2216CA9F5E56611806">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9146F84A493FDC4C74432045FD4EF89B">Matocq, Armand</mods:namePart>
</mods:name>
<mods:typeOfResource id="454A26FC20B4CDF7854535E025E8A468">text</mods:typeOfResource>
<mods:relatedItem id="68B184AA7D553B4E83A8DB30685E4BB8" type="host">
<mods:titleInfo id="F711CE00A57AFA12B19958AF42FC157B">
<mods:title id="F604E59C5391F6ACDA83E7049888A143">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="7734556D7934A6752DE72B5A7FA80227">
<mods:date id="E01A0A94B92B79122309DC269ED4BA9B">2008</mods:date>
<mods:detail id="F9F575B2E056F148557C256C937963B3" type="pubDate">
<mods:number id="61A8C5DDDCDA13B173EB73E24E2DD460">2008-12-08</mods:number>
</mods:detail>
<mods:detail id="6C76C8AC920D68226BC3266EE124CF9F" type="volume">
<mods:number id="7BE8160B6EF3E0BEF58645EAF95B7AA0">48</mods:number>
</mods:detail>
<mods:detail id="084292C6DC5597F986FACB281E0E1E65" type="issue">
<mods:number id="5E5001026EBD7D10796C2B9BDD720264">2</mods:number>
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<mods:start id="7062696D3A01A9E2413D420C3DA9C9AA">419</mods:start>
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</mods:extent>
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<mods:classification id="AA8A87CDD3C327C071D9C3D975B69EBA">journal article</mods:classification>
<mods:identifier id="B45D35D406EF21EF0FFAAC5E4878B7BB" type="DOI">10.5281/zenodo.5341416</mods:identifier>
<mods:identifier id="2B19EFFE973A5B8FEFDF33B6878EEFE8" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="8E76E71BB32EE005C860EC2648B63411" type="Zenodo-Dep">5341416</mods:identifier>
</mods:mods>
<treatment id="0E494614E344EE1839BCFA23FDD405C6" LSID="urn:lsid:plazi:treatment:0E494614E344EE1839BCFA23FDD405C6" httpUri="http://treatment.plazi.org/id/0E494614E344EE1839BCFA23FDD405C6" lastPageId="3" lastPageNumber="422" pageId="0" pageNumber="419">
<subSubSection id="CEFAA489E344EE1B39BCFA23FCCA0294" box="[489,788,1430,1456]" pageId="0" pageNumber="419" type="nomenclature">
<paragraph id="865FF702E344EE1B39BCFA23FCCA0294" blockId="0.[489,788,1430,1456]" box="[489,788,1430,1456]" pageId="0" pageNumber="419">
<heading id="DD17406EE344EE1B39BCFA23FCCA0294" bold="true" box="[489,788,1430,1456]" centered="true" fontSize="8" level="2" pageId="0" pageNumber="419" reason="3">
<emphasis id="B4942B10E344EE1B39BCFA23FCCA0294" bold="true" box="[489,788,1430,1456]" pageId="0" pageNumber="419">
<taxonomicName id="41E08C81E344EE1B39BCFA23FD66028B" ID-CoL="73826" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[489,696,1430,1455]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="419" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E344EE1B39BCFA23FD66028B" bold="true" box="[489,696,1430,1455]" italics="true" pageId="0" pageNumber="419">Megalocoleus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE344EE1B3AEAFA2DFCCA0294" box="[703,788,1432,1456]" pageId="0" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CEFAA489E344EE1B3A19FA03FD6F02ED" box="[588,689,1462,1481]" pageId="0" pageNumber="419" type="description">
<paragraph id="865FF702E344EE1B3A19FA03FD6F02ED" blockId="0.[151,1127,1462,1595]" box="[588,689,1462,1481]" pageId="0" pageNumber="419">
(
<figureCitation id="1EDBEB87E344EE1B3A06FA03FD7402ED" box="[595,682,1462,1481]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="1.[113,144,1110,1129]" captionStartId-1="2.[151,190,1060,1079]" captionTargetBox-1="[183,1090,220,1030]" captionTargetId-1="figure-215@2.[179,1098,210,1034]" captionTargetPageId-1="2" captionText-0="Fig. 1. Megalocoleus stysi sp. nov. Habitus of male and female." captionText-1="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi-1="http://doi.org/10.5281/zenodo.5341418" httpUri-1="https://zenodo.org/record/5341418/files/figure.png" pageId="0" pageNumber="419">Figs. 1-10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CEFAA489E344EE1B38C2FA6DFCB5011F" pageId="0" pageNumber="419" type="materials_examined">
<paragraph id="865FF702E344EE1B38C2FA6DFD8C02CF" blockId="0.[151,1127,1462,1595]" box="[151,594,1495,1515]" pageId="0" pageNumber="419">
<emphasis id="B4942B10E344EE1B38C2FA6DFECE02CF" bold="true" box="[151,272,1495,1515]" pageId="0" pageNumber="419">
<typeStatus id="595B49A0E344EE1B38C2FA6DFF1A02CF" box="[151,196,1496,1515]" pageId="0" pageNumber="419">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="FEF7B792E344EE1B3940FA6DFEBA02CF" box="[277,356,1496,1515]" name="Morocco" pageId="0" pageNumber="419">Morocco</collectingCountry>
, Kenitra, Foręt de Mamora.
</paragraph>
<paragraph id="865FF702E344EE1B38C2FA46FCB5011F" blockId="0.[151,1127,1462,1595]" pageId="0" pageNumber="419">
<emphasis id="B4942B10E344EE1B38C2FA46FEC60122" bold="true" box="[151,280,1522,1542]" pageId="0" pageNumber="419">Type material.</emphasis>
<materialsCitation id="3688FD5FE344EE1B3949FA46FBDA0122" box="[284,1028,1518,1543]" collectingDate="1967-05-05" collectionCode="MHNG" collectorName="de Mamora &amp; Eckerlein" country="Morocco" location="Kenitra" municipality="Foret de Mamora" pageId="0" pageNumber="419" specimenCount="1" typeStatus="holotype">
<typeStatus id="595B49A0E344EE1B3949FA46FEB00121" box="[284,366,1523,1542]" pageId="0" pageNumber="419" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingCountry id="FEF7B792E344EE1B39C6FA46FE3D0122" box="[403,483,1523,1542]" name="Morocco" pageId="0" pageNumber="419">Morocco</collectingCountry>
/
<location id="833FA1D9E344EE1B39A4FA46FDEE0122" LSID="urn:lsid:plazi:treatment:0E494614E344EE1839BCFA23FDD405C6:833FA1D9E344EE1B39A4FA46FDEE0122" box="[497,560,1523,1542]" country="Morocco" municipality="Foret de Mamora" name="Kenitra" pageId="0" pageNumber="419">Kenitra</location>
/
<collectingMunicipality id="663B6D78E344EE1B3A68FA46FD120122" box="[573,716,1523,1542]" pageId="0" pageNumber="419">
Forêt
<collectorName id="2B1592D4E344EE1B3A38FA46FD120122" box="[621,716,1523,1542]" pageId="0" pageNumber="419">de Mamora</collectorName>
</collectingMunicipality>
/
<date id="F25ED1C2E344EE1B3A8FFA46FCF70122" box="[730,809,1523,1542]" pageId="0" pageNumber="419" value="1967-05-05">
<collectingDate id="E21A282AE344EE1B3A8FFA46FCF70122" box="[730,809,1523,1542]" pageId="0" pageNumber="419" value="1967-05-05">5-V-1967</collectingDate>
</date>
/
<collectorName id="2B1592D4E344EE1B3B62FA46FC560122" box="[823,904,1523,1542]" pageId="0" pageNumber="419">Eckerlein</collectorName>
leg.’ (
<collectionCode id="E0F16FC7E344EE1B3BECFA46FC200122" box="[953,1022,1523,1542]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="419" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3688FD5FE344EE1B3C5FFA46FDC70104" collectingDate="1967-05-05" collectionCode="MHNG" collectorName="de Mamora &amp; Eckerlein" country="Morocco" location="Kenitra" municipality="Foret de Mamora" pageId="0" pageNumber="419" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<typeStatus id="595B49A0E344EE1B3C5FFA46FBBE0121" box="[1034,1120,1523,1542]" pageId="0" pageNumber="419" type="paratype">PARATYPES</typeStatus>
: 2 JJ
<specimenCount id="90E63C8BE344EE1B3899F9B8FF250104" box="[204,251,1543,1568]" pageId="0" pageNumber="419" type="female">6 ♀♀</specimenCount>
, same data as holotype (
<collectionCode id="E0F16FC7E344EE1B3998F9B8FDCD0104" box="[461,531,1549,1568]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="0" pageNumber="419" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3688FD5FE344EE1B3A74F9BBFDF9011F" collectingDate="1967-05-05" collectionCode="MHNG" collectorName="Matocq" country="Morocco" location="Kenitra" municipality="Foret de Mamora" pageId="0" pageNumber="419" specimenCount="12" specimenCount-female="1" typeStatus="paratype">
1 J
<specimenCount id="90E63C8BE344EE1B3A13F9BBFDB80104" box="[582,614,1543,1569]" pageId="0" pageNumber="419" type="female">1 ♀</specimenCount>
, same data as holotype (coll.
<collectorName id="2B1592D4E344EE1B3B36F9B8FC760104" box="[867,936,1549,1568]" pageId="0" pageNumber="419">Matocq</collectorName>
). The
<specimenCount id="90E63C8BE344EE1B3B89F9BBFB910104" box="[988,1103,1549,1569]" pageId="0" pageNumber="419" type="generic">11 specimens</specimenCount>
of the type series have lost most of their vestiture
</materialsCitation>
; setae can only be observed in places.
</paragraph>
</subSubSection>
<caption id="D29FA78AE345EE1A3824FBE3FD4E034D" box="[113,656,1110,1129]" pageId="1" pageNumber="420" startId="1.[113,144,1110,1129]">
<paragraph id="865FF702E345EE1A3824FBE3FD4E034D" blockId="1.[113,656,1110,1129]" box="[113,656,1110,1129]" pageId="1" pageNumber="420">
Fig. 1.
<taxonomicName id="41E08C81E345EE1A38FBFBE2FE8F034D" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[174,337,1111,1129]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E345EE1A38FBFBE2FE8F034D" box="[174,337,1111,1129]" italics="true" pageId="1" pageNumber="420">Megalocoleus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE345EE1A3903FBE3FE47034D" box="[342,409,1110,1129]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
Habitus of male and female.
</paragraph>
</caption>
<subSubSection id="CEFAA489E345EE193824FB07FC12022A" lastPageId="2" lastPageNumber="421" pageId="1" pageNumber="420" type="description">
<paragraph id="865FF702E345EE1A3824FB07FC83026E" blockId="1.[113,1090,1202,1642]" pageId="1" pageNumber="420">
<emphasis id="B4942B10E345EE1A3824FB07FF2803EE" bold="true" box="[113,246,1202,1226]" pageId="1" pageNumber="420">Description</emphasis>
(both sexes). Integument rather mat, body and appendages entirely greyish yellow except the last segment of rostrum and the last tarsal segments and claws of all legs, which are black. Dorsal vestiture with
<specimenCount id="90E63C8BE345EE1A3A40FB46FDA3022F" box="[533,637,1267,1291]" pageId="1" pageNumber="420" type="generic" typeStatus="types">two types</specimenCount>
of setae of unknown distribution (see the note above), fine and adpressed pale setae and semi-erect dark setae observed along lateral margin of hemelytra and on cuneus). Length (J and
<specimenCount id="90E63C8BE345EE1A3AF7FA99FD6B026F" box="[674,693,1324,1355]" pageId="1" pageNumber="420" type="female">♀</specimenCount>
) =
<quantity id="41185AE7E345EE1A3A8FFA87FC87026E" box="[730,857,1330,1354]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.5" metricValueMin="3.8" pageId="1" pageNumber="420" unit="mm" value="4.15" valueMax="4.5" valueMin="3.8">3.8-4.5 mm</quantity>
.
</paragraph>
<paragraph id="865FF702E345EE1A38C5FAE7FF79028E" blockId="1.[113,1090,1202,1642]" pageId="1" pageNumber="420">
Head short, not elongated (
<figureCitation id="1EDBEB87E345EE1A39F8FAE7FE30024E" box="[429,494,1362,1386]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="1" pageNumber="420">Fig. 2</figureCitation>
); length of antennal segments: I –
<quantity id="41185AE7E345EE1A3B0FFAE7FC64024E" box="[858,954,1362,1386]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="420" unit="mm" value="0.35">0.35 mm</quantity>
, II –
<quantity id="41185AE7E345EE1A3BBEFAE7FBE3024E" box="[1003,1085,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="420" unit="mm" value="1.1">1.1 mm</quantity>
, III –
<quantity id="41185AE7E345EE1A38FEFAC7FEDF02AE" box="[171,257,1394,1418]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="420" unit="mm" value="0.9">0.9 mm</quantity>
, IV –
<quantity id="41185AE7E345EE1A391CFAC7FE7702AE" box="[329,425,1394,1418]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="420" unit="mm" value="0.45">0.45 mm</quantity>
(
<figureCitation id="1EDBEB87E345EE1A39EFFAC7FE2302AE" box="[442,509,1394,1418]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="1" pageNumber="420">Fig. 3</figureCitation>
). Rostrum reaching the middle of abdominal sternite VIII.
</paragraph>
<paragraph id="865FF702E345EE1A38C5FA06FB9C02EE" blockId="1.[113,1090,1202,1642]" box="[144,1090,1458,1482]" pageId="1" pageNumber="420">Pronotum trapezoidal with slightly carinate lateral margin; xyphus concave and carinate.</paragraph>
<paragraph id="865FF702E345EE1A38C5FA67FC8C012E" blockId="1.[113,1090,1202,1642]" pageId="1" pageNumber="420">Hemelytra slightly translucent with more or less darkend spot in the middle (more pronounced in females); membrane smoky transparent with white veins.</paragraph>
<paragraph id="865FF702E345EE1A38C5F9A6FE07014E" blockId="1.[113,1090,1202,1642]" pageId="1" pageNumber="420">
Legs: femora without black spots; tibiae with black spines; tarsi: segments I and II greyish yellow, segment III and claw black; hind tarsi: segments II and III of hind tarsus subequal in length (
<figureCitation id="1EDBEB87E345EE1A3896F9E7FEDB014E" box="[195,261,1618,1642]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="1" pageNumber="420">Fig. 4</figureCitation>
); claws as in
<figureCitation id="1EDBEB87E345EE1A39C1F9E7FE0B014E" box="[404,469,1618,1642]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="1" pageNumber="420">Fig. 5</figureCitation>
.
</paragraph>
<caption id="D29FA78AE346EE1938C2FB91FE080348" ID-DOI="http://doi.org/10.5281/zenodo.5341418" ID-Zenodo-Dep="5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421" startId="2.[151,190,1060,1079]" targetBox="[183,1090,220,1030]" targetPageId="2">
<paragraph id="865FF702E346EE1938C2FB91FE080348" blockId="2.[151,1129,1060,1133]" pageId="2" pageNumber="421">
Figs. 2-10.
<taxonomicName id="41E08C81E346EE1938ADFB90FE450313" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[248,411,1061,1079]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE1938ADFB90FE450313" box="[248,411,1061,1079]" italics="true" pageId="2" pageNumber="421">Megalocoleus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE1939F4FB91FE3D0313" box="[417,483,1060,1079]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
(male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina.
</paragraph>
</caption>
<paragraph id="865FF702E346EE1938E0FB03FC12022A" blockId="2.[151,1127,1206,1646]" pageId="2" pageNumber="421">
Genitalia. Male: vesica slender, U-shaped; with a thin, straight and acute apex (
<figureCitation id="1EDBEB87E346EE193C40FB03FB8703EA" box="[1045,1113,1206,1230]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421">Fig. 6</figureCitation>
); secondary gonopore removed far from apex of vesica (
<figureCitation id="1EDBEB87E346EE193AA5FB63FCED03CA" box="[752,819,1238,1262]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421">Fig. 7</figureCitation>
); left paramere as in
<figureCitation id="1EDBEB87E346EE193C4AFB63FBBC03CA" box="[1055,1122,1238,1262]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421">Fig. 8</figureCitation>
. Female: vagina large and oval (
<figureCitation id="1EDBEB87E346EE1939B2FB43FDF7022A" box="[487,553,1270,1294]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421">Fig. 9</figureCitation>
). Ring sclerites subtriangular (
<figureCitation id="1EDBEB87E346EE193B25FB43FC1E022A" box="[880,960,1270,1294]" captionStart="Figs" captionStartId="2.[151,190,1060,1079]" captionTargetBox="[183,1090,220,1030]" captionTargetId="figure-215@2.[179,1098,210,1034]" captionTargetPageId="2" captionText="Figs. 2-10. Megalocoleus stysi sp. nov. (male holotype: 2 to 8). 2 – head, lateral view; 3 – antenna; 4 – hind tarsus; 5 – claw; 6 – vesica, lateral view; 7 – apex of vesica, different angle; 8 – left paramere; 9 – vagina, dorsal view; 10 – detail of left ring sclerite of vagina." figureDoi="http://doi.org/10.5281/zenodo.5341418" httpUri="https://zenodo.org/record/5341418/files/figure.png" pageId="2" pageNumber="421">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CEFAA489E346EE1838C2FAA3FB9F0550" lastPageId="3" lastPageNumber="422" pageId="2" pageNumber="421" type="diagnosis">
<paragraph id="865FF702E346EE1938C2FAA3FBFE02CA" blockId="2.[151,1127,1206,1646]" pageId="2" pageNumber="421">
<emphasis id="B4942B10E346EE1938C2FAA3FE4F020A" bold="true" box="[151,401,1302,1326]" pageId="2" pageNumber="421">Differential diagnosis.</emphasis>
<taxonomicName id="41E08C81E346EE1939CDFAA2FDBD020A" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[408,611,1303,1326]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE1939CDFAA2FDBD020A" box="[408,611,1303,1326]" italics="true" pageId="2" pageNumber="421">Megalocoleus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE193A3FFAA3FD630209" box="[618,701,1302,1325]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
shows some similarities with
<taxonomicName id="41E08C81E346EE193BAAFAA2FEF5026A" authority="(Fieber, 1861)" baseAuthorityName="Fieber" baseAuthorityYear="1861" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="lunula">
<emphasis id="B4942B10E346EE193BAAFAA2FBB8020A" box="[1023,1126,1303,1326]" italics="true" pageId="2" pageNumber="421">M. lunula</emphasis>
(Fieber, 1861)
</taxonomicName>
in the colouration and vestiture but differs from the latter species in the structure of the vesica. The vesica of
<taxonomicName id="41E08C81E346EE193992FAE2FDC4024A" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[455,538,1367,1390]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE193992FAE2FDC4024A" box="[455,538,1367,1390]" italics="true" pageId="2" pageNumber="421">M. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE193A76FAE3FDA80249" box="[547,630,1366,1389]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
is thin with a straight apex and the secondary gonopore is removed far from the apex of the vesica, while the vesica of
<taxonomicName id="41E08C81E346EE193BCDFAC2FC2002AA" baseAuthorityName="Fieber" baseAuthorityYear="1861" box="[920,1022,1399,1422]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="lunula">
<emphasis id="B4942B10E346EE193BCDFAC2FC2002AA" box="[920,1022,1399,1422]" italics="true" pageId="2" pageNumber="421">M. lunula</emphasis>
</taxonomicName>
is thicker with a curved apex and subapically placed secondary gonopore. The vesica of
<taxonomicName id="41E08C81E346EE193B94FA22FBCC028A" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[961,1042,1431,1454]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE193B94FA22FBCC028A" box="[961,1042,1431,1454]" italics="true" pageId="2" pageNumber="421">M. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE193C42FA23FBB90289" box="[1047,1127,1430,1453]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
is similar to that of
<taxonomicName id="41E08C81E346EE19393CFA02FD5202EA" authority="(Reuter, 1879)" baseAuthorityName="Reuter" baseAuthorityYear="1879" box="[361,652,1462,1486]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="krueperi">
<emphasis id="B4942B10E346EE19393CFA02FE3902EA" box="[361,487,1463,1486]" italics="true" pageId="2" pageNumber="421">M. krueperi</emphasis>
(Reuter, 1879)
</taxonomicName>
, but the latter species is easily distinguished by its general shape, short and incrassate antennae, and vestiture (see
<bibRefCitation id="E2718AF3E346EE193B2BFA63FBCA02CA" author="MATOCQ A." box="[894,1044,1494,1518]" pageId="2" pageNumber="421" pagination="69 - 101" refId="ref1544" refString="MATOCQ A. 2004: Revue des especes attribuees au genre Megalocoleus Reuter, 1890 (Heteroptera: Miridae). Annales de la Societe Entomologique de France (N. S.) 40: 69 - 101." type="journal article" year="2004">MATOCQ 2004</bibRefCitation>
).
</paragraph>
<paragraph id="865FF702E346EE1938E0FA42FF24014A" blockId="2.[151,1127,1206,1646]" pageId="2" pageNumber="421">
<taxonomicName id="41E08C81E346EE1938E0FA42FE5F012A" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[181,385,1527,1550]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE1938E0FA42FE5F012A" box="[181,385,1527,1550]" italics="true" pageId="2" pageNumber="421">Megalocoleus stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE1939DDFA43FE050129" box="[392,475,1526,1549]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
runs to couplet
<quantity id="41185AE7E346EE193AD3FA43FD62012A" box="[646,700,1526,1550]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="421" unit="in" value="12.0">12 in</quantity>
the key presented by
<bibRefCitation id="E2718AF3E346EE193BF0FA43FB92012A" author="MATOCQ A." box="[933,1100,1526,1550]" pageId="2" pageNumber="421" pagination="69 - 101" refId="ref1544" refString="MATOCQ A. 2004: Revue des especes attribuees au genre Megalocoleus Reuter, 1890 (Heteroptera: Miridae). Annales de la Societe Entomologique de France (N. S.) 40: 69 - 101." type="journal article" year="2004">MATOCQ (2004)</bibRefCitation>
. A third choice can be added at that point to accommodate
<taxonomicName id="41E08C81E346EE193ABCF9A2FCE5010A" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[745,827,1559,1582]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E346EE193ABCF9A2FCE5010A" box="[745,827,1559,1582]" italics="true" pageId="2" pageNumber="421">M. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE346EE193B17F9A3FC4B0109" box="[834,917,1558,1581]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
in the key. Asterisk (*) means that an examination of the male genitalia is necessary to identify the species with certainty.
</paragraph>
<paragraph id="865FF702E347EE183824FF69FB9F0677" blockId="3.[113,1089,219,628]" pageId="3" pageNumber="422">
12. Second segment of antenna slender, at least
<quantity id="41185AE7E347EE183A3AFF6EFD6107D7" box="[623,703,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="422" unit="mm" value="1.5">1.5 mm</quantity>
long. Female: pubescence generally pale dense and fine, dark hairs sparse. Male: pubescence brownish, flexible, not dense. Rostrum in both sexes very long, reaching or even surpassing posterior end of abdomen. Large species. ................................................................ *
<taxonomicName id="41E08C81E347EE183AAAFE89FB9F0677" authority="(Fieber, 1861)" baseAuthorityName="Fieber" baseAuthorityYear="1861" box="[767,1089,315,340]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B4942B10E347EE183AAAFE89FC7C0670" bold="true" box="[767,930,315,340]" italics="true" pageId="3" pageNumber="422">M. longirostris</emphasis>
(Fieber, 1861)
</taxonomicName>
</paragraph>
<paragraph id="865FF702E347EE183824FEE9FB9F06B0" blockId="3.[113,1089,219,628]" pageId="3" pageNumber="422">
– Second segment of antenna much shorter,
<quantity id="41185AE7E347EE183A0CFEE9FD770657" box="[601,681,348,372]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="422" unit="mm" value="1.1">1.1 mm</quantity>
long; apex of rostrum reaching middle of abdominal segment VIII. ............................................................... *
<emphasis id="B4942B10E347EE183BC5FEC9FB9F06B0" bold="true" box="[912,1089,379,404]" pageId="3" pageNumber="422">
<taxonomicName id="41E08C81E347EE183BC5FEC9FC3A06B7" authority="Matocq, 2008" authorityName="Matocq" authorityYear="2008" box="[912,996,379,403]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="stysi" status="sp. nov.">
<emphasis id="B4942B10E347EE183BC5FEC9FC3A06B7" bold="true" box="[912,996,379,403]" italics="true" pageId="3" pageNumber="422">M. stysi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFA7966BE347EE183BBEFEC9FB9F06B0" box="[1003,1089,380,404]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="865FF702E347EE183824FE29FB9F0697" blockId="3.[113,1089,219,628]" box="[113,1089,411,436]" pageId="3" pageNumber="422">
– Second segment of antenna shorter,
<quantity id="41185AE7E347EE183A75FE2EFDA10697" box="[544,639,411,435]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="422" unit="mm" value="1.25">1.25 mm</quantity>
long. ....................................................
<emphasis id="B4942B10E347EE183C73FE2EFB9F0697" bold="true" box="[1062,1089,411,435]" pageId="3" pageNumber="422">13</emphasis>
</paragraph>
<paragraph id="865FF702E347EE183824FE09FB9F0537" blockId="3.[113,1089,219,628]" pageId="3" pageNumber="422">
13. Brown pubescence dense, hairs thick; pale pubescence sparse and fine. Apex of rostrum surpassing abdominal segment VIII (male) or reaching its basal quarter (female). Corium sometimes with a round, more or less pronounced mark. ..... *
<taxonomicName id="41E08C81E347EE183B60FE49FB9F0537" authority="(Fieber, 1861)" baseAuthorityName="Fieber" baseAuthorityYear="1861" box="[821,1089,507,532]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="lunula">
<emphasis id="B4942B10E347EE183B60FE49FC7D0530" bold="true" box="[821,931,507,532]" italics="true" pageId="3" pageNumber="422">M. lunula</emphasis>
(Fieber, 1861)
</taxonomicName>
</paragraph>
<paragraph id="865FF702E347EE183824FDA9FB9F0550" blockId="3.[113,1089,219,628]" pageId="3" pageNumber="422">
– Brown and pale pubescence fine and rather dense, evenly distributed. Apex of rostrum surpassing abdominal segment VIII (male) or reaching its middle (female). Cuneus sometimes orange. ................................................................... *
<taxonomicName id="41E08C81E347EE183B13FDE9FB9F0550" authority="(Reuter, 1879)" baseAuthorityName="Reuter" baseAuthorityYear="1879" box="[838,1089,604,628]" class="Insecta" family="Miridae" genus="Megalocoleus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="naso">
<emphasis id="B4942B10E347EE183B13FDE9FC410557" bold="true" box="[838,927,604,627]" italics="true" pageId="3" pageNumber="422">M. naso</emphasis>
(Reuter, 1879)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CEFAA489E347EE183824FD3EFDD405C6" pageId="3" pageNumber="422" type="etymology">
<paragraph id="865FF702E347EE183824FD3EFDD405C6" blockId="3.[113,1089,649,739]" pageId="3" pageNumber="422">
<emphasis id="B4942B10E347EE183824FD3EFF2C0587" bold="true" box="[113,242,651,675]" pageId="3" pageNumber="422">Etymology.</emphasis>
The species is dedicated to Prof. Pavel Štys on the occasion of his 75
<superScript id="71955A4AE347EE183B80FD3CFC3F05BC" attach="left" box="[981,993,649,664]" fontSize="6" pageId="3" pageNumber="422">th</superScript>
birthday and in recognition of his important contribution to our knowledge of the
<taxonomicName id="41E08C81E347EE183B2FFD1EFC2505E6" box="[890,1019,683,706]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
.
<emphasis id="B4942B10E347EE183824FD7FFEDA05C6" bold="true" box="[113,260,714,738]" pageId="3" pageNumber="422">Distribution.</emphasis>
Northwestern
<collectingCountry id="FEF7B792E347EE1839F7FD7EFDD805C6" box="[418,518,715,738]" name="Morocco" pageId="3" pageNumber="422">Morocco</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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