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<document id="0335DB999848590409319E50C8C4895D" ID-DOI="10.11646/zootaxa.3852.1.5" ID-GBIF-Dataset="f1e56244-7924-416e-838b-c2e8cf3acdce" ID-ISSN="1175-5326" ID-Zenodo-Dep="230277" ID-ZooBank="D65F909C-A047-4A90-A6BC-36D402F8093D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460883309221" checkinUser="plazi" docAuthor="Miranda, Ivana, Schubart, Christoph D. &amp; Mantelatto, Fernando L." docDate="2014" docId="03E887D7FFE0FFA6E4A6FDA0FC1DFBAB" docLanguage="en" docName="zt03852p132.pdf" docOrigin="Zootaxa 3852 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachycheles chubutensis Boschi 1963" docType="treatment" docVersion="7" lastPageNumber="126" masterDocId="FFD1FFAFFFE6FFAEE431FF97FF9EFFD6" masterDocTitle="Morphological and molecular data support the distinctiveness of Pachycheles laevidactylus Ortmann, 1892 and Pachycheles chubutensis Boschi, 1963 (Anomura, Porcellanidae)" masterLastPageNumber="132" masterPageNumber="118" pageNumber="124" updateTime="1698358650689" updateUser="plazi">
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<mods:title id="6E99B13A829654AAB40C24B849717579">Morphological and molecular data support the distinctiveness of Pachycheles laevidactylus Ortmann, 1892 and Pachycheles chubutensis Boschi, 1963 (Anomura, Porcellanidae)</mods:title>
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<treatment id="03E887D7FFE0FFA6E4A6FDA0FC1DFBAB" ID-DOI="http://doi.org/10.5281/zenodo.6142192" ID-GBIF-Taxon="119585813" ID-Zenodo-Dep="6142192" LSID="urn:lsid:plazi:treatment:03E887D7FFE0FFA6E4A6FDA0FC1DFBAB" httpUri="http://treatment.plazi.org/id/03E887D7FFE0FFA6E4A6FDA0FC1DFBAB" lastPageId="8" lastPageNumber="126" pageId="6" pageNumber="124">
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<paragraph id="8BFE36C1FFE0FFA8E4A6FDA0FDF7FD84" blockId="6.[151,617,567,627]" box="[151,617,567,594]" pageId="6" pageNumber="124">
<heading id="D0B681ADFFE0FFA8E4A6FDA0FDF7FD84" bold="true" box="[151,617,567,594]" fontSize="11" level="1" pageId="6" pageNumber="124" reason="1">
<taxonomicName id="4C414D42FFE0FFA8E4A6FDA0FDF7FD84" ID-CoL="d8c1715e-ffb7-4d64-a35b-176871f07a2f" authority="Boschi, 1963" authorityName="Boschi" authorityYear="1963" box="[151,617,567,594]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="124" phylum="Arthropoda" rank="species" species="chubutensis">
<emphasis id="B935EAD3FFE0FFA8E4A6FDA0FDF7FD84" bold="true" box="[151,617,567,594]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4A6FDA0FE5BFD87" bold="true" box="[151,453,567,593]" italics="true" pageId="6" pageNumber="124">Pachycheles chubutensis</emphasis>
<bibRefCitation id="EFD04B30FFE0FFA8E5FDFDAFFDF7FD84" author="Boschi" box="[460,617,568,594]" pageId="6" pageNumber="124" refString="Boschi, E. E. (1963) Sobre dos especies de Pachycheles de la Argentina. Neotropica, 9, 31 - 37." type="journal article" year="1963">Boschi, 1963</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BFE36C1FFE0FFA8E4A6FDCDFEB5FDA5" blockId="6.[151,617,567,627]" box="[151,299,602,627]" pageId="6" pageNumber="124">
(
<figureCitation id="137A2A44FFE0FFA8E4AEFDCDFF72FDA5" box="[159,236,602,627]" captionStart="FIGURE 2" captionStartId="7.[151,250,1718,1740]" captionTargetBox="[160,1426,589,1696]" captionTargetId="figure@7.[151,1436,589,1697]" captionTargetPageId="7" captionText="FIGURE 2. Pachycheles chubutensis Boschi, 1963. A) General dorsal view (male, CCFurg 920); B) Minor cheliped, glabrous form (CCDB 2802, male, dorsal view); C) Telson with 7 plates (CCDB 2802, male, ventral view). Scale bar in A) = 1 cm." httpUri="https://zenodo.org/record/230279/files/figure.png" pageId="6" pageNumber="124">Figs. 2</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFE0FFA8E4A6FD0BFCEEFD39" pageId="6" pageNumber="124" type="reference_group">
<paragraph id="8BFE36C1FFE0FFA8E4A6FD0BFCEEFD39" blockId="6.[151,1436,668,751]" pageId="6" pageNumber="124">
<treatmentCitationGroup id="AB5111EFFFE0FFA8E4A6FD0BFCEEFD39" pageId="6" pageNumber="124">
<taxonomicName id="4C414D42FFE0FFA8E4A6FD0BFDD6FD64" ID-CoL="d8c1715e-ffb7-4d64-a35b-176871f07a2f" authority="Boschi, 1963: 35" authorityName="Boschi" authorityPageNumber="35" authorityYear="1963" box="[151,584,668,690]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="124" phylum="Arthropoda" rank="species" species="chubutensis">
<emphasis id="B935EAD3FFE0FFA8E4A6FD0BFE0DFD64" box="[151,403,668,690]" italics="true" pageId="6" pageNumber="124">Pachycheles chubutensis</emphasis>
<treatmentCitation id="0AE010D0FFE0FFA8E5ABFD0BFDD6FD64" author="Boschi" box="[410,584,668,690]" page="35" pageId="6" pageNumber="124" year="1963">
<bibRefCitation id="EFD04B30FFE0FFA8E5ABFD0BFDBDFD64" author="Boschi" box="[410,547,668,690]" pageId="6" pageNumber="124" refString="Boschi, E. E. (1963) Sobre dos especies de Pachycheles de la Argentina. Neotropica, 9, 31 - 37." type="journal article" year="1963">Boschi, 1963</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
, 36, figs. 2, 3; 1979: 137.—
<treatmentCitation id="0AE010D0FFE0FFA8E752FD0BFBF4FD64" author="Da" box="[867,1130,668,690]" page="133" pageId="6" pageNumber="124" year="1989">
<bibRefCitation id="EFD04B30FFE0FFA8E752FD0BFBA6FD64" author="Da" box="[867,1080,668,690]" pageId="6" pageNumber="124" refString="Da Silva, B. M. G., Braga, A. C. &amp; D´Incao, F. (1989) Porcellanidae (Decapoda, Anomura) de Santa Catarina e Rio Grande do Sul, Brasil. Iheringia, Serie Zoologia, 69, 131 - 146." type="journal article" year="1989">
Da Silva
<emphasis id="B935EAD3FFE0FFA8E7F2FD0AFC68FD64" box="[963,1014,668,690]" italics="true" pageId="6" pageNumber="124">et al.</emphasis>
, 1989
</bibRefCitation>
: 133
</treatmentCitation>
, 135.—
<treatmentCitation id="0AE010D0FFE0FFA8E087FD0BFF71FD06" author="Veloso" page="173" pageId="6" pageNumber="124" year="1993">
<bibRefCitation id="EFD04B30FFE0FFA8E087FD0BFA07FD64" author="Veloso" box="[1206,1433,668,690]" pageId="6" pageNumber="124" refString="Veloso, V. G. &amp; Melo, G. A. S. (1993) Taxonomia e distribuicao da Familia Porcellanidae (Crustacea, Decapoda, Anomura) no Litoral Brasileiro. Iheringia, Serie Zoloogia, 75, 171 - 186." type="journal article" year="1993">Veloso &amp; Melo, 1993</bibRefCitation>
: 173
</treatmentCitation>
, 177, 184.—
<treatmentCitation id="0AE010D0FFE0FFA8E55FFD2DFD35FD06" author="Harvey" box="[366,683,698,720]" page="713" pageId="6" pageNumber="124" year="1996">
<bibRefCitation id="EFD04B30FFE0FFA8E55FFD2DFDE4FD06" author="Harvey" box="[366,634,698,720]" pageId="6" pageNumber="124" refString="Harvey, A. W. &amp; De Santo, E. M. (1996) On the status of Pachycheles laevidactylus Ortmann, 1892 (Crustacea: Decapoda: Porcellanidae). Proceedings of the Biological Society of Washington, 109 (4), 707 - 714." type="journal article" year="1996">
Harvey &amp;
<collectingCountry id="F3567651FFE0FFA8E5EAFD2CFE67FD06" box="[475,505,699,720]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="124">De</collectingCountry>
Santo, 1996
</bibRefCitation>
: 713
</treatmentCitation>
.—
<treatmentCitation id="0AE010D0FFE0FFA8E6FBFD2DFCEAFD06" author="Melo" box="[714,884,698,720]" page="227" pageId="6" pageNumber="124" year="1999">
<bibRefCitation id="EFD04B30FFE0FFA8E6FBFD2DFCDFFD06" author="Melo" box="[714,833,698,720]" pageId="6" pageNumber="124" refString="Melo, G. A. S. (1999) Manual de identificacao dos Crustacea Decapoda do litoral brasileiro: Anomura, Thalassinidea, Palinuridea, Astacidea. Pleidae, Sao Paulo, Brazil, 551 pp." type="book" year="1999">Melo, 1999</bibRefCitation>
: 227
</treatmentCitation>
, 232, fig. 154.—
<treatmentCitation id="0AE010D0FFE0FFA8E02BFD2DFAADFD06" author="Rodriguez" box="[1050,1331,698,720]" page="554" pageId="6" pageNumber="124" year="2005">
<bibRefCitation id="EFD04B30FFE0FFA8E02BFD2DFA9CFD06" author="Rodriguez" box="[1050,1282,698,720]" pageId="6" pageNumber="124" refString="Rodriguez, I. T., Hernandez, G. &amp; Felder, D. L. (2005) Review of western Atlantic Porcellanidae with new records, systematic observations, and comments on biogeography. Caribbean Journal of Science, 41 (3), 544 - 582." type="journal article" year="2005">
Rodríguez
<emphasis id="B935EAD3FFE0FFA8E0BCFD2CFB5EFD06" box="[1165,1216,698,720]" italics="true" pageId="6" pageNumber="124">et al.</emphasis>
, 2005
</bibRefCitation>
: 554
</treatmentCitation>
, 555, 570, 572, 574.—
<treatmentCitation id="0AE010D0FFE0FFA8E50CFD4EFE6BFD39" author="Ferreira" box="[317,501,729,751]" page="59" pageId="6" pageNumber="124" year="2009">
<bibRefCitation id="EFD04B30FFE0FFA8E50CFD4EFE4CFD39" author="Ferreira" box="[317,466,729,751]" pageId="6" pageNumber="124" refString="Ferreira, L. A. A. (2009) Six additional records of porcellain crabs from Brazil (Anomura: Porcellanidae). Nauplius, 17 (1), 59 - 60." type="journal article" year="2009">Ferreira, 2009</bibRefCitation>
: 59
</treatmentCitation>
.—
<treatmentCitation id="0AE010D0FFE0FFA8E622FD4EFCF2FD39" author="Osawa" box="[531,876,729,751]" page="112" pageId="6" pageNumber="124" year="2010">
<bibRefCitation id="EFD04B30FFE0FFA8E622FD4EFCA5FD39" author="Osawa" box="[531,827,729,751]" pageId="6" pageNumber="124" refString="Osawa, M. &amp; McLaughlin, P. A. (2010) Annotated checklist of Anomuran Decapod Crustacean (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of Galatheoidea) Part II - Porcellanidae. The Raffles Bulletin of Zoology, 23, 109 - 129." type="journal article" year="2010">Osawa &amp; McLaughlin, 2010</bibRefCitation>
: 112
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFE0FFA8E4A6FC82FBADFBA4" pageId="6" pageNumber="124" type="materials_examined">
<paragraph id="8BFE36C1FFE0FFA8E4A6FC82FD50FCA0" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4A6FC82FEA8FCF8" bold="true" box="[151,310,789,814]" pageId="6" pageNumber="124">
<typeStatus id="54FA8863FFE0FFA8E4A6FC82FF4CFCF8" box="[151,210,789,814]" pageId="6" pageNumber="124">Type</typeStatus>
locality.
</emphasis>
Golfo San Matías, Chubut,
<collectingCountry id="F3567651FFE0FFA8E642FC82FD78FCF8" box="[627,742,789,814]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
. The
<typeStatus id="54FA8863FFE0FFA8E714FC82FC19FCF8" box="[805,903,789,814]" pageId="6" pageNumber="124" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C414D42FFE0FFA8E79CFC82FB5AFCF8" box="[941,1220,789,814]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="124" phylum="Arthropoda" rank="species" species="chubutensis">
<emphasis id="B935EAD3FFE0FFA8E79CFC82FB5AFCF8" box="[941,1220,789,814]" italics="true" pageId="6" pageNumber="124">Pachycheles chubutensis</emphasis>
</taxonomicName>
(male with
<quantity id="4CB99B24FFE0FFA8E17CFC81FA02FCFB" box="[1357,1436,790,814]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="124" unit="mm" value="10.0">10 mm</quantity>
carapace length) is in the Museo
<collectingCountry id="F3567651FFE0FFA8E62AFCAEFD12FC84" box="[539,652,825,850]" name="Argentina" pageId="6" pageNumber="124">Argentino</collectingCountry>
de Ciencias Naturales (Buenos Aires,
<collectingCountry id="F3567651FFE0FFA8E07EFCAEFB5EFC84" box="[1103,1216,825,850]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
) (Col. Museo Arg. Cienc. No. 21.244) (
<bibRefCitation id="EFD04B30FFE0FFA8E54AFCCAFD8BFCA0" author="Boschi" box="[379,533,861,886]" pageId="6" pageNumber="124" refString="Boschi, E. E. (1963) Sobre dos especies de Pachycheles de la Argentina. Neotropica, 9, 31 - 37." type="journal article" year="1963">Boschi, 1963</bibRefCitation>
). Not examined.
</paragraph>
<paragraph id="8BFE36C1FFE0FFA8E4F6FC16FBADFBA4" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6FC16FE2BFC4F" bold="true" box="[199,437,897,922]" pageId="6" pageNumber="124">Material examined.</emphasis>
<specimenCount id="9D47FD48FFE0FFA8E5F0FC15FDB5FC4C" box="[449,555,897,922]" pageId="6" pageNumber="124" type="female">1 female</specimenCount>
,
<specimenCount id="9D47FD48FFE0FFA8E60AFC15FD15FC4C" box="[571,651,897,922]" pageId="6" pageNumber="124" type="male">1 male</specimenCount>
(CCFurg 381), Porto Belo, Santa Catarina,
<collectingCountry id="F3567651FFE0FFA8E0ABFC16FB7FFC4C" box="[1178,1249,897,922]" name="Brazil" pageId="6" pageNumber="124">Brazil</collectingCountry>
, coll. Silva, B.,
<date id="FFFF1001FFE0FFA8E4A6FC31FEB4FC68" box="[151,298,934,958]" pageId="6" pageNumber="124" value="1987-03-23">23.Mar.1987</date>
;
<specimenCount id="9D47FD48FFE0FFA8E505FC31FE14FC68" box="[308,394,933,958]" pageId="6" pageNumber="124" type="male">2 males</specimenCount>
(CCFurg 370), Rio Grande do Sul (RS),
<collectingCountry id="F3567651FFE0FFA8E765FC32FC05FC68" box="[852,923,933,958]" name="Brazil" pageId="6" pageNumber="124">Brazil</collectingCountry>
, coll. Silva, B.,
<date id="FFFF1001FFE0FFA8E07DFC32FB46FC68" box="[1100,1240,933,958]" pageId="6" pageNumber="124" value="1986-06-27">27.Jun.1986</date>
;
<specimenCount id="9D47FD48FFE0FFA8E0D2FC31FAD9FC68" box="[1251,1351,933,958]" pageId="6" pageNumber="124" type="female">1 female</specimenCount>
,
<specimenCount id="9D47FD48FFE0FFA8E160FC31FA02FC68" box="[1361,1436,933,958]" pageId="6" pageNumber="124" type="male">1 male</specimenCount>
(CCFurg 920), Parcel do Carpinteiro, RS,
<collectingCountry id="F3567651FFE0FFA8E645FC5EFD25FC34" box="[628,699,969,994]" name="Brazil" pageId="6" pageNumber="124">Brazil</collectingCountry>
, coll. Feijó, F.A.,
<date id="FFFF1001FFE0FFA8E7B7FC5EFC76FC34" box="[902,1000,969,994]" pageId="6" pageNumber="124" value="1988-07">Jul.1988</date>
;
<specimenCount id="9D47FD48FFE0FFA8E7C4FC5DFBC8FC34" box="[1013,1110,969,994]" pageId="6" pageNumber="124" type="female">1 female</specimenCount>
(
<collectionCode id="ED50AE04FFE0FFA8E057FC5DFB2AFC34" box="[1126,1204,970,994]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="124">CCDB</collectionCode>
860), Mar del Plata,
<collectingCountry id="F3567651FFE0FFA8E4A6FC7AFE97FBD0" box="[151,265,1005,1030]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
, coll. Ocampo, E.,
<date id="FFFF1001FFE0FFA8E5D6FC7AFDE8FBD0" box="[487,630,1005,1030]" pageId="6" pageNumber="124" value="2003-02-14">14.Feb.2003</date>
;
<specimenCount id="9D47FD48FFE0FFA8E6B2FC79FD6FFBD0" box="[643,753,1005,1030]" pageId="6" pageNumber="124" type="female">2 females</specimenCount>
(
<collectionCode id="ED50AE04FFE0FFA8E730FC79FCD1FBD0" box="[769,847,1006,1030]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="124">CCDB</collectionCode>
2800), La Restinga, Mar del Plata,
<collectingCountry id="F3567651FFE0FFA8E0DDFC7AFAC0FBD0" box="[1260,1374,1005,1030]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
, coll. Ocampo, E.,
<date id="FFFF1001FFE0FFA8E505FB85FE59FBFC" box="[308,455,1042,1066]" pageId="6" pageNumber="124" value="2010-03-22">22.Mar.2010</date>
;
<specimenCount id="9D47FD48FFE0FFA8E5E6FB85FDA3FBFC" box="[471,573,1041,1066]" pageId="6" pageNumber="124" type="female">1 female</specimenCount>
(
<collectionCode id="ED50AE04FFE0FFA8E660FB85FD01FBFC" box="[593,671,1042,1066]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="124">CCDB</collectionCode>
2801), La Restinga, Mar del Plata,
<collectingCountry id="F3567651FFE0FFA8E063FB86FB5AFBFC" box="[1106,1220,1041,1066]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
, coll. E. Ocampo,
<date id="FFFF1001FFE0FFA8E4A6FBA1FE9CFB98" box="[151,258,1078,1102]" pageId="6" pageNumber="124" value="2008-10">Oct.2008</date>
;
<specimenCount id="9D47FD48FFE0FFA8E53EFBA1FEC2FB98" box="[271,348,1077,1102]" pageId="6" pageNumber="124" type="male">1 male</specimenCount>
(
<collectionCode id="ED50AE04FFE0FFA8E55CFBA1FE25FB98" box="[365,443,1078,1102]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="124">CCDB</collectionCode>
2802), La Restinga, Mar del Plata,
<collectingCountry id="F3567651FFE0FFA8E768FBA2FC55FB98" box="[857,971,1077,1102]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
, coll. Ocampo, E.,
<date id="FFFF1001FFE0FFA8E09BFBA1FADCFB98" box="[1194,1346,1078,1102]" pageId="6" pageNumber="124" value="2008-05-22">22.May.2008</date>
;
<specimenCount id="9D47FD48FFE0FFA8E17EFBA1FA02FB98" box="[1359,1436,1077,1102]" pageId="6" pageNumber="124" type="male">1 male</specimenCount>
(
<collectionCode id="ED50AE04FFE0FFA8E4AFFBCDFF72FBA4" box="[158,236,1114,1138]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="124">CCDB</collectionCode>
2804), La Restinga, Mar del Plata,
<collectingCountry id="F3567651FFE0FFA8E64DFBCEFD70FBA4" box="[636,750,1113,1138]" name="Argentina" pageId="6" pageNumber="124">Argentina</collectingCountry>
, coll. Ocampo, E.,
<date id="FFFF1001FFE0FFA8E7F5FBCDFBAEFBA4" box="[964,1072,1114,1138]" pageId="6" pageNumber="124" value="2008-10">Oct.2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFE0FFA8E4F6FBEAFF7EFB08" pageId="6" pageNumber="124" type="diagnosis">
<paragraph id="8BFE36C1FFE0FFA8E4F6FBEAFF7EFB08" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6FBEAFEDFFB43" bold="true" box="[199,321,1149,1174]" pageId="6" pageNumber="124">Diagnosis.</emphasis>
Frontal region with tuft of plumose setae. Chelipeds without prominent granules, external surface with plumose setae and some simple setae on margins.
<taxonomicName id="4C414D42FFE0FFA8E721FB36FCC4FB6C" box="[784,858,1185,1210]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
consisting of 7 plates. Second pleopods present in males.
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFE0FFA6E4F6FB7EFC85FE06" lastPageId="8" lastPageNumber="126" pageId="6" pageNumber="124" type="description">
<paragraph id="8BFE36C1FFE0FFA8E4F6FB7EFC7AFA60" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6FB7EFEEAFAD7" bold="true" box="[199,372,1257,1282]" pageId="6" pageNumber="124">Redescription.</emphasis>
<emphasis id="B935EAD3FFE0FFA8E54FFB7DFE6AFAD7" box="[382,500,1258,1281]" italics="true" pageId="6" pageNumber="124">Carapace.</emphasis>
Carapace slightly longer than broad; surface of branchial and postlateral regions wrinkled, more prominent in some specimens. Tufts of short and simple setae present on metagastric region of some specimens. Front trilobed in frontal view, with medium lobe more prominent in some specimens; a dense tuft of plumose setae present, smaller tufts can be found on hepatic region. Orbital region well defined, continuing on a curved crest that constitutes an accentuated angle with the lateral carapace borders. Lateral walls divided into a large piece followed by one or more (two to three) smaller posterior pieces.
</paragraph>
<paragraph id="8BFE36C1FFE0FFA8E4F6FA56FCF1FA28" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6FA56FEDEFA0C" box="[199,320,1473,1498]" italics="true" pageId="6" pageNumber="124">Antennule.</emphasis>
Basal segment of antennule armed with 3–5 strong spines on medial anterior margin (visible from dorsal-lateral view), 7 or 8 spines on anterolateral dorsal surface.
</paragraph>
<paragraph id="8BFE36C1FFE0FFA8E4F6F99DFEA4F958" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6F99DFEB2F9F7" box="[199,300,1546,1569]" italics="true" pageId="6" pageNumber="124">Antenna.</emphasis>
Basal segment of antenna with projection on the anterior margin, configuring a well defined cavity for the eyes together with the border of orbits; second segment with a small protuberance on the anterior margin; third segment smooth to slightly granulose on anterior margin; flagellum with tiny bristles just visible under great magnification.
</paragraph>
<paragraph id="8BFE36C1FFE0FFA8E4F6F90EFD08F820" blockId="6.[151,1437,789,2038]" pageId="6" pageNumber="124">
<emphasis id="B935EAD3FFE0FFA8E4F6F90EFE7CF964" box="[199,482,1689,1714]" italics="true" pageId="6" pageNumber="124">Pereiopod 1 (Chelipeds).</emphasis>
Chelipeds unequal, strong, with a thin and weak granulation, more intense from the medial region to the external margin of carpus, propodus, and dactylus. Merus with rounded projection on inner margin; presence of parallel series of small bristles following wrinkles in inner-outer border direction; dorsal surface of carpus, propodus and dactylus sometimes covered by tufts of plumose setae originating from base of granules, alternating with long and simple setae, the last ones especially on external borders. Density and distribution of these tufts is variable among individuals from different localities. Anterior margin of carpus armed with projecting lobe subdivided into 2–4 lobes of variable sizes; distal ones being better delimited. Inner border of pollex of major cheliped with 1–4 large tubercles; lines of short bristles present; dense tuft of plumose setae at the base of pollex filling the gap between fingers. Internal margin of dactylus with 7 or 8 small tubercles; both fingers of minor cheliped with a cutting inner border.
</paragraph>
<paragraph id="8BFE36C1FFE1FFA9E4F6FF00FB07FE06" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6FF00FEE1FF66" box="[199,383,151,176]" italics="true" pageId="7" pageNumber="125">Pereiopods 2–4.</emphasis>
Merus and carpus of first pereiopods slightly granulated, covered by wide plumose setae on dorsal surface; propodus with wide lines of plumose setae on dorsal and lateral margins, ventral surface with 3–5 spines, 2 on distal border, 1 subdistal, shaping a triangle with the first 2 and 1 or 2 other medial spines located proximally; dactylus ending in a strong spine, inner border with 4 or 5 spines decreasing in size. Second pereiopod with the same pattern of setae distribution for merus, carpus and propodus, more dense when compared to the first pair; ventral surface of propodus with three to five spines, two at distal border, followed by one medial and immediately above, forming triangle with first 2, and 1 or 2 medial, located proximally; dactylus ending in a strong claw; inner border with 4 spines decreasing in size. Third pereiopod equal to the second. Fourth pereiopod smooth, carpus with rolls of long and simple setae on dorsal, ventral and lateral surfaces, brush-like.
</paragraph>
<paragraph id="8BFE36C1FFE1FFA9E4F6FE4BFD5AFE23" blockId="7.[151,1436,151,536]" box="[199,708,476,501]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6FE4BFE8AFE22" box="[199,276,476,501]" italics="true" pageId="7" pageNumber="125">
<taxonomicName id="4C414D42FFE1FFA9E4F6FE4BFE93FE23" box="[199,269,476,501]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="7" pageNumber="125" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
.
</emphasis>
<taxonomicName id="4C414D42FFE1FFA9E52CFE4BFEF9FE23" box="[285,359,476,501]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="7" pageNumber="125" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
7-plated in males and females.
</paragraph>
<paragraph id="8BFE36C1FFE1FFA9E4F6FE68FD2BFDCE" blockId="7.[151,1436,151,536]" box="[199,693,511,536]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6FE68FEA9FDCE" box="[199,311,511,536]" italics="true" pageId="7" pageNumber="125">Pleopods.</emphasis>
Second pleopod present in males.
</paragraph>
<caption id="DF3E6649FFE1FFA9E4A6F921FAF5F93C" httpUri="https://zenodo.org/record/230279/files/figure.png" pageId="7" pageNumber="125" targetBox="[160,1426,589,1696]" targetPageId="7">
<paragraph id="8BFE36C1FFE1FFA9E4A6F921FAF5F93C" blockId="7.[151,1436,1718,1771]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4A6F921FE93F91D" bold="true" box="[151,269,1718,1740]" pageId="7" pageNumber="125">FIGURE 2</emphasis>
.
<taxonomicName id="4C414D42FFE1FFA9E528F921FD3BF91A" authority="Boschi, 1963" authorityName="Boschi" authorityYear="1963" box="[281,677,1718,1740]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="125" phylum="Arthropoda" rank="species" species="chubutensis">
<emphasis id="B935EAD3FFE1FFA9E528F921FD88F91A" box="[281,534,1718,1740]" italics="true" pageId="7" pageNumber="125">Pachycheles chubutensis</emphasis>
Boschi, 1963
</taxonomicName>
. A) General dorsal view (male, CCFurg 920); B) Minor cheliped, glabrous form (CCDB 2802, male, dorsal view); C)
<taxonomicName id="4C414D42FFE1FFA9E67BF943FD13F93C" box="[586,653,1748,1770]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="7" pageNumber="125" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
with 7 plates (CCDB 2802, male, ventral view). Scale bar in A) = 1 cm.
</paragraph>
</caption>
<paragraph id="8BFE36C1FFE1FFA9E4F6F885FBCBF8FC" blockId="7.[151,1436,1810,2015]" box="[199,1109,1810,1835]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6F885FEE3F8FD" bold="true" box="[199,381,1810,1835]" pageId="7" pageNumber="125">Measurements.</emphasis>
Males with carapace width 8.0–12.0 mm, females
<quantity id="4CB99B24FFE1FFA9E788F885FBCFF8FC" box="[953,1105,1810,1834]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.24" metricValueMin="0.8099999999999999" pageId="7" pageNumber="125" unit="mm" value="10.25" valueMax="12.4" valueMin="8.1">8.1–12.4 mm</quantity>
.
</paragraph>
<paragraph id="8BFE36C1FFE1FFA9E4F6F8A1FCE5F8A5" blockId="7.[151,1436,1810,2015]" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6F8A1FE8FF898" bold="true" box="[199,273,1846,1871]" pageId="7" pageNumber="125">Color.</emphasis>
All the material analyzed was preserved in alcohol or formalin, being impossible to report the color in life. There is no previous information to be found in the literature.
</paragraph>
<paragraph id="8BFE36C1FFE1FFA6E4F6F8E9FC85FE06" blockId="7.[151,1436,1810,2015]" lastBlockId="8.[151,1437,151,1149]" lastPageId="8" lastPageNumber="126" pageId="7" pageNumber="125">
<emphasis id="B935EAD3FFE1FFA9E4F6F8E9FEB5F840" bold="true" box="[199,299,1918,1943]" pageId="7" pageNumber="125">Ecology.</emphasis>
<taxonomicName id="4C414D42FFE1FFA9E505F8E9FDD2F841" box="[308,588,1918,1943]" class="Malacostraca" family="Porcellanidae" genus="Pachycheles" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="125" phylum="Arthropoda" rank="species" species="chubutensis">
<emphasis id="B935EAD3FFE1FFA9E505F8E9FDD2F841" box="[308,588,1918,1943]" italics="true" pageId="7" pageNumber="125">Pachycheles chubutensis</emphasis>
</taxonomicName>
occurs infralittorally, preferentially on hard, mussel-free bottoms (
<bibRefCitation id="EFD04B30FFE1FFA9E172F8E8FEF1F86C" author="Bremec" pageId="7" pageNumber="125" refString="Bremec, C. S. &amp; Cazzaniga, N. J. (1984) Consideraciones sobre Pachycheles haigae Rodrigues da Costa, 1960 y P. chubutensis Boschi, 1963 en Monte Hermoso (Republica Argentina) (Crustacea, Anomura, Porcellanidae). Iheringia, Serie Zoologia, 64, 149 - 162." type="journal article" year="1984">Bremec &amp; Cassaniga 1984</bibRefCitation>
;
<bibRefCitation id="EFD04B30FFE1FFA9E54FF835FD5CF86C" author="Gonzalez-Pisani" box="[382,706,1954,1979]" pageId="7" pageNumber="125" refString="Gonzalez-Pisani, X., Rubilar, T. &amp; Dupre, E. (2009) Embryonic development of Pachycheles chubutensis (Decapoda, Anomura). Journal of the Marine Biological Association of the United Kingdom, 89 (6), 1 - 8. http: // dx. doi. org / 10.1017 / s 0025315409000678" type="journal article" year="2009">
González-Pisani
<emphasis id="B935EAD3FFE1FFA9E671F834FDE2F86D" box="[576,636,1954,1979]" italics="true" pageId="7" pageNumber="125">et al.</emphasis>
2009
</bibRefCitation>
), associated to tubes of
<emphasis id="B935EAD3FFE1FFA9E7EEF835FB58F86C" box="[991,1222,1954,1978]" italics="true" pageId="7" pageNumber="125">Litofaga patagonica</emphasis>
(d'Orbigny, 1846) (
<bibRefCitation id="EFD04B30FFE1FFA9E4AFF851FE40F809" author="Gonzalez-Pisani" box="[158,478,1990,2015]" pageId="7" pageNumber="125" refString="Gonzalez-Pisani, X., Ward, C. P. &amp; Dupre, E. (2006) Descripcion del primer estadio larvario de Pachycheles chubutensis (Boschi, 1963) (Crustacea, Decapoda, Anomura) en Patagonia. Investigaciones Marinas, 34 (2), 191 - 196. http: // dx. doi. org / 10.4067 / s 0717 - 71782006000200020" type="journal article" year="2006">
González-Pisani
<emphasis id="B935EAD3FFE1FFA9E550F850FE02F809" box="[353,412,1990,2015]" italics="true" pageId="7" pageNumber="125">et al.</emphasis>
2006
</bibRefCitation>
) and also on sand bottoms at depths between
<quantity id="4CB99B24FFE1FFA9E7C5F850FBFBF808" box="[1012,1125,1991,2015]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.8" metricValueMin="0.3" pageId="7" pageNumber="125" unit="m" value="15.5" valueMax="28.0" valueMin="3.0">3 to 28 m</quantity>
(
<bibRefCitation id="EFD04B30FFE1FFA9E047F851FA94F809" author="Boschi" box="[1142,1290,1990,2015]" pageId="7" pageNumber="125" refString="Boschi, E. E. (1963) Sobre dos especies de Pachycheles de la Argentina. Neotropica, 9, 31 - 37." type="journal article" year="1963">Boschi 1963</bibRefCitation>
;
<bibRefCitation id="EFD04B30FFE1FFA9E129F851FA09F809" author="Melo" box="[1304,1431,1990,2015]" pageId="7" pageNumber="125" refString="Melo, G. A. S. (1999) Manual de identificacao dos Crustacea Decapoda do litoral brasileiro: Anomura, Thalassinidea, Palinuridea, Astacidea. Pleidae, Sao Paulo, Brazil, 551 pp." type="book" year="1999">Melo 1999</bibRefCitation>
;
<bibRefCitation id="EFD04B30FFEEFFA6E4A6FF00FEFAFF66" author="Scelzo" box="[151,356,151,176]" pageId="8" pageNumber="126" refString="Scelzo, M. A., Farias, N., Ocampo, E., Nunez, J. D. &amp; Cledon, Y. M. (2008) New extension range of the porcelain crabs Pachycheles laevidactylus and new localities for P. chubutensis, in marine Waters of Argentina Southwestern Atlantic Ocean. Abstracts of V Brazilian Crustacean Congress (CBC), Gramado, RS, Brazil, pp. 119." type="book chapter" year="2008">
Scelzo
<emphasis id="B935EAD3FFEEFFA6E4D9FF0EFEBFFF66" box="[232,289,151,176]" italics="true" pageId="8" pageNumber="126">et al.</emphasis>
2008
</bibRefCitation>
).
<bibRefCitation id="EFD04B30FFEEFFA6E547FF0FFD2AFF66" author="Bremec" box="[374,692,151,176]" pageId="8" pageNumber="126" refString="Bremec, C. S. &amp; Cazzaniga, N. J. (1984) Consideraciones sobre Pachycheles haigae Rodrigues da Costa, 1960 y P. chubutensis Boschi, 1963 en Monte Hermoso (Republica Argentina) (Crustacea, Anomura, Porcellanidae). Iheringia, Serie Zoologia, 64, 149 - 162." type="journal article" year="1984">Bremec &amp; Cassaniga (1984)</bibRefCitation>
have recorded the species occurring at the inner zone of the Bahía Blanca Estuary, a region that suffers considerable variations of salinity and temperature throughout the year. Despite their importance in the bioenergetics marine cycle, as an adult in the benthos, and as larvae in the water column, information about their biology and population ecology is scarce. Larvae are an important component of the zooplankton in the embayment of Chubut Province (
<bibRefCitation id="EFD04B30FFEEFFA6E727FEB0FBA1FE96" author="Dellatorre" box="[790,1087,295,320]" pageId="8" pageNumber="126" refString="Dellatorre, F. G. &amp; Baron, P. J. (2007) Megalopae and juvenile decapod settlement on artificial substrata in Northern Patagonia: seasonal and interannual variations. In: Book of Abstracts. The Crustacean Society Mid-Year Meeting, 2007, La Serena, Chile, pp. 87." type="book chapter" year="2007">Dellatorre &amp; Baron, 2007</bibRefCitation>
), while as an adult it is preyed upon by cephalopods (
<bibRefCitation id="EFD04B30FFEEFFA6E5A0FEDBFD56FEB2" author="Gonzalez-Pisani" box="[401,712,332,357]" pageId="8" pageNumber="126" refString="Gonzalez-Pisani, X., Rubilar, T. &amp; Dupre, E. (2009) Embryonic development of Pachycheles chubutensis (Decapoda, Anomura). Journal of the Marine Biological Association of the United Kingdom, 89 (6), 1 - 8. http: // dx. doi. org / 10.1017 / s 0025315409000678" type="journal article" year="2009">
González-Pisani
<emphasis id="B935EAD3FFEEFFA6E660FEDAFD16FEB3" box="[593,648,332,357]" italics="true" pageId="8" pageNumber="126">et al.</emphasis>
2009
</bibRefCitation>
) and some benthic fish (
<bibRefCitation id="EFD04B30FFEEFFA6E7E6FEDBFA8EFEB3" author="Gonzalez-Pisani" box="[983,1296,332,357]" pageId="8" pageNumber="126" refString="Gonzalez-Pisani, X., Ward, C. P. &amp; Dupre, E. (2006) Descripcion del primer estadio larvario de Pachycheles chubutensis (Boschi, 1963) (Crustacea, Decapoda, Anomura) en Patagonia. Investigaciones Marinas, 34 (2), 191 - 196. http: // dx. doi. org / 10.4067 / s 0717 - 71782006000200020" type="journal article" year="2006">
González-Pisani
<emphasis id="B935EAD3FFEEFFA6E0A7FEDAFB50FEB3" box="[1174,1230,332,357]" italics="true" pageId="8" pageNumber="126">et al.</emphasis>
2006
</bibRefCitation>
), integrating the trophic networks. Observations provided by
<bibRefCitation id="EFD04B30FFEEFFA6E6F0FEE7FB96FE5E" author="Bremec" box="[705,1032,367,392]" pageId="8" pageNumber="126" refString="Bremec, C. S. &amp; Cazzaniga, N. J. (1984) Consideraciones sobre Pachycheles haigae Rodrigues da Costa, 1960 y P. chubutensis Boschi, 1963 en Monte Hermoso (Republica Argentina) (Crustacea, Anomura, Porcellanidae). Iheringia, Serie Zoologia, 64, 149 - 162." type="journal article" year="1984">Bremec &amp; Cazzaniga (1984)</bibRefCitation>
suggest an annual life cycle of the species in
<collectingCountry id="F3567651FFEEFFA6E53CFE03FEE1FE7B" box="[269,383,404,429]" name="Argentina" pageId="8" pageNumber="126">Argentina</collectingCountry>
, with recruitment season in summer (September to March), when it can be found abundantly in the lower intertidal zone (
<bibRefCitation id="EFD04B30FFEEFFA6E5E2FE20FC91FE06" author="Gonzalez-Pisani" box="[467,783,439,464]" pageId="8" pageNumber="126" refString="Gonzalez-Pisani, X., Ward, C. P. &amp; Dupre, E. (2006) Descripcion del primer estadio larvario de Pachycheles chubutensis (Boschi, 1963) (Crustacea, Decapoda, Anomura) en Patagonia. Investigaciones Marinas, 34 (2), 191 - 196. http: // dx. doi. org / 10.4067 / s 0717 - 71782006000200020" type="journal article" year="2006">
González-Pisani
<emphasis id="B935EAD3FFEEFFA6E6A2FE2EFD52FE06" box="[659,716,439,464]" italics="true" pageId="8" pageNumber="126">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFEEFFA6E4F6FE4BFCD6FBE2" pageId="8" pageNumber="126" type="discussion">
<paragraph id="8BFE36C1FFEEFFA6E4F6FE4BFBB1FDB6" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="126">
<emphasis id="B935EAD3FFEEFFA6E4F6FE4BFEA5FE22" bold="true" box="[199,315,476,501]" pageId="8" pageNumber="126">Remarks.</emphasis>
Some differences were found in relation to the original text of the description and they were here incorporated. In the original description, the author mentioned the carapace as slightly broader than long, especially in males. This revision could not find any specimen fitting this proportion. The description of the basal antennule segment is presented here and also the description of the antenna is more detailed.
</paragraph>
<paragraph id="8BFE36C1FFEEFFA6E4F6FDFBFCD6FBE2" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="126">
The morphology of the lateral walls is variable, but in most cases it follows the original description. One exception is a specimen (CCDB 2801) which presented four smaller pieces after the most anterior and larger one. Presence, quantity and distribution of the setae on the cheliped surface is extremely variable, as cited firstly by
<bibRefCitation id="EFD04B30FFEEFFA6E4A6FD4FFE4FFD26" box="[151,465,727,752]" pageId="8" pageNumber="126" refString="Harvey, A. W. &amp; De Santo, E. M. (1996) On the status of Pachycheles laevidactylus Ortmann, 1892 (Crustacea: Decapoda: Porcellanidae). Proceedings of the Biological Society of Washington, 109 (4), 707 - 714." type="journal article">
Harvey &amp;
<collectingCountry id="F3567651FFEEFFA6E520FD4FFEACFD39" box="[273,306,728,751]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="126">De</collectingCountry>
Santo (1996)
</bibRefCitation>
, from less and short setae more concentrated on the outer margins of merus, carpus, propodus and dactylus to a dense coverage on these segments. All the material from
<collectingCountry id="F3567651FFEEFFA6E069FD6BFB02FCC3" box="[1112,1180,764,789]" name="Brazil" pageId="8" pageNumber="126">Brazil</collectingCountry>
had the whole surface of chelipeds covered by short and plumose setae together with long and simple setae, especially on external margins. In
<collectingCountry id="F3567651FFEEFFA6E52EFCD3FE0FFC8B" box="[287,401,836,861]" name="Argentina" pageId="8" pageNumber="126">Argentina</collectingCountry>
, we found animals with fewer and shorter setae distributed on the wrinkles and base of the granules. The number of tubercles on the inner border of pollex (major cheliped) ranged from one to four instead of three to four, being one unique medial tubercle very frequent. We extend here the range for the number of spines on propodus and dactylus of first pereiopods to three to five instead of four, as cited in the original description. Descriptions of the second, third and fourth pereiopods were added. The original description mentioned females, in general having a five-plated
<taxonomicName id="4C414D42FFEEFFA6E5EEFC60FDBBFBC6" box="[479,549,1015,1040]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="8" pageNumber="125" phylum="Arthropoda" rank="genus">telson</taxonomicName>
, sometimes with the small posterior plates half segmented; the first condition was not observed by us, while the second was observed once.
</paragraph>
</subSubSection>
<subSubSection id="C35B654AFFEEFFA6E4F6FBA8FC1DFBAB" pageId="8" pageNumber="126" type="distribution">
<paragraph id="8BFE36C1FFEEFFA6E4F6FBA8FC1DFBAB" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="126">
<emphasis id="B935EAD3FFEEFFA6E4F6FBA8FEFEFB8E" bold="true" box="[199,352,1087,1112]" pageId="8" pageNumber="126">Distribution.</emphasis>
<collectingCountry id="F3567651FFEEFFA6E556FBA8FE35FB8E" box="[359,427,1087,1112]" name="Brazil" pageId="8" pageNumber="126">Brazil</collectingCountry>
on Rio de Janeiro (
<bibRefCitation id="EFD04B30FFEEFFA6E6B7FBA8FCBAFB8E" author="Ferreira" box="[646,804,1087,1112]" pageId="8" pageNumber="126" refString="Ferreira, L. A. A. (2009) Six additional records of porcellain crabs from Brazil (Anomura: Porcellanidae). Nauplius, 17 (1), 59 - 60." type="journal article" year="2009">Ferreira 2009</bibRefCitation>
), from Santa Catarina to Rio Grande do Sul (
<bibRefCitation id="EFD04B30FFEEFFA6E12FFBD7FF6AFBAA" author="Da" pageId="8" pageNumber="126" refString="Da Silva, B. M. G., Braga, A. C. &amp; D´Incao, F. (1989) Porcellanidae (Decapoda, Anomura) de Santa Catarina e Rio Grande do Sul, Brasil. Iheringia, Serie Zoologia, 69, 131 - 146." type="journal article" year="1989">
Da Silva
<emphasis id="B935EAD3FFEEFFA6E1B9FBD6FF2AFBAB" italics="true" pageId="8" pageNumber="126">et al.</emphasis>
1989
</bibRefCitation>
) and Rawson, Chubut Province,
<collectingCountry id="F3567651FFEEFFA6E657FBF3FD4BFBAB" box="[614,725,1124,1149]" name="Argentina" pageId="8" pageNumber="126">Argentina</collectingCountry>
(
<bibRefCitation id="EFD04B30FFEEFFA6E6D5FBF3FCE8FBAB" author="Boschi" box="[740,886,1124,1149]" pageId="8" pageNumber="126" refString="Boschi, E. E. (1963) Sobre dos especies de Pachycheles de la Argentina. Neotropica, 9, 31 - 37." type="journal article" year="1963">Boschi 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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