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<document id="99CF02B508779CBDEFC9690FE582DBF2" ID-CLB-Dataset="31516" ID-DOI="10.11646/zootaxa.4079.2.8" ID-GBIF-Dataset="fe597a14-c52d-475b-b151-d9fda89fd971" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050736" ID-ZooBank="6AEDACFA-2506-4521-984B-A222E661B480" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811058477" checkinUser="plazi" docAuthor="Santiago-Jiménez, Quiyari J." docDate="2016" docId="03E887CA32134176FF3E32C769C4F901" docLanguage="en" docName="zootaxa.4079.2.8.pdf" docOrigin="Zootaxa 4079 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrozoque hoplandrioides Santiago-Jiménez 2016, sp. nov." docType="treatment" docVersion="7" masterDocId="FFD1FFB232104170FFA934666A3DFFF7" masterDocTitle="Macrozoque hoplandrioides (Coleoptera: Staphylinidae: Aleocharinae), a new genus and species of false Lomechusini from México" updateTime="1698492248874" updateUser="ExternalLinkService">
<mods:mods id="9733419C34BB6AABEC9F7596CB5F2533" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="714AA26642AADA0066DF275CD97AB70C">
<mods:title id="87AB0AA6C1B5082AFEC2D1AD75DB6AE7">Macrozoque hoplandrioides (Coleoptera: Staphylinidae: Aleocharinae), a new genus and species of false Lomechusini from México</mods:title>
</mods:titleInfo>
<mods:name id="5156CA9972BF456DB2ECC228F2531B00" type="personal">
<mods:role id="F4058BD60C4805E685E5D115092E6C6B">
<mods:roleTerm id="C92F29422A09F93A57C2C566AA06FEE1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3788E29A27BF0733DB4682E3D81E319E">Santiago-Jiménez, Quiyari J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B2CC88492413C3A76D0D873B6CD084E9">text</mods:typeOfResource>
<mods:relatedItem id="47146E674DE6086E1A84511189ED4011" type="host">
<mods:titleInfo id="D0152FF3EB73C85655B4EA2A69336BFA">
<mods:title id="DFF28DC14003DA38AA3BF45CFF1D6287">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6490FEB1922E6FB246E546E698C42B6C">
<mods:date id="4D98D91419E2273556C86047358C9F49">2016</mods:date>
<mods:detail id="D082575C8AAE2D4AF3D9AEC9DED3D214" type="pubDate">
<mods:number id="6B671C8443FCAD56F44CF847A8E49629">2016-02-11</mods:number>
</mods:detail>
<mods:detail id="727B46C7EE4FE63CF350F7EF59E00195" type="volume">
<mods:number id="C753919071A5939FBCDD9F08EC5CBF60">4079</mods:number>
</mods:detail>
<mods:detail id="546881BFBE8100A4B3BB43F144EB9EFB" type="issue">
<mods:number id="0A8619A8D85C0579DFEB70E67AD1441B">2</mods:number>
</mods:detail>
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<mods:classification id="69041C6DED756506131CF24A0593118C">journal volume</mods:classification>
<mods:identifier id="0CF77A9B0D287C7EF7DB3A78FA2D69DC" type="CLB-Dataset">31516</mods:identifier>
<mods:identifier id="5ECE1C1B3B6CFAC4EF9B2F5C440B28E6" type="DOI">10.11646/zootaxa.4079.2.8</mods:identifier>
<mods:identifier id="38F3DDF2350761D25AD6FC516F2AC995" type="GBIF-Dataset">fe597a14-c52d-475b-b151-d9fda89fd971</mods:identifier>
<mods:identifier id="4D6B445A3591689745CF911929E50366" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="239714287575EFFC7BCB0CD41E63CB8D" type="Zenodo-Dep">1050736</mods:identifier>
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</mods:mods>
<treatment id="03E887CA32134176FF3E32C769C4F901" ID-DOI="http://doi.org/10.5281/zenodo.6075743" ID-GBIF-Taxon="137307245" ID-Zenodo-Dep="6075743" LSID="urn:lsid:plazi:treatment:03E887CA32134176FF3E32C769C4F901" httpUri="http://treatment.plazi.org/id/03E887CA32134176FF3E32C769C4F901" lastPageId="6" pageId="3">
<subSubSection id="C35B655732134173FF3E32C76B30F92B" pageId="3" type="nomenclature">
<paragraph id="8BFE36DC32134173FF3E32C7690FF94C" blockId="3.[151,818,1697,1757]" box="[151,818,1697,1723]" pageId="3">
<heading id="D0B681B032134173FF3E32C7690FF94C" bold="true" box="[151,818,1697,1723]" fontSize="11" level="1" pageId="3" reason="1">
<emphasis id="B935EACE32134173FF3E32C7690FF94C" bold="true" box="[151,818,1697,1723]" pageId="3">
<taxonomicName id="4C414D5F32134173FF3E32C768F0F94C" ID-CoL="3XD28" authority="Santiago-Jimenez" authorityName="Santiago-Jiménez" authorityYear="2016" box="[151,717,1697,1723]" class="Insecta" family="Staphylinidae" genus="Macrozoque" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="hoplandrioides" status="sp. nov.">
<emphasis id="B935EACE32134173FF3E32C76BD4F94C" bold="true" box="[151,489,1697,1723]" italics="true" pageId="3">Macrozoque hoplandrioides</emphasis>
Santiago-Jiménez
</taxonomicName>
<taxonomicNameLabel id="A20657B532134173FD7C32C7690FF94C" box="[725,818,1697,1723]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE36DC32134173FF3E32A26B30F92B" blockId="3.[151,818,1697,1757]" box="[151,269,1732,1757]" pageId="3">
<figureCitation id="137A2A5932134173FF3E32A26B30F92B" box="[151,269,1732,1757]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 17" captionStart-2="FIGURE 18" captionStartId-0="4.[151,250,1909,1931]" captionStartId-1="5.[151,264,1855,1878]" captionStartId-2="6.[151,250,1259,1281]" captionTargetBox-0="[406,1180,755,1888]" captionTargetBox-1="[217,1369,193,1835]" captionTargetBox-2="[282,1306,197,1235]" captionTargetId-0="figure@4.[406,1180,755,1888]" captionTargetId-1="figure@5.[217,1369,193,1835]" captionTargetId-2="figure@6.[276,1310,193,1237]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Habitus of Macrozoque hoplandrioides gen. nov. and sp. nov., male. Scale bar = 2 mm." captionText-1="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." captionText-2="FIGURE 18. Collection site of Macrozoque hoplandrioides gen. nov. and sp. nov. (black circle)." httpUri-0="https://zenodo.org/record/1050738/files/figure.png" httpUri-1="https://zenodo.org/record/1050740/files/figure.png" httpUri-2="https://zenodo.org/record/1050742/files/figure.png" pageId="3">Figs. 118</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35B655732134173FF3E336A689DF867" pageId="3" type="materials_examined">
<paragraph id="8BFE36DC32134173FF3E336A689DF867" blockId="3.[151,1437,1804,2009]" pageId="3">
<materialsCitation id="3B293C8132134173FF3E336A6BE3F89B" ID-GBIF-Occurrence="1700783196" collectingDate="2013-06-10" collectingDateMax="2013-07-10" collectingDateMin="2013-06-10" collectionCode="IEXA" collectorName="El Gringo. Alt. &amp; E. Mora &amp; L. Delgado" country="Mexico" elevation="1600" latitude="16.67861" location="San Miguel Chimalapa" longLatPrecision="21" longitude="-94.26278" municipality="San Antonio" pageId="3" specimenCount="2" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis id="B935EACE32134173FF3E336A6BF3F8D2" bold="true" box="[151,462,1804,1829]" pageId="3">
<typeStatus id="54FA887E32134173FF3E336A6AEFF8D2" box="[151,210,1804,1829]" pageId="3">Type</typeStatus>
material.
<typeStatus id="54FA887E32134173FEF1336A6BF4F8D2" box="[344,457,1804,1829]" pageId="3" type="holotype">Holotype</typeStatus>
:
</emphasis>
male,
<emphasis id="B935EACE32134173FD8E336A689BF8D2" bold="true" box="[551,678,1804,1829]" pageId="3">
<collectingCountry id="F356764C32134173FD8E336A689CF8D2" box="[551,673,1804,1829]" name="Mexico" pageId="3">MÉXICO</collectingCountry>
:
</emphasis>
<collectingRegion id="4985F83E32134173FD1D336A6936F8D3" box="[692,779,1804,1828]" country="Mexico" name="Oaxaca" pageId="3">Oaxaca</collectingRegion>
,
<location id="8E9E600732134173FCB5336A6E10F8D2" LSID="urn:lsid:plazi:treatment:03E887CA32134176FF3E32C769C4F901:8E9E600732134173FCB5336A6E10F8D2" box="[796,1069,1804,1829]" country="Mexico" latitude="16.67861" longLatPrecision="21" longitude="-94.26278" municipality="San Antonio" name="San Miguel Chimalapa" pageId="3" stateProvince="Oaxaca">San Miguel Chimalapa</location>
,
<collectingMunicipality id="6B9AACA632134173FB96336A6EEBF8D2" box="[1087,1238,1804,1829]" pageId="3">San Antonio</collectingMunicipality>
,
<collectorName id="26B4530A32134173FB4E336A6FA0F8D2" box="[1255,1437,1804,1829]" pageId="3">El Gringo. Alt.</collectorName>
<elevation id="006CD1EF32134173FF3E33496AD3F8BF" box="[151,238,1839,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" unit="m" value="1600.0">
<quantity id="4CB99B3932134173FF3E33496AD3F8BF" box="[151,238,1839,1864]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" unit="m" value="1600.0">1,600m</quantity>
</elevation>
, Bosque Mesófilo.
<collectingDate id="EFBBE9F432134173FE62334968A2F8BF" box="[459,671,1839,1864]" pageId="3" value="2013-06-10" valueMax="2013-07-10" valueMin="2013-06-10">
<date id="FFFF101C32134173FE62334968A2F8BF" box="[459,671,1839,1864]" pageId="3" value="2013-06-10" valueMax="2013-07-10" valueMin="2013-06-10">10.VI10.VII.2013</date>
</collectingDate>
,
<geoCoordinate id="EE75501B32134173FD043349690EF8BF" box="[685,819,1839,1864]" direction="north" orientation="latitude" pageId="3" precision="15" value="16.67861">16°40'43&quot;N</geoCoordinate>
,
<geoCoordinate id="EE75501B32134173FCE8334969F6F8BF" box="[833,971,1839,1864]" direction="west" orientation="longitude" pageId="3" precision="15" value="-94.26278">94°15'46&quot;W</geoCoordinate>
, ex. trampa de intercepción de vuelo.
<collectorName id="26B4530A32134173FA2D33566AE9F89B" pageId="3">E. Mora</collectorName>
y
<collectorName id="26B4530A32134173FF5133336B47F89A" box="[248,378,1876,1901]" pageId="3">L. Delgado</collectorName>
(
<collectionCode id="ED50AE1932134173FE2433326BEBF89B" box="[397,470,1876,1900]" pageId="3">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B293C8132134173FE453332689DF867" ID-GBIF-Occurrence="1700783182" collectingDate="2013-06-10" collectingDateMax="2013-07-10" collectingDateMin="2013-06-10" collectionCode="IEXA" collectorName="El Gringo. Alt. &amp; E. Mora &amp; L. Delgado" country="Mexico" elevation="1600" latitude="16.67861" location="San Miguel Chimalapa" longLatPrecision="21" longitude="-94.26278" municipality="San Antonio" pageId="3" specimenCount="10" specimenCount-male="8" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis id="B935EACE32134173FE4533326850F89A" bold="true" box="[492,621,1876,1901]" pageId="3">
<typeStatus id="54FA887E32134173FE4533326855F89A" box="[492,616,1876,1901]" pageId="3" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as holotype (
<specimenCount id="9D47FD5532134173FC26333269D4F89A" box="[911,1001,1876,1901]" pageId="3" type="male">8 males</specimenCount>
(
<collectionCode id="ED50AE1932134173FC5433326E78F89B" box="[1021,1093,1876,1900]" pageId="3">IEXA</collectionCode>
),
<specimenCount id="9D47FD5532134173FBF433326EDDF89A" box="[1117,1248,1876,1901]" pageId="3" type="generic">1 specimen</specimenCount>
in 96% ethanol (MUZ-UV),
<specimenCount id="9D47FD5532134173FE8E331E6B9BF867" box="[295,422,1911,1936]" pageId="3" type="generic">1 specimen</specimenCount>
on slide (MUZ-UV)).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35B655732134176FF6E33FA6BACF9B5" lastPageId="6" pageId="3" type="description">
<paragraph id="8BFE36DC32134174FF6E33FA6AC2FF0F" blockId="3.[151,1437,1804,2009]" lastBlockId="4.[151,1436,151,717]" lastPageId="4" pageId="3">
<emphasis id="B935EACE32134173FF6E33FA6B64F842" bold="true" box="[199,345,1948,1973]" pageId="3">Description.</emphasis>
Body length
<quantity id="4CB99B3932134173FE5833FA6841F843" box="[497,636,1948,1973]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" metricValueMax="9.5" metricValueMin="9.3" pageId="3" unit="mm" value="9.4" valueMax="9.5" valueMin="9.3">9.39.5 mm</quantity>
. Most of body black; pro- and mesofemora in apical half brown except black apex; pro- and mesotibiae and tarsi brown; basal two thirds of metafemora brown-yellowish, apical third black; metatibiae and metatarsi brown-yellowish; antennomeres 111 brown, 411 darker, but apex of 11 brownyellowish. Head, pronotum, elytra and abdominal sternites with sparse pubescence; abdominal tergites almost glabrous.
</paragraph>
<paragraph id="8BFE36DC32144174FF6E35626E35FEB7" blockId="4.[151,1436,151,717]" pageId="4">
<emphasis id="B935EACE32144174FF6E35626B39FEEA" box="[199,260,260,285]" italics="true" pageId="4">Head</emphasis>
(
<figureCitation id="137A2A5932144174FEB335626B5BFEEB" box="[282,358,260,285]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 2</figureCitation>
). Dorsal surface without impression, protuberance or carinae. Antennomeres (
<figureCitation id="137A2A5932144174FAB835626F61FEEB" box="[1297,1372,260,285]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 9</figureCitation>
) 18 elongate, antenommere 3 very long; 9 subquadrate; 10 transverse; 11 elongate.
</paragraph>
<paragraph id="8BFE36DC32144174FF6E352A6865FE92" blockId="4.[151,1436,151,717]" box="[199,600,332,357]" pageId="4">
<emphasis id="B935EACE32144174FF6E352A6B25FE92" box="[199,280,332,357]" italics="true" pageId="4">Thorax</emphasis>
. As described for the genus.
</paragraph>
<paragraph id="8BFE36DC32144174FF6E3509697AFE03" blockId="4.[151,1436,151,717]" pageId="4">
<emphasis id="B935EACE32144174FF6E35096B0CFE7F" box="[199,305,367,392]" italics="true" pageId="4">Abdomen</emphasis>
. As described for the genus. Other conspicuous characters are: sternite IV without reservoir or glands; each lateral margin of sternite VII with eight small denticles (
<figureCitation id="137A2A5932144174FC0935F269CEFE5B" box="[928,1011,404,429]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 14</figureCitation>
); each apico-lateral margin of sternite VIII with six small denticles (
<figureCitation id="137A2A5932144174FE5B35D16875FE27" box="[498,584,439,464]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 14</figureCitation>
); apical margin of tergite VIII with one lateral point on each side and six minute denticles medially making margin crenulate (
<figureCitation id="137A2A5932144174FD4F35BA6904FE03" box="[742,825,476,501]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BFE36DC32144174FF6E359968FDFD3A" blockId="4.[151,1436,151,717]" pageId="4">
<emphasis id="B935EACE32144174FF6E35996836FDEF" box="[199,523,511,536]" italics="true" pageId="4">Secondary sexual structures.</emphasis>
Possible secondary sexual structures of male include the following: each elytron with bifurcate carina parallel to external margin, elytral disc with impression parallel to external margin; abdominal terga IIIIV with deep impression posterior to basal impression, the same impression moderately deep on tergites VVI; parasternite III with a protuberance on apico-lateral corner; tergite IV with pit-like impression on each side of midline; base of sternite VII with external gland (
<figureCitation id="137A2A5932144174FD7236E96910FD5F" box="[731,813,655,680]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="4">Fig. 13</figureCitation>
); tergite VIII with protuberance on midline. The lack of these characters in female needs to be confirmed.
</paragraph>
<caption id="DF3E665432144174FF3E33136EAFF87C" box="[151,1170,1909,1932]" httpUri="https://zenodo.org/record/1050738/files/figure.png" pageId="4" targetBox="[406,1180,755,1888]" targetPageId="4">
<paragraph id="8BFE36DC32144174FF3E33136EAFF87C" blockId="4.[151,1170,1909,1932]" box="[151,1170,1909,1932]" pageId="4">
<emphasis id="B935EACE32144174FF3E33136B2EF87C" bold="true" box="[151,275,1909,1931]" pageId="4">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="4C414D5F32144174FE203311689BF87C" authority="Santiago-Jiménez, 2016" authorityName="Santiago-Jiménez" authorityYear="2016" box="[393,678,1909,1932]" class="Insecta" family="Staphylinidae" genus="Macrozoque" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="hoplandrioides" status="gen. nov.">
<emphasis id="B935EACE32144174FE203311689BF87C" box="[393,678,1909,1932]" italics="true" pageId="4">Macrozoque hoplandrioides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657B532144174FD04331069B0F87C" box="[685,909,1909,1931]" pageId="4" rank="species">
<emphasis id="B935EACE32144174FD0433106936F87C" bold="true" box="[685,779,1910,1931]" pageId="4">gen. nov.</emphasis>
and
<emphasis id="B935EACE32144174FC94331069B0F87C" bold="true" box="[829,909,1910,1931]" pageId="4">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="DF3E665432154175FF3E33596B53F818" httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="5" targetBox="[217,1369,193,1835]" targetPageId="5">
<paragraph id="8BFE36DC32154175FF3E33596B53F818" blockId="5.[151,1436,1855,2031]" pageId="5">
<emphasis id="B935EACE32154175FF3E33596B7AF8A2" bold="true" box="[151,327,1855,1878]" pageId="5">FIGURES 217.</emphasis>
Structures of
<taxonomicName id="4C414D5F32154175FE7C332768CFF8A1" authority="Santiago-Jiménez, 2016" authorityName="Santiago-Jiménez" authorityYear="2016" box="[469,754,1856,1878]" class="Insecta" family="Staphylinidae" genus="Macrozoque" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="hoplandrioides" status="gen. nov.">
<emphasis id="B935EACE32154175FE7C332768CFF8A1" box="[469,754,1856,1878]" italics="true" pageId="5">Macrozoque hoplandrioides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657B532154175FD51332669E4F8A2" box="[760,985,1856,1878]" pageId="5" rank="species">
<emphasis id="B935EACE32154175FD513326696AF8A2" bold="true" box="[760,855,1856,1877]" pageId="5">gen. nov.</emphasis>
and
<emphasis id="B935EACE32154175FC21332669E4F8A2" bold="true" box="[904,985,1856,1877]" pageId="5">sp. nov.</emphasis>
</taxonomicNameLabel>
2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VIIVIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VIIVIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="DF3E665432164176FF3E308D6E4FFAF6" box="[151,1138,1259,1281]" httpUri="https://zenodo.org/record/1050742/files/figure.png" pageId="6" targetBox="[282,1306,197,1235]" targetPageId="6">
<paragraph id="8BFE36DC32164176FF3E308D6E4FFAF6" blockId="6.[151,1138,1259,1281]" box="[151,1138,1259,1281]" pageId="6">
<emphasis id="B935EACE32164176FF3E308D6B22FAF7" bold="true" box="[151,287,1259,1281]" pageId="6">FIGURE 18.</emphasis>
Collection site of
<taxonomicName id="4C414D5F32164176FE73308A68CAFAF6" authority="Santiago-Jiménez, 2016" authorityName="Santiago-Jiménez" authorityYear="2016" box="[474,759,1259,1281]" class="Insecta" family="Staphylinidae" genus="Macrozoque" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="hoplandrioides" status="gen. nov.">
<emphasis id="B935EACE32164176FE73308A68CAFAF6" box="[474,759,1259,1281]" italics="true" pageId="6">Macrozoque hoplandrioides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657B532164176FD57308D69E3FAF7" box="[766,990,1259,1281]" pageId="6" rank="species">
<emphasis id="B935EACE32164176FD57308D6961FAF7" bold="true" box="[766,860,1259,1280]" pageId="6">gen. nov.</emphasis>
and
<emphasis id="B935EACE32164176FC27308D69E3FAF7" bold="true" box="[910,990,1259,1280]" pageId="6">sp. nov.</emphasis>
</taxonomicNameLabel>
(black circle).
</paragraph>
</caption>
<paragraph id="8BFE36DC32164176FF6E314B6BDBF9E9" blockId="6.[151,1436,1325,1782]" pageId="6">
<emphasis id="B935EACE32164176FF6E314B6B06FAB1" box="[199,315,1325,1350]" italics="true" pageId="6">Aedeagus.</emphasis>
Internal sac of medial lobe (
<figureCitation id="137A2A5932164176FD2E314B6932FAB1" box="[647,783,1325,1350]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="6">Figs. 1617</figureCitation>
) without evident spinules, but with triangular sclerotized plate at apex (
<figureCitation id="137A2A5932164176FE9731376BA8FA9D" box="[318,405,1361,1386]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="6">Fig. 16</figureCitation>
); medial lobe with long compressor plate (extended almost over entire length of median lobe); apical lobe curved ventrad (visible in lateral view) and pointed; basal ridge convex. Paramere as in
<figureCitation id="137A2A5932164176FAED31136FAAFA79" box="[1348,1431,1397,1422]" captionStart="FIGURES 2 17" captionStartId="5.[151,264,1855,1878]" captionTargetBox="[217,1369,193,1835]" captionTargetId="figure@5.[217,1369,193,1835]" captionTargetPageId="5" captionText="FIGURES 2 17. Structures of Macrozoque hoplandrioides gen. nov. and sp. nov. 2, head, dorsal view; 3, left mandible dorsal view; 4, right mandible, dorsal view; 5, left maxilla, dorsal view; 6, pronotum, dorsal view; 7, prementum and labial palpi, dorsal view; 8, labrum, dorsal view; 9, antenna; 10, right elytra, dorsal view; 11, meso- and metaventrite, ventral view; 12, tergites VII VIII, dorsal view; 13, sternite VII, showing the external gland (arrows), ventral view; 14, sternites VII VIII, ventral view; 15, paramere, lateral view; 16, medial lobe of aedeagus, lateral view; 17, medial lobe of aedeagus, dorsal view. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1050740/files/figure.png" pageId="6">Fig. 15</figureCitation>
; anterodorsal margin of paramerite with prominent sensory pores present beneath the velar sac; hinge zone of paramerite faint, extended from dorsal surface to near articulation between condylite and paramerite; apical process of paramerite clearly articulated anterior to edge of velum; condylite with row of sensory pores; velum short (less than half length of paramere).
</paragraph>
<paragraph id="8BFE36DC32164176FF6E324F6BACF9B5" blockId="6.[151,1436,1325,1782]" box="[199,401,1577,1602]" pageId="6">
<emphasis id="B935EACE32164176FF6E324F6B1EF9B5" box="[199,291,1577,1602]" italics="true" pageId="6">Female.</emphasis>
Unkown.
</paragraph>
</subSubSection>
<subSubSection id="C35B655732164176FF6E322B6B69F959" pageId="6" type="discussion">
<paragraph id="8BFE36DC32164176FF6E322B6B69F959" blockId="6.[151,1436,1325,1782]" pageId="6">
<emphasis id="B935EACE32164176FF6E322B6B06F991" bold="true" box="[199,315,1613,1638]" pageId="6">Remarks.</emphasis>
<taxonomicName id="4C414D5F32164176FEEB32286840F991" authorityName="Santiago-Jiménez" authorityYear="2016" box="[322,637,1613,1638]" class="Insecta" family="Staphylinidae" genus="Macrozoque" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="hoplandrioides">
<emphasis id="B935EACE32164176FEEB32286840F991" box="[322,637,1613,1638]" italics="true" pageId="6">Macrozoque hoplandrioides</emphasis>
</taxonomicName>
is the only described species in the genus. The characters that could be useful at species level are the shape of the aedeagus, and impressions and protuberances on the abdomen as described above.
</paragraph>
</subSubSection>
<subSubSection id="C35B655732164176FF6E32DF69C4F901" pageId="6" type="etymology">
<paragraph id="8BFE36DC32164176FF6E32DF69C4F901" blockId="6.[151,1436,1325,1782]" pageId="6">
<emphasis id="B935EACE32164176FF6E32DF6B73F925" bold="true" box="[199,334,1721,1746]" pageId="6">Etymology.</emphasis>
The general shape of the abdomen of this species resembles species of
<taxonomicName id="4C414D5F32164176FBC632DF6ECCF925" authorityName="Kraatz" authorityYear="1857" box="[1135,1265,1721,1746]" class="Insecta" family="Staphylinidae" genus="Hoplandria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B935EACE32164176FBC632DF6ECCF925" box="[1135,1265,1721,1746]" italics="true" pageId="6">Hoplandria</emphasis>
</taxonomicName>
. The name is a combination of
<taxonomicName id="4C414D5F32164176FEE132BB6BF6F901" authorityName="Kraatz" authorityYear="1857" box="[328,459,1757,1782]" class="Insecta" family="Staphylinidae" genus="Hoplandria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="B935EACE32164176FEE132BB6BF6F901" box="[328,459,1757,1782]" italics="true" pageId="6">Hoplandria</emphasis>
</taxonomicName>
(stem:
<emphasis id="B935EACE32164176FDB632BB68AEF901" box="[543,659,1757,1782]" italics="true" pageId="6">Hoplandri</emphasis>
-), and -
<emphasis id="B935EACE32164176FD4432BB6915F901" box="[749,808,1757,1782]" italics="true" pageId="6">oides</emphasis>
that means “like”.
</paragraph>
</subSubSection>
</treatment>
</document>