treatments-xml/data/03/E8/87/03E88785FFD2421CA568C97897C0613C.xml

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<mods:title id="E7DAF6CFE7B095AD882BAD0126A30D73">Review of the Sphaeropthalma imperialis species­group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes</mods:title>
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<treatment id="03E88785FFD2421CA568C97897C0613C" ID-DOI="http://doi.org/10.5281/zenodo.6252843" ID-GBIF-Taxon="119346082" ID-Zenodo-Dep="6252843" LSID="urn:lsid:plazi:treatment:03E88785FFD2421CA568C97897C0613C" httpUri="http://treatment.plazi.org/id/03E88785FFD2421CA568C97897C0613C" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
<subSubSection id="C35B6518FFD24210A568C97896E4620C" box="[264,715,1124,1150]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BFE3693FFD24210A568C97896E4620C" blockId="5.[264,715,1124,1150]" box="[264,715,1124,1150]" pageId="5" pageNumber="6">
<heading id="D0B681FFFFD24210A568C97896E4620C" bold="true" box="[264,715,1124,1150]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="4C414D10FFD24210A568C97896E4620C" authority="Cresson" authorityName="Cresson" box="[264,715,1124,1150]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B935EA81FFD24210A568C97896E4620C" bold="true" box="[264,715,1124,1150]" pageId="5" pageNumber="6">
<emphasis id="B935EA81FFD24210A568C9789663620C" bold="true" box="[264,588,1124,1150]" italics="true" pageId="5" pageNumber="6">Sphaeropthalma edwardsii</emphasis>
(Cresson)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6518FFD24210A568C9B390F56334" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BFE3693FFD24210A568C9B395B16377" blockId="5.[264,1324,1198,1350]" pageId="5" pageNumber="6">
<taxonomicName id="4C414D10FFD24210A568C9B3964462B7" authority="Edwardsii Cresson, 1875" authorityName="Edwardsii Cresson" authorityYear="1875" box="[264,619,1198,1222]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B935EA81FFD24210A568C9B395E662B4" box="[264,457,1199,1222]" italics="true" pageId="5" pageNumber="6">Mutilla Edwardsii</emphasis>
<bibRefCitation id="EFD04B62FFD24210A5B0C9B2964462B7" author="Cresson" box="[464,619,1198,1221]" pageId="5" pageNumber="19" refString="Cresson, E. T. (1875) Descriptions of new species of Mutilla. Transactions of the American Entomological Society, 5, 119 - 120." type="journal article" year="1875">Cresson, 1875</bibRefCitation>
</taxonomicName>
. Amer. Ent. Soc., Trans. 5: 119. Male. 
<typeStatus id="54FA8831FFD24210A067C9B2904062B7" box="[1031,1135,1198,1221]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
: Oregon (ANSP). 
<taxonomicName id="4C414D10FFD24210A568C9D397A16297" authority="Cresson" authorityName="Cresson" box="[264,910,1230,1254]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="edwardsii" subGenus="Photopsis" subSpecies="edwardsii">
<emphasis id="B935EA81FFD24210A568C9D395986294" box="[264,439,1231,1254]" italics="true" pageId="5" pageNumber="6">Sphaeropthalma</emphasis>
(
<emphasis id="B935EA81FFD24210A5A9C9D3961D6294" box="[457,562,1231,1254]" italics="true" pageId="5" pageNumber="6">Photopsis</emphasis>
) 
<emphasis id="B935EA81FFD24210A623C9D397336294" box="[579,796,1231,1254]" italics="true" pageId="5" pageNumber="6">edwardsii edwardsii</emphasis>
(Cresson)
</taxonomicName>
, 
<bibRefCitation id="EFD04B62FFD24210A7FBC9D290176297" author="Schuster" box="[923,1080,1230,1253]" pageId="5" pageNumber="20" refString="Schuster, R. M. (1958) A revision of the Sphaeropthalmine Mutillidae of America North of Mexico. II. Entomologica Americana, 37, 1 - 130." type="journal article" year="1958">Schuster 1958</bibRefCitation>
. Ent. Amer. (n. s.) 37: 36. Male.
</paragraph>
<paragraph id="8BFE3693FFD24210A568C81390F56334" blockId="5.[264,1324,1198,1350]" pageId="5" pageNumber="6">
<taxonomicName id="4C414D10FFD24210A568C81397E16357" authority="Schuster, 1958" authorityName="Schuster" authorityYear="1958" box="[264,974,1294,1318]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="edwardsii" subGenus="Photopsis" subSpecies="flammifera">
<emphasis id="B935EA81FFD24210A568C81395986354" box="[264,439,1295,1318]" italics="true" pageId="5" pageNumber="6">Sphaeropthalma</emphasis>
(
<emphasis id="B935EA81FFD24210A5A8C813961E6354" box="[456,561,1295,1318]" italics="true" pageId="5" pageNumber="6">Photopsis</emphasis>
) 
<emphasis id="B935EA81FFD24210A621C81397096354" box="[577,806,1295,1318]" italics="true" pageId="5" pageNumber="6">edwardsii flammifera</emphasis>
<bibRefCitation id="EFD04B62FFD24210A74DC81297E16357" author="Schuster" box="[813,974,1294,1317]" pageId="5" pageNumber="20" refString="Schuster, R. M. (1958) A revision of the Sphaeropthalmine Mutillidae of America North of Mexico. II. Entomologica Americana, 37, 1 - 130." type="journal article" year="1958">Schuster, 1958</bibRefCitation>
</taxonomicName>
. Ent. Amer. (n. s.) 37: 36. Male. 
<typeStatus id="54FA8831FFD24210A558C832958F6337" box="[312,416,1326,1349]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
: California, Antioch, 
<date id="FFFF1053FFD24210A6E6C83297226337" box="[646,781,1326,1349]" pageId="5" pageNumber="6" value="1941-09-14">14.Sep.1941</date>
, coll. J.R. Fisher (UMSP). 
<emphasis id="B935EA81FFD24210A04FC83290F56334" bold="true" box="[1071,1242,1326,1350]" pageId="5" pageNumber="6">New Synonym.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35B6518FFD24210A568C86B90A36103" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BFE3693FFD24210A568C86B955063E2" blockId="5.[264,1323,1399,1905]" box="[264,383,1399,1424]" pageId="5" pageNumber="6">
<emphasis id="B935EA81FFD24210A568C86B955063E2" box="[264,383,1399,1424]" italics="true" pageId="5" pageNumber="6">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BFE3693FFD24210A558C88390A36103" blockId="5.[264,1323,1399,1905]" pageId="5" pageNumber="6">
The male of 
<taxonomicName id="4C414D10FFD24210A5B1C883964C63CA" box="[465,611,1439,1464]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B935EA81FFD24210A5B1C883964C63CA" box="[465,611,1439,1464]" italics="true" pageId="5" pageNumber="6">S. edwardsii</emphasis>
</taxonomicName>
can be separated from all other 
<taxonomicName id="4C414D10FFD24210A786C883908863CA" box="[998,1191,1439,1464]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B935EA81FFD24210A786C883908863CA" box="[998,1191,1439,1464]" italics="true" pageId="5" pageNumber="6">Sphaeropthalma</emphasis>
</taxonomicName>
species by its coloration. The pubescence varies from yellow to scarlet, while the integumental coloration varies from orange to piceous and the wings are dark brown to black (
<figureCitation id="137A2A16FFD24210A0A5C8F39132607B" box="[1221,1309,1519,1545]" captionStart="FIGURES 26 – 29" captionStartId="8.[264,382,1049,1073]" captionTargetBox="[269,1316,326,1025]" captionTargetId="figure@8.[269,1317,326,1025]" captionTargetPageId="8" captionText="FIGURES 26 – 29. Males, lateral view: 26. Sphaeropthalma edwardsii; 27. S. imperialis; 28. S. megagnathos; 29. S. marpesia." httpUri="https://zenodo.org/record/172967/files/figure.png" pageId="5" pageNumber="6">Fig. 26</figureCitation>
). This species also has the following unique combination of characters. The mandible is diagnostic being moderately dilated, distally little or scarcely wider than at tooth, the ventral basal tooth of the mandible is small, and the apex is vertical. Also, the clypeus is moderately depressed below the dorsal mandibular margin, the sternal felt line is absent, and the genitalic morphology is unique (
<figureCitation id="137A2A16FFD24210A68DCBAB975F60A3" box="[749,880,1719,1745]" captionStart="FIGURES 8 – 19" captionStartId="6.[264,382,1274,1298]" captionTargetBox="[271,1322,322,1273]" captionTargetId="figure@6.[271,1330,300,1286]" captionTargetPageId="6" captionText="FIGURES 8 – 19. Genitalia: penial valve, lateral view; internal lateral view, penial valve removed; ventral view left, dorsal view right; Sphaeropthalma edwardsii: 8 – 10. S. imperialis: 11 – 13. S. marpesia: 14 – 16; S. megagnathos: 17 – 19." httpUri="https://zenodo.org/record/172965/files/figure.png" pageId="5" pageNumber="6">Figs. 8–10</figureCitation>
). The female of this species is easily recognized by the unique combination of having a small ventral angulation located basally on the mandible, a distinctly petiolate metasomal segment 1, a granulate pygidium, the presence of plumose setae especially on the fringes of the metasomal tergites, and the dense long yellow setae that obscures the integumental sculpturing (
<figureCitation id="137A2A16FFD24210A047CA4B90506103" box="[1063,1151,1879,1905]" captionStart="FIGURES 20 – 25" captionStartId="7.[264,383,1710,1734]" captionTargetBox="[271,1315,526,1686]" captionTargetId="figure@7.[271,1315,526,1686]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Sphaeropthalma imperialis female, dark form: 20. lateral view; 21. anterior portion of second metasomal tergite; S. imperialis female, light form: 22. lateral view; 23. pygidium, dorsal view; 24. S. edwardsii female, lateral view; 25. S. marpesia female, lateral view." httpUri="https://zenodo.org/record/172966/files/figure.png" pageId="5" pageNumber="6">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3E661BFFD14213A568C9E696E46324" httpUri="https://zenodo.org/record/172965/files/figure.png" pageId="6" pageNumber="7" targetBox="[271,1322,322,1273]" targetPageId="6">
<paragraph id="8BFE3693FFD14213A568C9E696E46324" blockId="6.[264,1323,1274,1367]" pageId="6" pageNumber="7">
<emphasis id="B935EA81FFD14213A568C9E695ED6360" bold="true" box="[264,450,1274,1298]" pageId="6" pageNumber="7">FIGURES 8–19.</emphasis>
Genitalia: penial valve, lateral view; internal lateral view, penial valve removed; ventral view left, dorsal view right; 
<taxonomicName id="4C414D10FFD14213A6C5C80197E76346" box="[677,968,1309,1332]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B935EA81FFD14213A6C5C80197E76346" box="[677,968,1309,1332]" italics="true" pageId="6" pageNumber="7">Sphaeropthalma edwardsii</emphasis>
</taxonomicName>
: 8–10. 
<taxonomicName id="4C414D10FFD14213A044C80190806346" box="[1060,1199,1309,1332]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B935EA81FFD14213A044C80190806346" box="[1060,1199,1309,1332]" italics="true" pageId="6" pageNumber="7">S. imperialis</emphasis>
</taxonomicName>
: 11–13. 
<taxonomicName id="4C414D10FFD14213A176C80195446325" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B935EA81FFD14213A176C80195446325" italics="true" pageId="6" pageNumber="7">S. marpesia</emphasis>
</taxonomicName>
: 14–16; 
<taxonomicName id="4C414D10FFD14213A5AAC85C965C6325" box="[458,627,1344,1367]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="megagnathos">
<emphasis id="B935EA81FFD14213A5AAC85C965C6325" box="[458,627,1344,1367]" italics="true" pageId="6" pageNumber="7">S. megagnathos</emphasis>
</taxonomicName>
: 17–19.
</paragraph>
</caption>
<subSubSection id="C35B6518FFD1421DA558C896977D6393" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BFE3693FFD14213A558C896959A617E" blockId="6.[264,1324,1418,1884]" pageId="6" pageNumber="7">
<emphasis id="B935EA81FFD14213A558C89695BF63D1" box="[312,400,1418,1443]" italics="true" pageId="6" pageNumber="7">Female</emphasis>
. 
<emphasis id="B935EA81FFD14213A5CCC896970663D1" box="[428,809,1418,1443]" italics="true" pageId="6" pageNumber="7">Coloration and Setal Pattern</emphasis>
(
<figureCitation id="137A2A16FFD14213A726C896978363D6" box="[838,940,1418,1444]" captionStart="FIGURES 20 – 25" captionStartId="7.[264,383,1710,1734]" captionTargetBox="[271,1315,526,1686]" captionTargetId="figure@7.[271,1315,526,1686]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Sphaeropthalma imperialis female, dark form: 20. lateral view; 21. anterior portion of second metasomal tergite; S. imperialis female, light form: 22. lateral view; 23. pygidium, dorsal view; 24. S. edwardsii female, lateral view; 25. S. marpesia female, lateral view." httpUri="https://zenodo.org/record/172966/files/figure.png" pageId="6" pageNumber="7">Fig. 24</figureCitation>
). Integument fuscous, except metasomal segments 5–6 ferruginous. Dorsum of head, mesosoma, and metasoma with long dense reddish to yellow­orange brachyplumose setae obscuring integument; setae denser on posterior margin of head and tergal margins. Slightly shorter plumose setae also present on dorsum. Head below antennal tubercles, scape, genal region, pleura, and legs with white brachyplumose setae, not as dense as on dorsum. Fringes of tergites with dense white yellow plumose setae; disk with sparse yellow plumose setae, as well as aforementioned setae. T3–T5 with anterior margin with dense white plumose setae. Sternites with sparse white brachyplumose setae. Fringes of sternites with dense white plumose setae.
</paragraph>
<paragraph id="8BFE3693FFD14212A558CA0690AC67A4" blockId="6.[264,1324,1418,1884]" lastBlockId="7.[264,1323,284,470]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B935EA81FFD14213A558CA0695586141" box="[312,375,1818,1843]" italics="true" pageId="6" pageNumber="7">Head</emphasis>
. Rounded posteriorly, not as wide as mesosoma; coarsely punctate. Eye round. Antennal tubercles striate. Scrobe with lateral carina beginning just below eye, becoming weaker above height of antennal tubercle but continuing to antennal tubercle. Mandible bidentate, with slight ventral angulation basally; dorsal carina continuing to internal tooth. Genal carina absent. Flagellomere I ~2.25X pedicel length; flagellomere II ~1.75X length of pedicel. Clypeus truncate with slight median emargination; tuberculate posteriorly, appearing longitudinally carinate from anterior margin to posterior tubercle.
</paragraph>
<caption id="DF3E661BFFD04212A568CBB291326179" httpUri="https://zenodo.org/record/172966/files/figure.png" pageId="7" pageNumber="8" targetBox="[271,1315,526,1686]" targetPageId="7">
<paragraph id="8BFE3693FFD04212A568CBB291326179" blockId="7.[264,1323,1710,1803]" pageId="7" pageNumber="8">
<emphasis id="B935EA81FFD04212A568CBB295FC60B4" bold="true" box="[264,467,1710,1734]" pageId="7" pageNumber="8">FIGURES 20–25.</emphasis>
<taxonomicName id="4C414D10FFD04212A5BECBB2972B60B7" box="[478,772,1710,1733]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B935EA81FFD04212A5BECBB2972B60B7" box="[478,772,1710,1733]" italics="true" pageId="7" pageNumber="8">Sphaeropthalma imperialis</emphasis>
</taxonomicName>
female, dark form: 20. lateral view; 21. anterior portion of second metasomal tergite; 
<taxonomicName id="4C414D10FFD04212A6DCCBCD9765609A" box="[700,842,1745,1768]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B935EA81FFD04212A6DCCBCD9765609A" box="[700,842,1745,1768]" italics="true" pageId="7" pageNumber="8">S. imperialis</emphasis>
</taxonomicName>
female, light form: 22. lateral view; 23. pygidium, dorsal view; 24. 
<taxonomicName id="4C414D10FFD04212A64DCBE8969F6179" box="[557,688,1780,1803]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B935EA81FFD04212A64DCBE8969F6179" box="[557,688,1780,1803]" italics="true" pageId="7" pageNumber="8">S. edwardsii</emphasis>
</taxonomicName>
female, lateral view; 25. 
<taxonomicName id="4C414D10FFD04212A7A0CBE890126179" box="[960,1085,1780,1803]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B935EA81FFD04212A7A0CBE890126179" box="[960,1085,1780,1803]" italics="true" pageId="7" pageNumber="8">S. marpesia</emphasis>
</taxonomicName>
female, lateral view.
</paragraph>
</caption>
<caption id="DF3E661BFFDF421DA568C905967E6221" httpUri="https://zenodo.org/record/172967/files/figure.png" pageId="8" pageNumber="9" targetBox="[269,1316,326,1025]" targetPageId="8">
<paragraph id="8BFE3693FFDF421DA568C905967E6221" blockId="8.[264,1323,1049,1107]" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA568C90595FD6243" bold="true" box="[264,466,1049,1073]" pageId="8" pageNumber="9">FIGURES 26–29.</emphasis>
Males, lateral view: 26. 
<taxonomicName id="4C414D10FFDF421DA68EC906903F6243" box="[750,1040,1050,1073]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="B935EA81FFDF421DA68EC906903F6243" box="[750,1040,1050,1073]" italics="true" pageId="8" pageNumber="9">Sphaeropthalma edwardsii</emphasis>
</taxonomicName>
; 27. 
<taxonomicName id="4C414D10FFDF421DA02EC90690F66243" box="[1102,1241,1050,1073]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B935EA81FFDF421DA02EC90690F66243" box="[1102,1241,1050,1073]" italics="true" pageId="8" pageNumber="9">S. imperialis</emphasis>
</taxonomicName>
; 28. 
<taxonomicName id="4C414D10FFDF421DA176C90695B96221" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="megagnathos">
<emphasis id="B935EA81FFDF421DA176C90695B96221" italics="true" pageId="8" pageNumber="9">S. megagnathos</emphasis>
</taxonomicName>
; 29. 
<taxonomicName id="4C414D10FFDF421DA5ACC92096656221" box="[460,586,1084,1107]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marpesia">
<emphasis id="B935EA81FFDF421DA5ACC92096656221" box="[460,586,1084,1107]" italics="true" pageId="8" pageNumber="9">S. marpesia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BFE3693FFDF421DA558C99B909F6333" blockId="8.[264,1323,1159,1505]" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA558C99B959A62D2" box="[312,437,1159,1184]" italics="true" pageId="8" pageNumber="9">Mesosoma</emphasis>
. Slightly longer than wide. Humeral angles dentate. Dorsum with distinct lateral margin. Coarsely punctate. Propleuron coarsely to moderately punctate. Mesopleuron with medial area moderately punctate and produced outward from dorsal to ventral margin; otherwise glabrous and nitid. Metapleuron and lateral faces of propodeum glabrous and nitid. Propodeum with distinct vertical and dorsal faces; reticulate.
</paragraph>
<paragraph id="8BFE3693FFDF421DA558C853977D6393" blockId="8.[264,1323,1159,1505]" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA558C853959D631A" box="[312,434,1359,1384]" italics="true" pageId="8" pageNumber="9">Metasoma</emphasis>
. Segment 1 distinctly petiolate with segment 2. T1 moderately punctate. T2 coarsely confluently punctate. T3–T5 and S3–S5 micropunctate, appearing granulate. S2 similar in punctation to T2, with anterior medial tumid region. Pygidium laterally defined by carinae, conspicuously granulate to colliculate.
</paragraph>
</subSubSection>
<subSubSection id="C35B6518FFDF421DA568CB0B975A602B" box="[264,885,1559,1625]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BFE3693FFDF421DA568CB0B95B86042" blockId="8.[264,885,1559,1625]" box="[264,407,1559,1584]" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA568CB0B95B86042" box="[264,407,1559,1584]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
</paragraph>
<paragraph id="8BFE3693FFDF421DA558CB23975A602B" blockId="8.[264,885,1559,1625]" box="[312,885,1599,1625]" pageId="8" pageNumber="9">Arizona, California, Idaho, Nevada, and Oregon.</paragraph>
</subSubSection>
<subSubSection id="C35B6518FFDF421CA568CB93967F631C" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BFE3693FFDF421DA568CB9395C960DA" blockId="8.[264,1323,1679,1905]" box="[264,486,1679,1704]" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA568CB9395C960DA" box="[264,486,1679,1704]" italics="true" pageId="8" pageNumber="9">Material examined</emphasis>
</paragraph>
<paragraph id="8BFE3693FFDF421CA558CBAB967F631C" blockId="8.[264,1323,1679,1905]" lastBlockId="9.[264,1323,283,1390]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B935EA81FFDF421DA558CBAB95C760A3" bold="true" box="[312,488,1719,1745]" pageId="8" pageNumber="9">
<collectingCountry id="F3567603FFDF421DA558CBAB955860A3" box="[312,375,1719,1745]" name="United States of America" pageId="8" pageNumber="9">USA</collectingCountry>
: Arizona
</emphasis>
: 
<emphasis id="B935EA81FFDF421DA598CBAB968460A2" box="[504,683,1719,1744]" italics="true" pageId="8" pageNumber="9">Santa Cruz Co.</emphasis>
, Madera Canyon, 1ɗ, 
<date id="FFFF1053FFDF421DA7D6CBAB907E60A3" box="[950,1105,1719,1745]" pageId="8" pageNumber="9" value="1968-08-26">26.Aug.1968</date>
, coll. K. Lorenzen (
<collectionCode id="ED50AE56FFDF421DA570CBC39541608B" box="[272,366,1759,1785]" pageId="8" pageNumber="9">KAWC</collectionCode>
). 
<emphasis id="B935EA81FFDF421DA5E4CBC3962B608B" bold="true" box="[388,516,1759,1785]" pageId="8" pageNumber="9">California</emphasis>
: 
<emphasis id="B935EA81FFDF421DA676CBC3969E608A" box="[534,689,1759,1784]" italics="true" pageId="8" pageNumber="9">Alameda Co.</emphasis>
, Tesla, 1ɗ 1Ψ, 
<date id="FFFF1053FFDF421DA70ECBC3902C608B" box="[878,1027,1759,1785]" pageId="8" pageNumber="9" value="1948-09-25">25.Sep.1948</date>
, coll. P.D. Hurd (
<collectionCode id="ED50AE56FFDF421DA0B6CBC39132608B" box="[1238,1309,1759,1785]" pageId="8" pageNumber="9">CISC</collectionCode>
); Tresia, 1ɗ, 
<date id="FFFF1053FFDF421DA5FDCA1B95C56153" box="[413,490,1799,1825]" pageId="8" pageNumber="9" value="1951-01">I.1951</date>
, coll. Thompson (
<collectionCode id="ED50AE56FFDF421DA6B2CA1B97366153" box="[722,793,1799,1825]" pageId="8" pageNumber="9">CISC</collectionCode>
); Tresia, 2ɗ, 
<date id="FFFF1053FFDF421DA7A8CA1B90736153" box="[968,1116,1799,1825]" pageId="8" pageNumber="9" value="1951-09-25">25.Sep.1951</date>
, coll. P.D. Hurd (
<collectionCode id="ED50AE56FFDF421DA570CA339579613B" box="[272,342,1839,1865]" pageId="8" pageNumber="9">CISC</collectionCode>
; 
<collectionCode id="ED50AE56FFDF421DA505CA339593613B" box="[357,444,1839,1865]" pageId="8" pageNumber="9">EMUS</collectionCode>
); 
<emphasis id="B935EA81FFDF421DA5B4CA33969E613A" box="[468,689,1839,1864]" italics="true" pageId="8" pageNumber="9">Contra Costa Co.,</emphasis>
Antioch, 1ɗ, 
<date id="FFFF1053FFDF421DA700CA3397DA613B" box="[864,1013,1839,1865]" pageId="8" pageNumber="9" value="1941-09-13">13.Sep.1941</date>
, coll. J.R. Fisher (
<collectionCode id="ED50AE56FFDF421DA0B6CA339132613B" box="[1238,1309,1839,1865]" pageId="8" pageNumber="9">CISC</collectionCode>
); Antioch, 1ɗ 1Ψ, 
<date id="FFFF1053FFDF421DA5B5CA4B965E6103" box="[469,625,1879,1905]" pageId="8" pageNumber="9" value="1952-08-22">22.Aug.1952</date>
, coll. G.A. Marsh (
<collectionCode id="ED50AE56FFDF421DA736CA4B97816103" box="[854,942,1879,1905]" pageId="8" pageNumber="9">EMUS</collectionCode>
); Antioch, 1ɗ 1Ψ, 
<date id="FFFF1053FFDF421DA0F0CA4B91096103" box="[1168,1318,1879,1905]" pageId="8" pageNumber="9" value="1954-09-26">26.Sep.1954</date>
, coll. M.S. Wasbauer (
<collectionCode id="ED50AE56FFDE421CA651CC0096596744" box="[561,630,284,310]" pageId="9" pageNumber="10">CISC</collectionCode>
, 
<collectionCode id="ED50AE56FFDE421CA6F0CC0096C96744" box="[656,742,284,310]" pageId="9" pageNumber="10">EMUS</collectionCode>
); Rock City, Mt Diablo, 1Ψ, 
<date id="FFFF1053FFDE421CA0F0CC0091056744" box="[1168,1322,284,310]" pageId="9" pageNumber="10" value="1940-05-24">24.May.1940</date>
(
<collectionCode id="ED50AE56FFDE421CA571CC589574672C" box="[273,347,324,350]" pageId="9" pageNumber="10">CISC</collectionCode>
);. 
<taxonomicName id="4C414D10FFDE421CA50CCC58965D672C" authority="Co., Arbuckle" authorityName="Co., Arbuckle" box="[364,626,324,350]" class="Insecta" family="Staphylinidae" genus="Colusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B935EA81FFDE421CA50CCC5895C5672F" box="[364,490,324,349]" italics="true" pageId="9" pageNumber="10">Colusa Co</emphasis>
., Arbuckle
</taxonomicName>
, 1ɗ, 
<date id="FFFF1053FFDE421CA6D1CC589762672C" box="[689,845,324,350]" pageId="9" pageNumber="10" value="1962-08-17">17.Aug.1962</date>
, coll. P.M. Marsh (
<collectionCode id="ED50AE56FFDE421CA052CC5890A5672C" box="[1074,1162,324,350]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA0FFCC589134672F" box="[1183,1307,324,349]" italics="true" pageId="9" pageNumber="10">Fresno Co</emphasis>
., Jacalitos Canyon, 2ɗ, 
<date id="FFFF1053FFDE421CA66ECC7096B667F4" box="[526,665,364,390]" pageId="9" pageNumber="10" value="1956-07-31">31.Jul.1956</date>
, coll. R.O. Schuster.Aug. (
<collectionCode id="ED50AE56FFDE421CA7BACC70901D67F4" box="[986,1074,364,390]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); L. Panoche Cr., 1ɗ, 
<date id="FFFF1053FFDE421CA568CC8895B667DC" box="[264,409,404,430]" pageId="9" pageNumber="10" value="1952-06-28">28.Jun.1952</date>
, coll. H.F. Robinson (
<collectionCode id="ED50AE56FFDE421CA6DECC88973C67DC" box="[702,787,404,430]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA756CC88979D67DF" box="[822,946,404,429]" italics="true" pageId="9" pageNumber="10">Kern Co.,</emphasis>
1ɗ, 
<date id="FFFF1053FFDE421CA79DCC8890B267DC" box="[1021,1181,404,430]" pageId="9" pageNumber="10" value="1962-06-05" valueMax="1962-06-06" valueMin="1962-06-05">5–6.Jun.1962</date>
, coll. R.E. Bellamy (
<collectionCode id="ED50AE56FFDE421CA51FCCA095F867A4" box="[383,471,444,470]" pageId="9" pageNumber="10">EMUS</collectionCode>
); Bakersfield, 9ɗ, 
<date id="FFFF1053FFDE421CA6DECCA0976867A4" box="[702,839,444,470]" pageId="9" pageNumber="10" value="1951-07-20">20.Jul.1951</date>
– 
<date id="FFFF1053FFDE421CA738CCA097CF67A4" box="[856,992,444,470]" pageId="9" pageNumber="10" value="1951-07-28">28.Jul.1951</date>
, coll. L.W. Isaak (
<collectionCode id="ED50AE56FFDE421CA0A6CCA0913167A4" box="[1222,1310,444,470]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); Taft, 1ɗ, 
<date id="FFFF1053FFDE421CA519CCF8962B678C" box="[377,516,484,510]" pageId="9" pageNumber="10" value="1956-07-25">25.Jul.1956</date>
, coll. T.R. Haig (
<collectionCode id="ED50AE56FFDE421CA6B2CCF89705678C" box="[722,810,484,510]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); Wasco, 1ɗ, 
<date id="FFFF1053FFDE421CA7AFCCF8904E678C" box="[975,1121,484,510]" pageId="9" pageNumber="10" value="1951-06-26">26.Jun.1951</date>
, coll. L.W. Isaak (
<collectionCode id="ED50AE56FFDE421CA570CF1095476454" box="[272,360,524,550]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA5E5CF10962C6457" box="[389,515,524,549]" italics="true" pageId="9" pageNumber="10">Kings Co.</emphasis>
, Helm, 1ɗ, 
<date id="FFFF1053FFDE421CA6CDCF1097176454" box="[685,824,524,550]" pageId="9" pageNumber="10" value="1955-07-24">24.Jul.1955</date>
, coll. H. Michalk (
<collectionCode id="ED50AE56FFDE421CA056CF1090A16454" box="[1078,1166,524,550]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 4.8 mi W Kettleman City, 1ɗ, 
<date id="FFFF1053FFDE421CA66BCF2896B2643C" box="[523,669,564,590]" pageId="9" pageNumber="10" value="1970-06-13">13.Jun.1970</date>
, coll. F.G. Andrews &amp; M.S. Wasbauer (
<collectionCode id="ED50AE56FFDE421CA0F4CF2890C8643C" box="[1172,1255,564,590]" pageId="9" pageNumber="10">CDFA</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA160CF2895BA6407" italics="true" pageId="9" pageNumber="10">Los Angeles Co</emphasis>
., Palmdale area, 1ɗ, 
<date id="FFFF1053FFDE421CA6CCCF40971A6404" box="[684,821,604,630]" pageId="9" pageNumber="10" value="1964-07-10">10.Jul.1964</date>
(
<collectionCode id="ED50AE56FFDE421CA72BCF4097896404" LSID="urn:lsid:biocol.org:col:34802" box="[843,934,604,630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="10">LACM</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA7A2CF4090646407" box="[962,1099,604,629]" italics="true" pageId="9" pageNumber="10">Merced Co</emphasis>
., Los Banos, 1Ψ, 
<date id="FFFF1053FFDE421CA568CF9895B464EC" box="[264,411,644,670]" pageId="9" pageNumber="10" value="1948-10-22">22.Oct.1948</date>
, coll. H.R. Greenfield (
<collectionCode id="ED50AE56FFDE421CA6DACF98972E64EC" box="[698,769,644,670]" pageId="9" pageNumber="10">CISC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA778CF9897A664EF" box="[792,905,644,669]" italics="true" pageId="9" pageNumber="10">Mono Co</emphasis>
., 1ɗ, 
<date id="FFFF1053FFDE421CA7B6CF98904E64EC" box="[982,1121,644,670]" pageId="9" pageNumber="10" value="1969-07-14">14.Jul.1969</date>
, coll. O. Shields (
<collectionCode id="ED50AE56FFDE421CA570CFB0954464B4" LSID="urn:lsid:biocol.org:col:34802" box="[272,363,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="9" pageNumber="10">LACM</collectionCode>
); San Luis Obispo Co., 
<quantity id="4CB99B76FFDE421CA6FBCFB096FA64B4" box="[667,725,684,710]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="9" pageNumber="10" unit="mi" value="4.0">4 mi</quantity>
S. Shandon, 1Ψ, 
<date id="FFFF1053FFDE421CA7D6CFB0906564B4" box="[950,1098,684,710]" pageId="9" pageNumber="10" value="1979-10-10">10.Oct.1979</date>
, coll. R.J. Skalski (
<collectionCode id="ED50AE56FFDE421CA570CFC89547649C" box="[272,360,724,750]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA51CCFC89621649F" box="[380,526,724,749]" italics="true" pageId="9" pageNumber="10">Siskiyou Co.</emphasis>
, Gazelle, 1ɗ, 
<date id="FFFF1053FFDE421CA6D3CFC89711649C" box="[691,830,724,750]" pageId="9" pageNumber="10" value="1978-07-29">29.Jul.1978</date>
, coll. F.D. Horn (
<collectionCode id="ED50AE56FFDE421CA068CFC8904F649C" box="[1032,1120,724,750]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<quantity id="4CB99B76FFDE421CA013CFC89087649C" box="[1139,1192,724,750]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="9" pageNumber="10" unit="mi" value="5.0">5 mi</quantity>
N Gazelle, 1ɗ, 
<date id="FFFF1053FFDE421CA559CFE095FA6564" box="[313,469,764,790]" pageId="9" pageNumber="10" value="1980-08-13">13.Aug.1980</date>
, 1ɗ, 
<date id="FFFF1053FFDE421CA672CFE096B26564" box="[530,669,764,790]" pageId="9" pageNumber="10" value="1981-07-16">16.Jul.1981</date>
, coll. F.D. Horn (
<collectionCode id="ED50AE56FFDE421CA70ACFE097926564" box="[874,957,764,790]" pageId="9" pageNumber="10">CDFA</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA7B1CFE090636567" box="[977,1100,764,789]" italics="true" pageId="9" pageNumber="10">Solano Co</emphasis>
., Cold Canyon, 
<quantity id="4CB99B76FFDE421CA16DCFE09502654C" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="10" unit="km" value="19.0">19 km</quantity>
NNW Vacaville, 2ɗ, 
<date id="FFFF1053FFDE421CA623CE3896E8654C" box="[579,711,804,830]" pageId="9" pageNumber="10" value="1992-06-30">30Jun1992</date>
, coll. B.L. Fisher (
<collectionCode id="ED50AE56FFDE421CA7D6CE389021654C" box="[950,1038,804,830]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); Cold Canyon, 
<quantity id="4CB99B76FFDE421CA0BCCE389104654C" box="[1244,1323,804,830]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="10" unit="km" value="19.0">19 km</quantity>
NNW Vacaville, 2ɗ, 
<date id="FFFF1053FFDE421CA664CE5096A86514" box="[516,647,844,870]" pageId="9" pageNumber="10" value="1992-06-28">28Jun1992</date>
, coll. P.S. Ward (
<collectionCode id="ED50AE56FFDE421CA734CE5097836514" box="[852,940,844,870]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); Cold Canyon, 
<quantity id="4CB99B76FFDE421CA00DCE50908C6514" box="[1133,1187,844,870]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="9" pageNumber="10" unit="mi" value="9.0">9 mi</quantity>
W Winers, 3ɗ, 
<date id="FFFF1053FFDE421CA524CE6895FF65FC" box="[324,464,884,910]" pageId="9" pageNumber="10" value="1992-06-22">22.Jun.1992</date>
– 
<date id="FFFF1053FFDE421CA581CE68964A65FC" box="[481,613,884,910]" pageId="9" pageNumber="10" value="1993-06-20">20Jun1993</date>
, coll. J.A. De Benedictis (
<collectionCode id="ED50AE56FFDE421CA7ABCE68900C65FC" box="[971,1059,884,910]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA022CE6890E165FF" box="[1090,1230,884,909]" italics="true" pageId="9" pageNumber="10">Solano Co.</emphasis>
, Gates Canyon, 1ɗ, 
<date id="FFFF1053FFDE421CA5C4CE80960065C4" box="[420,559,924,950]" pageId="9" pageNumber="10" value="1980-07-21">21.Jul.1980</date>
, coll. J.B. Johnson (
<collectionCode id="ED50AE56FFDE421CA77DCE80975A65C4" box="[797,885,924,950]" pageId="9" pageNumber="10">EMUS</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA7E9CE80900465C7" box="[905,1067,924,949]" italics="true" pageId="9" pageNumber="10">Stanislaus Co</emphasis>
., Del Puerto Canyon, 
<quantity id="4CB99B76FFDE421CA568CED8951365AC" box="[264,316,964,990]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="9" pageNumber="10" unit="mi" value="5.0">5 mi</quantity>
W Patterson, 1ɗ, 
<date id="FFFF1053FFDE421CA673CED896A765AC" box="[531,648,964,990]" pageId="9" pageNumber="10" value="2005-06-09">9Jun2005</date>
, coll. K.A. Williams (
<collectionCode id="ED50AE56FFDE421CA7E7CED897CA65AC" box="[903,997,964,990]" pageId="9" pageNumber="10">KAWC</collectionCode>
); Del Puerto Canyon, 
<quantity id="4CB99B76FFDE421CA087CED8910465AC" box="[1255,1323,964,990]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="9" pageNumber="10" unit="mi" value="20.0">20 mi</quantity>
W Payson, 2ɗ, 
<date id="FFFF1053FFDE421CA5B3CEF0965F6274" box="[467,624,1004,1030]" pageId="9" pageNumber="10" value="2001-05-27">27.May.2001</date>
, coll. K.A. &amp; E.E. Williams (
<collectionCode id="ED50AE56FFDE421CA790CEF090616274" box="[1008,1102,1004,1030]" pageId="9" pageNumber="10">KAWC</collectionCode>
); La Grange, 8ɗ, 
<date id="FFFF1053FFDE421CA568C90895A6625C" box="[264,393,1044,1070]" pageId="9" pageNumber="10" value="1955-06-07">7.Jun.1955</date>
– 
<date id="FFFF1053FFDE421CA5FAC9089631625C" box="[410,542,1044,1070]" pageId="9" pageNumber="10" value="1970-09-08">8.Sep.1970</date>
, coll. R.P. Allen (
<collectionCode id="ED50AE56FFDE421CA772C9089745625C" box="[786,874,1044,1070]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); LaGrange, 1ɗ, 
<date id="FFFF1053FFDE421CA030C90890C4625C" box="[1104,1259,1044,1070]" pageId="9" pageNumber="10" value="1961-08-30">30.Aug.1961</date>
, 1ɗ, 
<date id="FFFF1053FFDE421CA568C92095536224" box="[264,380,1084,1110]" pageId="9" pageNumber="10" value="1968-06-06">6Jun1968</date>
, 2ɗ, 
<date id="FFFF1053FFDE421CA5A2C92096616224" box="[450,590,1084,1110]" pageId="9" pageNumber="10" value="1975-08-04">4.Aug.1975</date>
– 
<date id="FFFF1053FFDE421CA63FC92096CE6224" box="[607,737,1084,1110]" pageId="9" pageNumber="10" value="1975-09-05">5.Sep.1975</date>
, coll. R.P. Allen (
<collectionCode id="ED50AE56FFDE421CA7A0C920903C6224" box="[960,1043,1084,1110]" pageId="9" pageNumber="10">CDFA</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA04CC92090816227" box="[1068,1198,1084,1109]" italics="true" pageId="9" pageNumber="10">Trinity Co.</emphasis>
, 
<quantity id="4CB99B76FFDE421CA0A2C92091256224" box="[1218,1290,1084,1110]" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="9" pageNumber="10" unit="mi" value="25.0">25 mi</quantity>
N Weaverville, 1Ψ, 
<date id="FFFF1053FFDE421CA5B6C978965E620C" box="[470,625,1124,1150]" pageId="9" pageNumber="10" value="1980-08-12">12.Aug.1980</date>
, coll. C.Y. Kitayama (
<collectionCode id="ED50AE56FFDE421CA717C97897E0620C" box="[887,975,1124,1150]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA784C9789070620F" box="[996,1119,1124,1149]" italics="true" pageId="9" pageNumber="10">Tulare Co.</emphasis>
, Poso creek near Posey, 1ɗ, 
<date id="FFFF1053FFDE421CA5F1C990961262D4" box="[401,573,1164,1190]" pageId="9" pageNumber="10" value="2000-03-23">
23. 
<collectionCode id="ED50AE56FFDE421CA5D2C99095D462D4" box="[434,507,1164,1190]" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="9" pageNumber="10">MAR</collectionCode>
.2000
</date>
, coll. T.D. Waite (
<collectionCode id="ED50AE56FFDE421CA77DC990975A62D4" box="[797,885,1164,1190]" pageId="9" pageNumber="10">BYUC</collectionCode>
); 
<emphasis id="B935EA81FFDE421CA7ECC99097DF62D7" box="[908,1008,1164,1189]" italics="true" pageId="9" pageNumber="10">Yolo Co.</emphasis>
, 
<quantity id="4CB99B76FFDE421CA064C990901362D4" box="[1028,1084,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="9" pageNumber="10" unit="mi" value="4.0">4 mi</quantity>
SW Dunnigan, 1ɗ, 
<date id="FFFF1053FFDE421CA568C9A895B262BC" box="[264,413,1204,1230]" pageId="9" pageNumber="10" value="1959-09-15">15.Sep.1959</date>
, coll. B.N. Chaniotis (
<collectionCode id="ED50AE56FFDE421CA6C0C9A896D762BC" box="[672,760,1204,1230]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<quantity id="4CB99B76FFDE421CA76CC9A8976F62BC" box="[780,832,1204,1230]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="9" pageNumber="10" unit="mi" value="8.0">8 mi</quantity>
NW Winters, 1ɗ, 
<date id="FFFF1053FFDE421CA078C9A8908262BC" box="[1048,1197,1204,1230]" pageId="9" pageNumber="10" value="1959-09-10">10.Sep.1959</date>
, coll. B.N. Chaniotis (
<collectionCode id="ED50AE56FFDE421CA5F1C9C095C66284" box="[401,489,1244,1270]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); Winters, 1ɗ, 
<date id="FFFF1053FFDE421CA6D3C9C0971A6284" box="[691,821,1244,1270]" pageId="9" pageNumber="10" value="1962-06-27">27Jun1962</date>
, coll. P.M. Marsh (
<collectionCode id="ED50AE56FFDE421CA057C9C090A06284" box="[1079,1167,1244,1270]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<quantity id="4CB99B76FFDE421CA0CCC9C090C66284" box="[1196,1257,1244,1270]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="9" pageNumber="10" unit="mi" value="8.0">8 mi</quantity>
NW Winters, 4ɗ, 
<date id="FFFF1053FFDE421CA5D0C818961B636C" box="[432,564,1284,1310]" pageId="9" pageNumber="10" value="1959-09-02">2.Sep.1959</date>
– 
<date id="FFFF1053FFDE421CA625C81896FF636C" box="[581,720,1284,1310]" pageId="9" pageNumber="10" value="1959-08-06">6.Aug.1959</date>
, coll. J. Fowler (
<collectionCode id="ED50AE56FFDE421CA7CDC818902A636C" box="[941,1029,1284,1310]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
); 
<quantity id="4CB99B76FFDE421CA07FC8189075636C" box="[1055,1114,1284,1310]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="9" pageNumber="10" unit="mi" value="7.0">7 mi</quantity>
W Zamora, 1ɗ, 
<date id="FFFF1053FFDE421CA568C83095A46334" box="[264,395,1324,1350]" pageId="9" pageNumber="10" value="1959-06-31">31Jun1959</date>
, coll. J. Fowler (
<collectionCode id="ED50AE56FFDE421CA63AC830969D6334" box="[602,690,1324,1350]" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="9" pageNumber="10">UCDC</collectionCode>
). 
<emphasis id="B935EA81FFDE421CA6A8C83097096334" bold="true" box="[712,806,1324,1350]" pageId="9" pageNumber="10">Nevada</emphasis>
: 
<emphasis id="B935EA81FFDE421CA759C83097E96337" box="[825,966,1324,1349]" italics="true" pageId="9" pageNumber="10">Washoe Co.</emphasis>
, Red Rock, 1ɗ, 
<date id="FFFF1053FFDE421CA0F4C83091096334" box="[1172,1318,1324,1350]" pageId="9" pageNumber="10" value="1969-06-20">20.Jun.1969</date>
, coll. R.C. Bechtel (
<collectionCode id="ED50AE56FFDE421CA589C848966B631C" box="[489,580,1364,1390]" httpUri="http://grbio.org/cool/c9nj-8iig" name="Nevada State Department of Agriculture" pageId="9" pageNumber="10">NVDA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6518FFDE421CA568C8B897C0613C" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BFE3693FFDE421CA568C8B8954163CF" blockId="9.[264,1323,1444,1870]" box="[264,366,1444,1469]" pageId="9" pageNumber="10">
<emphasis id="B935EA81FFDE421CA568C8B8954163CF" box="[264,366,1444,1469]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
</paragraph>
<paragraph id="8BFE3693FFDE421CA558C8D0959C6044" blockId="9.[264,1323,1444,1870]" pageId="9" pageNumber="10">The sex association is based on collection data, distribution, and similarities in coloration and morphology. Furthermore, the female resembles the other females in this species­group.</paragraph>
<paragraph id="8BFE3693FFDE421CA558CB5897C0613C" blockId="9.[264,1323,1444,1870]" pageId="9" pageNumber="10">Retaining subspecies designations is unwarranted. Study of the genitalia has revealed no distinguishable differences. Furthermore, the two subspecies have overlapping ranges. In the males, coloration of the pubescence can vary from yellow or golden yellow to brilliant scarlet or carmine­fulvous and the integumental coloration can vary from orange or brownish­orange to piceous. Head sculpture also varies from having almost confluent, coarse punctures to shallow punctures that range from moderately separated to almost contiguous. Females vary in setal coloration similarly to males.</paragraph>
</subSubSection>
</treatment>
</document>