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<document id="86EFDA88427F6EFAAFEB8256BD9438C3" ID-DOI="10.5324/cjcr.v0i30.2240" ID-ISSN="2387-5372" ID-Zenodo-Dep="7987298" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1685469670408" checkinUser="felipe" docAuthor="Epler, John H." docDate="2017" docId="03E2824B3D7DB923143C6C84FE35FDF4" docLanguage="en" docName="ChiroJChiroRes.30.4-18.pdf" docOrigin="CHIRONOMUS Journal of Chironomidae Research 30 (30)" docSource="https://www.ntnu.no/ojs/index.php/chironomus/article/view/2240" docStyle="DocumentStyle:B21B57A70ED88EDAB7971C4BDED0DA62.1:ChiroJChiroRes.2015-.journal_article" docStyleId="B21B57A70ED88EDAB7971C4BDED0DA62" docStyleName="ChiroJChiroRes.2015-.journal_article" docStyleVersion="1" docTitle="Riethia Kieffer 1917" docType="treatment" docVersion="1" lastPageNumber="10" masterDocId="FFDBFA333D7BB92417046A28FFD5FF8F" masterDocTitle="AN ANNOTATED PRELIMINARY LIST OF THE CHIRONOMIDAE (DIPTERA) OF ZURQUÍ, COSTA RICA Abstract Introduction" masterLastPageNumber="18" masterPageNumber="4" pageNumber="10" updateTime="1685624128335" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AC6E9B2B80C1A0FDDA6EF6B8AF2FFF78">AN ANNOTATED PRELIMINARY LIST OF THE CHIRONOMIDAE (DIPTERA) OF ZURQUÍ, COSTA RICA Abstract Introduction</mods:title>
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<mods:namePart id="8B931ACA30EE30862EF9903683D80449">Epler, John H.</mods:namePart>
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<treatment id="03E2824B3D7DB923143C6C84FE35FDF4" LSID="urn:lsid:plazi:treatment:03E2824B3D7DB923143C6C84FE35FDF4" httpUri="http://treatment.plazi.org/id/03E2824B3D7DB923143C6C84FE35FDF4" lastPageId="7" lastPageNumber="10" pageId="6" pageNumber="10">
<subSubSection id="C35160D63D7DB922143C6C84FC5DF94B" box="[824,904,1708,1732]" pageId="6" pageNumber="10" type="nomenclature">
<paragraph id="8BF4335D3D7DB922143C6C84FC5DF94B" blockId="6.[824,1361,1708,2021]" box="[824,904,1708,1732]" pageId="6" pageNumber="10">
<taxonomicName id="4C4B48DE3D7DB922143C6C84FC5DF94B" ID-CoL="79ZG" ID-ENA="391820" authorityName="Kieffer" authorityYear="1917" box="[824,904,1708,1732]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7DB922143C6C84FC5DF94B" bold="true" box="[824,904,1708,1732]" italics="true" pageId="6" pageNumber="10">Riethia</emphasis>
</taxonomicName>
</paragraph>
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<subSubSection id="C35160D63D7DB92314946C85FE35FDF4" lastPageId="7" lastPageNumber="11" pageId="6" pageNumber="10" type="description">
<paragraph id="8BF4335D3D7DB92314946C85FE35FDF4" blockId="6.[824,1361,1708,2021]" lastBlockId="7.[227,764,227,635]" lastPageId="7" lastPageNumber="11" pageId="6" pageNumber="10">
- Fifteen male specimens of a genus from the putative tribe
<taxonomicName id="4C4B48DE3D7DB92214F56CE5FB15F96A" box="[1009,1216,1741,1765]" pageId="6" pageNumber="10" rank="tribe" tribe="Pseudochironomini">Pseudochironomini</taxonomicName>
were collected; this genus is almost certainly
<taxonomicName id="4C4B48DE3D7DB922139D6CC5FB33F88A" authorityName="Kieffer" authorityYear="1917" box="[1177,1254,1773,1797]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7DB922139D6CC5FB33F88A" box="[1177,1254,1773,1797]" italics="true" pageId="6" pageNumber="10">Riethia</emphasis>
</taxonomicName>
, but there is a small chance they could be
<taxonomicName id="4C4B48DE3D7DB92213926D26FB37F8AA" box="[1174,1250,1806,1829]" class="Insecta" family="Chironomidae" genus="Manoa" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7DB92213926D26FB37F8AA" box="[1174,1250,1806,1829]" italics="true" pageId="6" pageNumber="10">Manoa</emphasis>
</taxonomicName>
. Males of the two genera cannot at this time be separated utilizing morphological characters; only females and pupae may be separated at the generic level (
<bibRefCitation id="EFDA4EAC3D7DB922143B6DA5FB92F82A" author="Jacobsen, R. E. &amp; Perry, S. A." box="[831,1095,1933,1957]" pageId="6" pageNumber="10" pagination="314 - 325" refId="ref9227" refString="Jacobsen, R. E. &amp; Perry, S. A. 2002. A new species of Manoa (Diptera: Chironomidae) from Everglades National Park. - Journal of the North American Benthological Society 21: 314 - 325." type="journal article" year="2002">Jacobsen &amp; Perry 2002</bibRefCitation>
;
<bibRefCitation id="EFDA4EAC3D7DB92213506DA5FCA7F84A" author="Trivinho-Strixino, S. &amp; Roque, F. O. &amp; Cranston, P. S." pageId="6" pageNumber="10" pagination="247 - 259" refId="ref10711" refString="Trivinho-Strixino, S., Roque, F. O. &amp; Cranston, P. S. 2009. Redescription of Riethia truncatocaudata (Edwards, 1931) (Diptera: Chironomidae), with description of female, pupa and larva and generic diagnosis for Riethia. - Aquatic Insects 31: 247 - 259." type="journal article" year="2009">Trivinho-Strixino et al. 2009</bibRefCitation>
). Im calling the Zurquí specimens
<taxonomicName id="4C4B48DE3D7DB92212006D85FA84F84A" authorityName="Kieffer" authorityYear="1917" box="[1284,1361,1965,1989]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7DB92212006D85FA84F84A" box="[1284,1361,1965,1989]" italics="true" pageId="6" pageNumber="10">Riethia</emphasis>
</taxonomicName>
based on their close resemblance to
<taxonomicName id="4C4B48DE3D7DB92313C96DE6FE63FF74" authority="(Edwards)" baseAuthorityName="Edwards" baseAuthorityYear="1931" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" lastPageId="7" lastPageNumber="11" order="Diptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="species" species="truncatocaudata">
<emphasis id="B93FEF4F3D7DB92313C96DE6FEECFF74" italics="true" lastPageId="7" lastPageNumber="11" pageId="6" pageNumber="10">R. truncatocaudata</emphasis>
(Edwards)
</taxonomicName>
as redescribed in
<bibRefCitation id="EFDA4EAC3D7CB92315926ACBFE1DFE94" author="Trivinho-Strixino, S. &amp; Roque, F. O. &amp; Cranston, P. S." pageId="7" pageNumber="11" pagination="247 - 259" refId="ref10711" refString="Trivinho-Strixino, S., Roque, F. O. &amp; Cranston, P. S. 2009. Redescription of Riethia truncatocaudata (Edwards, 1931) (Diptera: Chironomidae), with description of female, pupa and larva and generic diagnosis for Riethia. - Aquatic Insects 31: 247 - 259." type="journal article" year="2009">Trivinho-Strixino et al. (2009)</bibRefCitation>
and
<taxonomicName id="4C4B48DE3D7CB92315066B2CFD22FEB4" authority="Neubern, Trivinho-Strixino &amp; Silva (Neubern et al. 2011)" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="11" phylum="Arthropoda" rank="species" species="manauara">
<emphasis id="B93FEF4F3D7CB92315066B2CFD45FE94" box="[514,656,260,283]" italics="true" pageId="7" pageNumber="11">R. manauara</emphasis>
Neubern, Trivinho-Strixino &amp; Silva (
<bibRefCitation id="EFDA4EAC3D7CB923150A6B0BFD25FEB4" author="Neubern, C. S. O. &amp; Trivinho-Strixino, S. &amp; Silva, F. L." box="[526,752,291,315]" pageId="7" pageNumber="11" pagination="595 - 599" refId="ref9664" refString="Neubern, C. S. O., Trivinho-Strixino, S. &amp; Silva, F. L. 2011. Riethia manauara n. sp., an Amazonian chironomid (Diptera: Chironomidae) from Brazil. - Neotropical Entomology 40: 595 - 599." type="journal article" year="2011">Neubern et al. 2011</bibRefCitation>
)
</taxonomicName>
; the medial apices of the inferior volsellae of all three species have large, flattened, apically pectinate setae directed mediad (and dorsad in
<taxonomicName id="4C4B48DE3D7CB92315AC6BACFEB2FE34" baseAuthorityName="Edwards" baseAuthorityYear="1931" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="11" phylum="Arthropoda" rank="species" species="truncatocaudata">
<emphasis id="B93FEF4F3D7CB92315AC6BACFEB2FE34" italics="true" pageId="7" pageNumber="11">R. truncatocaudata</emphasis>
</taxonomicName>
). Such setae are unknown in
<taxonomicName id="4C4B48DE3D7CB92315AD6B8CFD20FE34" box="[681,757,420,443]" class="Insecta" family="Chironomidae" genus="Manoa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B93FEF4F3D7CB92315AD6B8CFD20FE34" box="[681,757,420,443]" italics="true" pageId="7" pageNumber="11">Manoa</emphasis>
</taxonomicName>
. Note that the legs of the Zurquí specimens are unbanded and there are other differences, especially in the superior volsella; thus, they do not represent
<taxonomicName id="4C4B48DE3D7CB92317E7680CFE60FDB4" baseAuthorityName="Edwards" baseAuthorityYear="1931" box="[227,437,547,571]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="11" phylum="Arthropoda" rank="species" species="truncatocaudata">
<emphasis id="B93FEF4F3D7CB92317E7680CFE60FDB4" box="[227,437,547,571]" italics="true" pageId="7" pageNumber="11">R. truncatocaudata</emphasis>
</taxonomicName>
. Likewise, differences in the genitalia indicate that
<emphasis id="B93FEF4F3D7CB92316D4686CFE35FDD4" box="[464,480,580,603]" italics="true" pageId="7" pageNumber="11">
<collectionCode id="ED5AAB983D7CB92316D4686CFE35FDD4" box="[464,480,580,603]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="11">R</collectionCode>
</emphasis>
. sp. ZUR-1 is not conspecific with
<taxonomicName id="4C4B48DE3D7CB9231649684CFE0CFDF4" box="[333,473,612,635]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="11" phylum="Arthropoda" rank="species" species="manauara">
<emphasis id="B93FEF4F3D7CB9231649684CFE0CFDF4" box="[333,473,612,635]" italics="true" pageId="7" pageNumber="11">R. manauara</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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