824 lines
81 KiB
XML
824 lines
81 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5678983" ID-GBIF-Dataset="d2f92698-5e17-472a-b73b-90c76705b4ce" ID-GBIF-Taxon="133186814" ID-ISSN="0936-9902" ID-Zenodo-Dep="5678983" checkinTime="1504771707859" checkinUser="plazi" docAuthor="Maurice Kottelat" docDate="2017" docId="03BE5B458638FFAFFF46FD87FE98FE8F" docLanguage="en" docName="2017 Schistura titan.pdf" docOrigin="Ichthyol. Explor. Freshwaters 28 (1)" docStyle="DocumentStyle{}" docTitle="Schistura titan Kottelat, 2017, new species" docType="treatment" docVersion="9" lastPageId="7" lastPageNumber="71" masterDocId="FF87233D863AFFA8FF8FFFE8FF9CFFC7" masterDocTitle="Schistura titan, a new species of loach from Dakchung Plateau, southern Laos (Teleostei: Nemacheilidae)" masterLastPageNumber="83" masterPageNumber="65" pageId="2" pageNumber="67" updateTime="1636704778628" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Schistura titan, a new species of loach from Dakchung Plateau, southern Laos (Teleostei: Nemacheilidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Maurice Kottelat</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Ichthyol. Explor. Freshwaters</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2017</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>65</mods:start>
|
||
<mods:end>83</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.5281/zenodo.886884</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">d2f92698-5e17-472a-b73b-90c76705b4ce</mods:identifier>
|
||
<mods:identifier type="ISSN">0936-9902</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">886884</mods:identifier>
|
||
<mods:identifier type="ZooBank">C1E005DB-59E5-47AA-A0B5-E01F15A5E21F</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5678983" ID-GBIF-Taxon="133186814" ID-Zenodo-Dep="5678983" LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F" httpUri="http://treatment.plazi.org/id/03BE5B458638FFAFFF46FD87FE98FE8F" lastPageId="7" lastPageNumber="71" pageId="2" pageNumber="67">
|
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<subSubSection box="[201,508,623,645]" pageId="2" pageNumber="67" type="nomenclature">
|
||
<paragraph blockId="2.[201,508,623,674]" box="[201,508,623,645]" pageId="2" pageNumber="67">
|
||
<heading bold="true" box="[201,508,623,645]" centered="true" fontSize="9" level="1" pageId="2" pageNumber="67" reason="4">
|
||
<emphasis bold="true" box="[201,508,623,645]" pageId="2" pageNumber="67">
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[201,362,623,645]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="67" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis bold="true" box="[201,362,623,645]" italics="true" pageId="2" pageNumber="67">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[375,508,623,645]" pageId="2" pageNumber="67" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[318,391,652,674]" pageId="2" pageNumber="67" type="description">
|
||
<paragraph blockId="2.[201,508,623,674]" box="[318,391,652,674]" pageId="2" pageNumber="67">
|
||
<heading box="[318,391,652,674]" centered="true" fontSize="9" level="3" pageId="2" pageNumber="67" reason="4">
|
||
(
|
||
<figureCitation box="[325,382,652,674]" captionStart="Fig. 1" captionStartId="1.[129,161,1376,1395]" captionTargetBox="[148,1160,181,1327]" captionTargetId="figure@1.[129,1187,716,1116]" captionTargetPageId="1" captionText="Fig. 1. Schistura titan; Laos: Xekong Province: Dakchung Plateau: Houay Champong; aı MHNG 2767.057, holotypeı 74.8 mm SL; bı CMK 22344, paratype, 95.0 mm SL (distal part of dorsal fin and upper part of caudal fin damaged and regrown)." httpUri="https://zenodo.org/record/886886/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
|
||
)
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="67" type="materials_examined">
|
||
<paragraph blockId="2.[98,611,709,904]" pageId="2" pageNumber="67">
|
||
<materialsCitation ID-GBIF-Occurrence="1612095192" collectingDate="2011-05-22" collectionCode="MHNG" collectorName="M. Kottelat" country="Laos" county="Houay Champong" elevation="1170" latitude="15.573334" location="Houay Xe Kaman" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" pageId="2" pageNumber="67" specimenCode="MHNG 2767.057, 74.8" specimenCount="1" stateProvince="Xekong Province" typeStatus="holotype">
|
||
<emphasis bold="true" box="[98,207,710,731]" pageId="2" pageNumber="67">
|
||
<typeStatus box="[98,202,710,731]" pageId="2" pageNumber="67">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode box="[219,455,709,731]" pageId="2" pageNumber="67">
|
||
MHNG 2767.057,
|
||
<quantity box="[413,503,709,731]" metricMagnitude="-2" metricUnit="m" metricValue="7.4799999999999995" pageId="2" pageNumber="67" unit="mm" value="74.8">74.8 mm</quantity>
|
||
</specimenCode>
|
||
SL;
|
||
<collectingCountry box="[554,607,710,731]" name="Laos" pageId="2" pageNumber="67">Laos</collectingCountry>
|
||
:
|
||
<collectingRegion box="[98,293,738,760]" country="Laos" name="Xekong" pageId="2" pageNumber="67">Xekong Province</collectingRegion>
|
||
:
|
||
<collectingCounty box="[309,519,738,760]" pageId="2" pageNumber="67">Houay Champong</collectingCounty>
|
||
, in
|
||
<collectingMunicipality pageId="2" pageNumber="67">Ban Dak</collectingMunicipality>
|
||
, immediately upstream of confluence with
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC888638FFAAFFEDFCF4FEB8FCF6" box="[98,292,796,817]" country="Laos" county="Houay Champong" latitude="15.573334" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" name="Houay Xe Kaman" pageId="2" pageNumber="67" stateProvince="Xekong Province">Houay Xe Kaman</location>
|
||
,
|
||
<quantity box="[303,369,796,818]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="67" unit="km" value="15.0">15 km</quantity>
|
||
from
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC888638FFAAFE3DFCF4FDBEFCF5" box="[434,546,796,818]" country="Laos" county="Houay Champong" latitude="15.573334" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" name="Dakchung" pageId="2" pageNumber="67" stateProvince="Xekong Province">Dakchung</location>
|
||
,
|
||
<quantity box="[558,611,796,818]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="67" unit="km" value="4.0">4 km</quantity>
|
||
after
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC888638FFAAFF14FCD0FF6AFC89" box="[155,246,824,846]" country="Laos" county="Houay Champong" latitude="15.573334" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" name="Dak Noi" pageId="2" pageNumber="67" stateProvince="Xekong Province">Dak Noi</location>
|
||
and
|
||
<quantity box="[302,374,824,846]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="67" unit="km" value="4.4">4.4 km</quantity>
|
||
before
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC888638FFAAFE48FCD1FDC3FC89" box="[455,607,824,846]" country="Laos" county="Houay Champong" latitude="15.573334" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" name="Ban Sang Mai" pageId="2" pageNumber="67" stateProvince="Xekong Province">Ban Sang Mai</location>
|
||
,
|
||
<geoCoordinate box="[98,217,853,875]" direction="north" orientation="latitude" pageId="2" pageNumber="67" precision="15" value="15.573334">15°34'24" N</geoCoordinate>
|
||
<geoCoordinate box="[225,352,853,875]" direction="east" orientation="longitude" pageId="2" pageNumber="67" precision="15" value="107.2175">107°13'03"E</geoCoordinate>
|
||
,
|
||
<elevation box="[364,477,853,875]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="67" unit="m" value="1170.0">
|
||
<quantity box="[364,439,853,875]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="67" unit="m" value="1170.0">1170 m</quantity>
|
||
asl
|
||
</elevation>
|
||
;
|
||
<collectorName box="[489,610,853,875]" pageId="2" pageNumber="67">M. Kottelat</collectorName>
|
||
et al.,
|
||
<collectingDate box="[162,298,882,904]" pageId="2" pageNumber="67" value="2011-05-22">
|
||
<date box="[162,298,882,904]" pageId="2" pageNumber="67" value="2011-05-22">22 May 2011</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="2.[98,611,940,990]" pageId="2" pageNumber="67">
|
||
<materialsCitation ID-GBIF-Occurrence="1612095193" collectingDate="2011-05-22" collectionCode="MHNG" collectorName="M. Kottelat" country="Laos" county="Houay Champong" elevation="1170" latitude="15.573334" location="Houay Xe Kaman" longLatPrecision="21" longitude="107.2175" municipality="Ban Dak" pageId="2" pageNumber="67" specimenCount="1" stateProvince="Xekong Province" typeStatus="paratype">
|
||
<emphasis bold="true" box="[98,200,940,961]" pageId="2" pageNumber="67">
|
||
<typeStatus box="[98,196,940,961]" pageId="2" pageNumber="67">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<accessionNumber box="[212,337,940,961]" httpUri="http://www.uniprot.org/uniprot/CMK22344" pageId="2" pageNumber="67">CMK 22344</accessionNumber>
|
||
, 1, 95.0 mm SL; collected with holotype.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="67" type="diagnosis">
|
||
<paragraph blockId="2.[98,611,1026,1480]" pageId="2" pageNumber="67">
|
||
<emphasis bold="true" box="[98,215,1026,1048]" pageId="2" pageNumber="67">Diagnosis.</emphasis>
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[227,374,1026,1048]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="67" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[227,374,1026,1048]" italics="true" pageId="2" pageNumber="67">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all the species of the genus in the Mekong drainage by the presence of a posterior chamber of the air bladder in the abdominal cavity. Besides, the following combination of characters is unique in the Mekong drainage: 8 + 7 branched caudal-fin rays; large size (up to at least
|
||
<quantity box="[426,498,1199,1221]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="67" unit="mm" value="95.0">95 mm</quantity>
|
||
SL); 17 or 18 bars on flank, meeting contralaterals on the back behind dorsal fin, not meeting, irregular or dissociated into small blotches in predorsal area; nostril flap pointed, reaching anterior margin of eye; 8
|
||
<emphasis bold="true" box="[159,163,1342,1354]" pageId="2" pageNumber="67">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[171,178,1351,1363]" pageId="2" pageNumber="67">2</emphasis>
|
||
branched dorsal-fin rays; pelvic-fin origin behind dorsal-fin origin; black pattern at caudalfin base: faint black bar on lower
|
||
<emphasis bold="true" box="[436,440,1400,1412]" pageId="2" pageNumber="67">2</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[448,455,1408,1420]" pageId="2" pageNumber="67">3</emphasis>
|
||
, not extending on ventral crest and a faint blotch at base of upper simple and posterior three procurrent rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="68" pageId="2" pageNumber="67" type="description">
|
||
<paragraph blockId="2.[98,611,1516,1682]" lastBlockId="2.[644,1157,592,1679]" pageId="2" pageNumber="67">
|
||
<emphasis bold="true" box="[98,232,1516,1538]" pageId="2" pageNumber="67">Description.</emphasis>
|
||
See
|
||
<figureCitation box="[284,368,1516,1538]" captionStart="Fig. 1" captionStartId="1.[129,161,1376,1395]" captionTargetBox="[148,1160,181,1327]" captionTargetId="figure@1.[129,1187,716,1116]" captionTargetPageId="1" captionText="Fig. 1. Schistura titan; Laos: Xekong Province: Dakchung Plateau: Houay Champong; aı MHNG 2767.057, holotypeı 74.8 mm SL; bı CMK 22344, paratype, 95.0 mm SL (distal part of dorsal fin and upper part of caudal fin damaged and regrown)." httpUri="https://zenodo.org/record/886886/files/figure.png" pageId="2" pageNumber="67">Figure 1</figureCitation>
|
||
for general appearance and
|
||
<tableCitation box="[147,225,1544,1566]" captionStart="Table 1" captionStartId="3.[128,184,637,656]" captionTargetBox="[128,621,750,1666]" captionText="Table 1. Morphometric data of Schistura titan (MHNG 2767.057, holotype; CMK 22344, paratype). H, holotype; P, paratype." httpUri="http://table.plazi.org/id/DF68BADB8639FFABFF0FFD95FEB2FD03" pageId="2" pageNumber="67" tableUuid="DF68BADB8639FFABFF0FFD95FEB2FD03">Table 1</tableCitation>
|
||
for morphometric data of
|
||
<typeStatus box="[516,610,1544,1566]" pageId="2" pageNumber="67">holotype</typeStatus>
|
||
and
|
||
<typeStatus box="[152,252,1574,1595]" pageId="2" pageNumber="67">paratype</typeStatus>
|
||
. An elongate nemacheilid with body depth regularly increasing up to slightly in front of dorsal-fin origin. Behind dorsal fin, body depth almost uniform until caudal-fin base, where it is slightly increasing (in
|
||
<typeStatus box="[977,1073,592,613]" pageId="2" pageNumber="67">paratype</typeStatus>
|
||
). Dorsal profile with a small concavity between head and body. Head slightly depressed; body compressed anteriorly to very compressed posteriorly. Interorbital area convex. In lateral view, eye below dorsal profile of head. Cheeks not swollen. Snout rounded. Caudal peduncle 1.3-1.5 times longer than deep, uniform depth in
|
||
<typeStatus box="[947,1041,793,815]" pageId="2" pageNumber="67">holotype</typeStatus>
|
||
or increasing until caudal-fin base in
|
||
<typeStatus box="[925,1023,822,843]" pageId="2" pageNumber="67">paratype</typeStatus>
|
||
. In
|
||
<typeStatus box="[1056,1153,822,843]" pageId="2" pageNumber="67">paratype</typeStatus>
|
||
, high dorsal crest on posterior
|
||
<emphasis bold="true" box="[977,981,850,862]" pageId="2" pageNumber="67">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[990,997,859,871]" pageId="2" pageNumber="67">4</emphasis>
|
||
of post-dorsal area and high ventral crest on posterior half of caudal peduncle; crests present in
|
||
<typeStatus box="[1019,1113,908,930]" pageId="2" pageNumber="67">holotype</typeStatus>
|
||
but less developed. Dorsal crest continuous with upper margin of caudal fin, supported by procurrent rays. Largest recorded size 95.0 mm SL.
|
||
</paragraph>
|
||
<paragraph blockId="2.[644,1157,592,1679]" pageId="2" pageNumber="67">
|
||
Dorsal fin with 4 unbranched and 8
|
||
<emphasis bold="true" box="[1135,1139,1023,1035]" pageId="2" pageNumber="67">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[1149,1156,1031,1043]" pageId="2" pageNumber="67">2</emphasis>
|
||
branched rays; distal margin slightly convex, second branched ray longest. Pectoral fin with 1 unbranched and 11 branched rays, rounded, reaching about halfway of distance to pelvicfin base. Pelvic fin with 1 unbranched and 7 branched rays, reaching to or almost to anus, rounded; posterior margin convex; origin below base of first branched dorsal-fin ray; axillary lobe present, small, entirely free. Anus situated about 1-1
|
||
<emphasis bold="true" box="[684,688,1311,1323]" pageId="2" pageNumber="67">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[697,704,1319,1331]" pageId="2" pageNumber="67">2</emphasis>
|
||
eye diameter in front of anal fin, distance between anus and anal-fin origin about 2 times in distance between anus and pelvic-fin base. Anal fin with 3 unbranched and 5
|
||
<emphasis bold="true" box="[1028,1032,1397,1409]" pageId="2" pageNumber="67">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[1040,1047,1406,1418]" pageId="2" pageNumber="67">2</emphasis>
|
||
branched rays; distal margin convex. Caudal fin with 8 + 7 branched rays in
|
||
<typeStatus box="[848,943,1456,1478]" pageId="2" pageNumber="67">holotype</typeStatus>
|
||
(6 +
|
||
<quantity box="[997,1042,1456,1478]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="67" unit="in" value="7.0">7 in</quantity>
|
||
<typeStatus box="[1054,1152,1456,1477]" pageId="2" pageNumber="67">paratype</typeStatus>
|
||
, but the upper part of the fin was damaged and rays are regrown and deformed); about 9 dorsal and about 6 ventral procurrent rays (difficult to count with transmitted light); emarginate, lobes rounded, subequal.
|
||
</paragraph>
|
||
<paragraph blockId="2.[644,1157,592,1679]" lastBlockId="3.[128,641,160,587]" lastPageId="3" lastPageNumber="68" pageId="2" pageNumber="67">Body entirely scaled, except between bases of pectoral fins in large specimen; scales deeply embedded. Lateral line complete, with 89-98 pores. Cephalic lateral line system with 6 or 7 supraorbital, 4 +11-13 infraorbital, 9 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
|
||
<paragraph blockId="3.[128,641,160,587]" lastBlockId="3.[675,1187,162,328]" pageId="3" pageNumber="68">
|
||
Anterior naris pierced in front side of a long pointed flap-like tube reaching to anterior margin of eye. Posterior naris adjacent to anterior one. Mouth arched, gape about 2.1-2.5 times wider than long (
|
||
<figureCitation box="[250,310,392,415]" captionStart="Fig. 2" captionStartId="2.[644,676,492,511]" captionTargetBox="[668,1134,192,474]" captionTargetId="figure@2.[644,1157,159,474]" captionTargetPageId="2" captionText="Fig. 2. Schistura titan, CMK 22344, paratype, 95.0 mm SL; ventral view of mouth." httpUri="https://zenodo.org/record/886888/files/figure.png" pageId="3" pageNumber="68">Fig. 2</figureCitation>
|
||
). Lips thick, fleshy. Upper lip without median notch, with several shallow furrows. Processus dentiformis present. Lower lip with median interruption; median part with a few deep furrows, lateral part smooth. Tip of lower jaw not exposed. A very shallow median concavity at tip of lower jaw. Inner rostral barbel reaching at most corner of mouth; outer one reaching anterior margin of eye. Maxillary barbel reaching at most about middle of postorbital area. Intestine with a bend immediately behind stomach (
|
||
<figureCitation box="[1113,1172,248,271]" captionStart="Fig. 3" captionStartId="4.[644,676,725,744]" captionTargetBox="[762,1038,159,707]" captionTargetId="figure@4.[762,1038,159,707]" captionTargetPageId="4" captionText="Fig. 3. Schistura titan, CMK 22344, paratype, 95.0 mm SL; digestive tract. Scale bar 5 mm." httpUri="https://zenodo.org/record/886890/files/figure.png" pageId="3" pageNumber="68">Fig. 3</figureCitation>
|
||
). Posterior chamber of air bladder present, large, extending to above pelvic-fin base.
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1187,364,558]" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[675,910,364,386]" pageId="3" pageNumber="68">Sexual dimorphism.</emphasis>
|
||
None of the characters usually associated with sexual dimorphism in
|
||
<taxonomicName box="[675,833,421,443]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="68" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
were observed (no suborbital flap, modified pectoral fins, pads of unculi or tubercles on cheek and fins). This is probably uninformative because both available specimens are possibly females (see below).
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1188,594,1278]" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[675,798,594,616]" pageId="3" pageNumber="68">Coloration.</emphasis>
|
||
After about two months in formalin. Head and body background colour pale olive brown, throat, belly, lower part of caudal peduncle pale yellowish; except otherwise stated, markings dark brown. Body with 17 or 18 bars (6 predorsal, 4 subdorsal, 7 or 8 postdorsal); bars of quite regular width and shape, slightly wider than interspaces, reaching downwards slightly beyond lateral line, continuous over back with contralaterals on caudal peduncle, irregular or broken down into small blotches on back in front of dorsal.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="DF68BADB8639FFABFF0FFD95FEB2FD03" httpUri="http://table.plazi.org/id/DF68BADB8639FFABFF0FFD95FEB2FD03" pageId="3" pageNumber="68" targetBox="[128,621,750,1666]" targetIsTable="true" targetPageId="3">
|
||
<paragraph blockId="3.[128,641,637,708]" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[128,205,637,656]" pageId="3" pageNumber="68">Table 1.</emphasis>
|
||
Morphometric data of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[428,556,637,656]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="68" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[428,556,637,656]" italics="true" pageId="3" pageNumber="68">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
(MHNG 2767.057, holotype; CMK 22344, paratype). H, holotype; P, paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="68">
|
||
<table box="[128,621,750,1666]" gridcols="3" gridrows="35" pageId="3" pageNumber="68">
|
||
<tr box="[128,621,750,769]" gridrow="0" pageId="3" pageNumber="68" rowspan-0="1">
|
||
<th box="[482,530,750,769]" gridcol="1" gridrow="0" pageId="3" pageNumber="68">H</th>
|
||
<th box="[573,621,750,769]" gridcol="2" gridrow="0" pageId="3" pageNumber="68">P</th>
|
||
</tr>
|
||
<tr box="[128,621,784,804]" gridrow="1" pageId="3" pageNumber="68">
|
||
<th box="[128,458,784,804]" gridcol="0" gridrow="1" pageId="3" pageNumber="68">Standard length (mm)</th>
|
||
<td box="[482,530,784,804]" gridcol="1" gridrow="1" pageId="3" pageNumber="68">74.8</td>
|
||
<td box="[573,621,784,804]" gridcol="2" gridrow="1" pageId="3" pageNumber="68">95.0</td>
|
||
</tr>
|
||
<tr box="[128,621,810,829]" gridrow="2" pageId="3" pageNumber="68">
|
||
<th box="[128,458,810,829]" gridcol="0" gridrow="2" pageId="3" pageNumber="68">Total length (mm)</th>
|
||
<td box="[482,530,810,829]" gridcol="1" gridrow="2" pageId="3" pageNumber="68">90.1</td>
|
||
<td box="[573,621,810,829]" gridcol="2" gridrow="2" pageId="3" pageNumber="68">111.3</td>
|
||
</tr>
|
||
<tr box="[128,621,843,862]" gridrow="3" pageId="3" pageNumber="68" rowspan-1="1" rowspan-2="1">
|
||
<th box="[128,458,843,862]" gridcol="0" gridrow="3" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[128,416,843,862]" pageId="3" pageNumber="68">In percent of standard length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,621,869,888]" gridrow="4" pageId="3" pageNumber="68">
|
||
<th box="[128,458,869,888]" gridcol="0" gridrow="4" pageId="3" pageNumber="68">Total length</th>
|
||
<td box="[482,530,869,888]" gridcol="1" gridrow="4" pageId="3" pageNumber="68">120.4</td>
|
||
<td box="[573,621,869,888]" gridcol="2" gridrow="4" pageId="3" pageNumber="68">117.1</td>
|
||
</tr>
|
||
<tr box="[128,621,894,913]" gridrow="5" pageId="3" pageNumber="68">
|
||
<th box="[128,458,894,913]" gridcol="0" gridrow="5" pageId="3" pageNumber="68">Head length (dorsal)</th>
|
||
<td box="[482,530,894,913]" gridcol="1" gridrow="5" pageId="3" pageNumber="68">21.2</td>
|
||
<td box="[573,621,894,913]" gridcol="2" gridrow="5" pageId="3" pageNumber="68">20.5</td>
|
||
</tr>
|
||
<tr box="[128,621,919,939]" gridrow="6" pageId="3" pageNumber="68">
|
||
<th box="[128,458,919,939]" gridcol="0" gridrow="6" pageId="3" pageNumber="68">Head length (lateral)</th>
|
||
<td box="[482,530,919,939]" gridcol="1" gridrow="6" pageId="3" pageNumber="68">24.7</td>
|
||
<td box="[573,621,919,939]" gridcol="2" gridrow="6" pageId="3" pageNumber="68">25.2</td>
|
||
</tr>
|
||
<tr box="[128,621,945,964]" gridrow="7" pageId="3" pageNumber="68">
|
||
<th box="[128,458,945,964]" gridcol="0" gridrow="7" pageId="3" pageNumber="68">Predorsal length</th>
|
||
<td box="[482,530,945,964]" gridcol="1" gridrow="7" pageId="3" pageNumber="68">55.6</td>
|
||
<td box="[573,621,945,964]" gridcol="2" gridrow="7" pageId="3" pageNumber="68">53.9</td>
|
||
</tr>
|
||
<tr box="[128,621,970,989]" gridrow="8" pageId="3" pageNumber="68">
|
||
<th box="[128,458,970,989]" gridcol="0" gridrow="8" pageId="3" pageNumber="68">Prepelvic length</th>
|
||
<td box="[482,530,970,989]" gridcol="1" gridrow="8" pageId="3" pageNumber="68">55.3</td>
|
||
<td box="[573,621,970,989]" gridcol="2" gridrow="8" pageId="3" pageNumber="68">53.8</td>
|
||
</tr>
|
||
<tr box="[128,621,996,1015]" gridrow="9" pageId="3" pageNumber="68">
|
||
<th box="[128,458,996,1015]" gridcol="0" gridrow="9" pageId="3" pageNumber="68">Pre-anus length</th>
|
||
<td box="[482,530,996,1015]" gridcol="1" gridrow="9" pageId="3" pageNumber="68">72.3</td>
|
||
<td box="[573,621,996,1015]" gridcol="2" gridrow="9" pageId="3" pageNumber="68">70.3</td>
|
||
</tr>
|
||
<tr box="[128,621,1021,1040]" gridrow="10" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1021,1040]" gridcol="0" gridrow="10" pageId="3" pageNumber="68">Pre-anal length</th>
|
||
<td box="[482,530,1021,1040]" gridcol="1" gridrow="10" pageId="3" pageNumber="68">77.9</td>
|
||
<td box="[573,621,1021,1040]" gridcol="2" gridrow="10" pageId="3" pageNumber="68">76.2</td>
|
||
</tr>
|
||
<tr box="[128,621,1046,1066]" gridrow="11" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1046,1066]" gridcol="0" gridrow="11" pageId="3" pageNumber="68">Head depth</th>
|
||
<td box="[482,530,1046,1066]" gridcol="1" gridrow="11" pageId="3" pageNumber="68">14.0</td>
|
||
<td box="[573,621,1046,1066]" gridcol="2" gridrow="11" pageId="3" pageNumber="68">14.7</td>
|
||
</tr>
|
||
<tr box="[128,621,1072,1091]" gridrow="12" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1072,1091]" gridcol="0" gridrow="12" pageId="3" pageNumber="68">Body depth at dorsal-fin origin</th>
|
||
<td box="[482,530,1072,1091]" gridcol="1" gridrow="12" pageId="3" pageNumber="68">16.7</td>
|
||
<td box="[573,621,1072,1091]" gridcol="2" gridrow="12" pageId="3" pageNumber="68">18.4</td>
|
||
</tr>
|
||
<tr box="[128,621,1097,1116]" gridrow="13" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1097,1116]" gridcol="0" gridrow="13" pageId="3" pageNumber="68">Depth of caudal peduncle</th>
|
||
<td box="[482,530,1097,1116]" gridcol="1" gridrow="13" pageId="3" pageNumber="68">12.0</td>
|
||
<td box="[573,621,1097,1116]" gridcol="2" gridrow="13" pageId="3" pageNumber="68">11.2</td>
|
||
</tr>
|
||
<tr box="[128,621,1123,1142]" gridrow="14" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1123,1142]" gridcol="0" gridrow="14" pageId="3" pageNumber="68">Length of caudal peduncle</th>
|
||
<td box="[482,530,1123,1142]" gridcol="1" gridrow="14" pageId="3" pageNumber="68">16.0</td>
|
||
<td box="[573,621,1123,1142]" gridcol="2" gridrow="14" pageId="3" pageNumber="68">16.2</td>
|
||
</tr>
|
||
<tr box="[128,621,1148,1167]" gridrow="15" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1148,1167]" gridcol="0" gridrow="15" pageId="3" pageNumber="68">Head width</th>
|
||
<td box="[482,530,1148,1167]" gridcol="1" gridrow="15" pageId="3" pageNumber="68">16.5</td>
|
||
<td box="[573,621,1148,1167]" gridcol="2" gridrow="15" pageId="3" pageNumber="68">18.0</td>
|
||
</tr>
|
||
<tr box="[128,621,1173,1193]" gridrow="16" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1173,1193]" gridcol="0" gridrow="16" pageId="3" pageNumber="68">Body width at dorsal-fin origin</th>
|
||
<td box="[482,530,1173,1193]" gridcol="1" gridrow="16" pageId="3" pageNumber="68">12.0</td>
|
||
<td box="[573,621,1173,1193]" gridcol="2" gridrow="16" pageId="3" pageNumber="68">13.6</td>
|
||
</tr>
|
||
<tr box="[128,621,1199,1218]" gridrow="17" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1199,1218]" gridcol="0" gridrow="17" pageId="3" pageNumber="68">Snout length</th>
|
||
<td box="[482,530,1199,1218]" gridcol="1" gridrow="17" pageId="3" pageNumber="68">10.9</td>
|
||
<td box="[573,621,1199,1218]" gridcol="2" gridrow="17" pageId="3" pageNumber="68">10.4</td>
|
||
</tr>
|
||
<tr box="[128,621,1224,1244]" gridrow="18" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1224,1244]" gridcol="0" gridrow="18" pageId="3" pageNumber="68">Eye diameter</th>
|
||
<td box="[482,530,1224,1244]" gridcol="1" gridrow="18" pageId="3" pageNumber="68">4.0</td>
|
||
<td box="[573,621,1224,1244]" gridcol="2" gridrow="18" pageId="3" pageNumber="68">3.8</td>
|
||
</tr>
|
||
<tr box="[128,621,1250,1269]" gridrow="19" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1250,1269]" gridcol="0" gridrow="19" pageId="3" pageNumber="68">Interorbital width</th>
|
||
<td box="[482,530,1250,1269]" gridcol="1" gridrow="19" pageId="3" pageNumber="68">8.3</td>
|
||
<td box="[573,621,1250,1269]" gridcol="2" gridrow="19" pageId="3" pageNumber="68">9.1</td>
|
||
</tr>
|
||
<tr box="[128,621,1275,1294]" gridrow="20" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1275,1294]" gridcol="0" gridrow="20" pageId="3" pageNumber="68">Length of dorsal fin</th>
|
||
<td box="[482,530,1275,1294]" gridcol="1" gridrow="20" pageId="3" pageNumber="68">16.3</td>
|
||
<td box="[573,621,1275,1294]" gridcol="2" gridrow="20" pageId="3" pageNumber="68">14.0</td>
|
||
</tr>
|
||
<tr box="[128,621,1300,1320]" gridrow="21" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1300,1320]" gridcol="0" gridrow="21" pageId="3" pageNumber="68">Length of upper caudal-fin lobe</th>
|
||
<td box="[482,530,1300,1320]" gridcol="1" gridrow="21" pageId="3" pageNumber="68">21.5</td>
|
||
<td box="[573,621,1300,1320]" gridcol="2" gridrow="21" pageId="3" pageNumber="68">18.7</td>
|
||
</tr>
|
||
<tr box="[128,621,1326,1345]" gridrow="22" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1326,1345]" gridcol="0" gridrow="22" pageId="3" pageNumber="68">Length of median caudal-fin rays</th>
|
||
<td box="[482,530,1326,1345]" gridcol="1" gridrow="22" pageId="3" pageNumber="68">17.8</td>
|
||
<td box="[573,621,1326,1345]" gridcol="2" gridrow="22" pageId="3" pageNumber="68">14.0</td>
|
||
</tr>
|
||
<tr box="[128,621,1351,1370]" gridrow="23" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1351,1370]" gridcol="0" gridrow="23" pageId="3" pageNumber="68">Length of lower caudal-fin lobe</th>
|
||
<td box="[482,530,1351,1370]" gridcol="1" gridrow="23" pageId="3" pageNumber="68">19.3</td>
|
||
<td box="[573,621,1351,1370]" gridcol="2" gridrow="23" pageId="3" pageNumber="68">17.3</td>
|
||
</tr>
|
||
<tr box="[128,621,1377,1396]" gridrow="24" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1377,1396]" gridcol="0" gridrow="24" pageId="3" pageNumber="68">Length of anal fin</th>
|
||
<td box="[482,530,1377,1396]" gridcol="1" gridrow="24" pageId="3" pageNumber="68">17.3</td>
|
||
<td box="[573,621,1377,1396]" gridcol="2" gridrow="24" pageId="3" pageNumber="68">16.3</td>
|
||
</tr>
|
||
<tr box="[128,621,1402,1421]" gridrow="25" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1402,1421]" gridcol="0" gridrow="25" pageId="3" pageNumber="68">Length of pelvic fin</th>
|
||
<td box="[482,530,1402,1421]" gridcol="1" gridrow="25" pageId="3" pageNumber="68">15.6</td>
|
||
<td box="[573,621,1402,1421]" gridcol="2" gridrow="25" pageId="3" pageNumber="68">13.0</td>
|
||
</tr>
|
||
<tr box="[128,621,1427,1447]" gridrow="26" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1427,1447]" gridcol="0" gridrow="26" pageId="3" pageNumber="68">Length of pectoral fin</th>
|
||
<td box="[482,530,1427,1447]" gridcol="1" gridrow="26" pageId="3" pageNumber="68">17.9</td>
|
||
<td box="[573,621,1427,1447]" gridcol="2" gridrow="26" pageId="3" pageNumber="68">16.5</td>
|
||
</tr>
|
||
<tr box="[128,621,1461,1480]" gridrow="27" pageId="3" pageNumber="68" rowspan-1="1" rowspan-2="1">
|
||
<th box="[128,458,1461,1480]" gridcol="0" gridrow="27" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[128,445,1461,1480]" pageId="3" pageNumber="68">In percent of dorsal head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,621,1486,1505]" gridrow="28" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1486,1505]" gridcol="0" gridrow="28" pageId="3" pageNumber="68">Snout length</th>
|
||
<td box="[482,530,1486,1505]" gridcol="1" gridrow="28" pageId="3" pageNumber="68">51</td>
|
||
<td box="[573,621,1486,1505]" gridcol="2" gridrow="28" pageId="3" pageNumber="68">50</td>
|
||
</tr>
|
||
<tr box="[128,621,1512,1531]" gridrow="29" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1512,1531]" gridcol="0" gridrow="29" pageId="3" pageNumber="68">Eye diameter</th>
|
||
<td box="[482,530,1512,1531]" gridcol="1" gridrow="29" pageId="3" pageNumber="68">19</td>
|
||
<td box="[573,621,1512,1531]" gridcol="2" gridrow="29" pageId="3" pageNumber="68">19</td>
|
||
</tr>
|
||
<tr box="[128,621,1537,1556]" gridrow="30" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1537,1556]" gridcol="0" gridrow="30" pageId="3" pageNumber="68">Interorbital width</th>
|
||
<td box="[482,530,1537,1556]" gridcol="1" gridrow="30" pageId="3" pageNumber="68">39</td>
|
||
<td box="[573,621,1537,1556]" gridcol="2" gridrow="30" pageId="3" pageNumber="68">44</td>
|
||
</tr>
|
||
<tr box="[128,621,1570,1590]" gridrow="31" pageId="3" pageNumber="68" rowspan-1="1" rowspan-2="1">
|
||
<th box="[128,458,1570,1590]" gridcol="0" gridrow="31" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[128,446,1570,1590]" pageId="3" pageNumber="68">In percent of lateral head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,621,1596,1615]" gridrow="32" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1596,1615]" gridcol="0" gridrow="32" pageId="3" pageNumber="68">Snout length</th>
|
||
<td box="[482,530,1596,1615]" gridcol="1" gridrow="32" pageId="3" pageNumber="68">44</td>
|
||
<td box="[573,621,1596,1615]" gridcol="2" gridrow="32" pageId="3" pageNumber="68">41</td>
|
||
</tr>
|
||
<tr box="[128,621,1621,1641]" gridrow="33" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1621,1641]" gridcol="0" gridrow="33" pageId="3" pageNumber="68">Eye diameter</th>
|
||
<td box="[482,530,1621,1641]" gridcol="1" gridrow="33" pageId="3" pageNumber="68">16</td>
|
||
<td box="[573,621,1621,1641]" gridcol="2" gridrow="33" pageId="3" pageNumber="68">15</td>
|
||
</tr>
|
||
<tr box="[128,621,1647,1666]" gridrow="34" pageId="3" pageNumber="68">
|
||
<th box="[128,458,1647,1666]" gridcol="0" gridrow="34" pageId="3" pageNumber="68">Interorbital width</th>
|
||
<td box="[482,530,1647,1666]" gridcol="1" gridrow="34" pageId="3" pageNumber="68">34</td>
|
||
<td box="[573,621,1647,1666]" gridcol="2" gridrow="34" pageId="3" pageNumber="68">36</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1188,594,1278]" pageId="3" pageNumber="68">
|
||
A faint black bar at caudal-fin base, on about lower
|
||
<emphasis bold="true" box="[746,750,968,980]" pageId="3" pageNumber="68">2</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[759,766,976,988]" pageId="3" pageNumber="68">3</emphasis>
|
||
of fin base, not extending on ventral crest. A faint blotch at base of upper simple and 3 posterior procurrent rays.
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1188,594,1278]" pageId="3" pageNumber="68">Dorsal fin hyaline; a small black spot at base of unbranched rays and branched rays 1-2, separated by a small hyaline area from a blackish brown elongated blotch along base of remaining rays. In large specimen, rays darker near first branching; a small elongated patch of black pigments on lowermost third of all membranes. Other fins hyaline.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="69" pageId="3" pageNumber="68" type="biology_ecology">
|
||
<paragraph blockId="3.[675,1187,1314,1682]" lastBlockId="4.[98,611,162,299]" lastPageId="4" lastPageNumber="69" pageId="3" pageNumber="68">
|
||
<emphasis bold="true" box="[675,883,1314,1336]" pageId="3" pageNumber="68">Notes on biology.</emphasis>
|
||
Both specimens were dissected. The holotype is a female with only a few yellow eggs about
|
||
<quantity box="[865,942,1372,1393]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="68" unit="mm" value="0.8">0.8 mm</quantity>
|
||
diameter. The paratype has a distended belly as usual for females but its sex could not be determined. They were collected on
|
||
<date box="[709,844,1458,1480]" pageId="3" pageNumber="68" value="2011-05-22">22 May 2011</date>
|
||
and this suggests that they had already spawned earlier in the dry season. The stomach of the paratype was large, filled by three unidentified insect larvae. Both specimens were infected by small worms free in the abdominal cavity as well as attached to the air bladder.
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[675,823,1631,1653]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="68" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[675,823,1631,1653]" italics="true" pageId="3" pageNumber="68">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
was obtained in a stream stretch under tree cover, with moderate flow over stone bottom but with important sand and mud deposit coming from activity further upstream (possibly gold mining judging from
|
||
<date box="[367,523,220,242]" pageId="4" pageNumber="69" value="2012-12">December 2012</date>
|
||
pictures on Google Earth, which would explain the very low number of fish present).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="69" type="distribution">
|
||
<paragraph blockId="4.[98,611,335,846]" pageId="4" pageNumber="69">
|
||
<emphasis bold="true" box="[98,241,335,357]" pageId="4" pageNumber="69">Distribution.</emphasis>
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[254,401,335,357]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[254,401,335,357]" italics="true" pageId="4" pageNumber="69">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
was observed only at the
|
||
<typeStatus box="[166,213,364,385]" pageId="4" pageNumber="69">type</typeStatus>
|
||
locality on Dakchung Plateau at Ban Dak (
|
||
<figureCitation box="[158,218,392,415]" captionStart="Fig. 4" captionStartId="5.[129,161,1009,1028]" captionTargetBox="[130,1181,165,994]" captionTargetId="figure@5.[118,1187,159,987]" captionText="Fig. 4. Dakchung Plateau (North East; A) and Bolaven Plateau (West; B) in southern Laos, and Xe Kong River flowing from North to South between them.: type locality of Schistura titan." httpUri="https://zenodo.org/record/886892/files/figure.png" pageId="4" pageNumber="69">Fig. 4</figureCitation>
|
||
). The toponymy of the area is quite confused. The villagers identified their village as Ban Dak and the streams as Houay Champong and Houay Xe Kaman. Ban Dak appears to be Ban Lak Tai on the 1: 100 0 0 0 topographic map (Service Géographique d’ État, 1987, sheet D-48- 23) and Houay Xe Kaman as Houay Poyo. The map shows Houay Poyo as a tributary of Nam Poay-O, in turn a tributary of Xe Kaman. Houay Xe Kaman is not the same as Xe Kaman, the main river on Dakchung Plateau. To further confuse the issue, Nam Poay-O was called Xe Kaman by the Ban Dak inhabitants that were interviewed. Ban Dak appears as Dak Long on Google Earth, but on the map this is the name of a village about
|
||
<quantity box="[98,151,824,846]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="69" unit="km" value="2.0">2 km</quantity>
|
||
west of Ban Dak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption httpUri="https://zenodo.org/record/886890/files/figure.png" pageId="4" pageNumber="69" targetBox="[762,1038,159,707]" targetPageId="4">
|
||
<paragraph blockId="4.[644,1157,725,770]" pageId="4" pageNumber="69">
|
||
<emphasis bold="true" box="[644,702,725,744]" pageId="4" pageNumber="69">Fig. 3.</emphasis>
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[713,843,725,744]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[713,843,725,744]" italics="true" pageId="4" pageNumber="69">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
, CMK 22344, paratype, 95.0 mm SL; digestive tract. Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="4" pageNumber="69" type="etymology">
|
||
<paragraph blockId="4.[98,611,881,962]" pageId="4" pageNumber="69">
|
||
<emphasis bold="true" box="[98,223,882,903]" pageId="4" pageNumber="69">Etymology.</emphasis>
|
||
From the Greek τιτάν (one of the Titans, giant deities), a reference to the large size of the specimens. A noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="71" pageId="4" pageNumber="69" type="discussion">
|
||
<paragraph blockId="4.[98,611,997,1682]" lastBlockId="4.[644,1157,824,1681]" pageId="4" pageNumber="69">
|
||
<emphasis bold="true" box="[98,201,998,1019]" pageId="4" pageNumber="69">Remarks.</emphasis>
|
||
There is no clue to the affinities of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[98,257,1026,1048]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[98,257,1026,1048]" italics="true" pageId="4" pageNumber="69">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
. It is peculiar in having 8 + 7 branched caudal-fin rays. The vast majority of species of
|
||
<taxonomicName authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[209,301,1084,1105]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="genus" sensu="lato" status="s. l.">
|
||
<emphasis box="[209,301,1084,1105]" italics="true" pageId="4" pageNumber="69">Schistura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[309,341,1084,1106]" pageId="4" pageNumber="69" sensu="lato">s. l.</taxonomicNameLabel>
|
||
have 9 +8 branched rays (as do the majority of
|
||
<taxonomicName box="[331,490,1112,1134]" class="Actinopterygii" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="order">Cypriniformes</taxonomicName>
|
||
). A smaller number of caudal-fin rays is usually found in small-size species, like
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[372,463,1170,1191]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="crabro">
|
||
<emphasis box="[372,463,1170,1191]" italics="true" pageId="4" pageNumber="69">S. crabro</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1998" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="punctifasciata">
|
||
<emphasis italics="true" pageId="4" pageNumber="69">S. punctifasciata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[158,280,1199,1220]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="fusinotata">
|
||
<emphasis box="[158,280,1199,1220]" italics="true" pageId="4" pageNumber="69">S. fusinotata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Ou, Montaña, Winemiller & Conway" authorityYear="2011" box="[294,408,1199,1220]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="diminuta">
|
||
<emphasis box="[294,408,1199,1220]" italics="true" pageId="4" pageNumber="69">S. diminuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[422,556,1199,1220]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="daubentoni">
|
||
<emphasis box="[422,556,1199,1220]" italics="true" pageId="4" pageNumber="69">S. daubentoni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[98,175,1228,1249]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="rikiki">
|
||
<emphasis box="[98,175,1228,1249]" italics="true" pageId="4" pageNumber="69">S. rikiki</emphasis>
|
||
</taxonomicName>
|
||
. Very few species of the size of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[538,611,1228,1249]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[538,611,1228,1249]" italics="true" pageId="4" pageNumber="69">S. titan</emphasis>
|
||
</taxonomicName>
|
||
have a reduced number of branched caudal-fin rays;
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[158,280,1286,1307]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="antennata">
|
||
<emphasis box="[158,280,1286,1307]" italics="true" pageId="4" pageNumber="69">S. antennata</emphasis>
|
||
</taxonomicName>
|
||
(from the Song Lam drainage in Central
|
||
<collectingCountry box="[210,301,1314,1336]" name="Vietnam" pageId="4" pageNumber="69">Vietnam</collectingCountry>
|
||
) and
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[360,497,1314,1335]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="namboensis">
|
||
<emphasis box="[360,497,1314,1335]" italics="true" pageId="4" pageNumber="69">S. namboensis</emphasis>
|
||
</taxonomicName>
|
||
(from Srepok and Sesan [Mekong drainage] and coastal drainages in southern Central
|
||
<collectingCountry box="[418,507,1372,1394]" name="Vietnam" pageId="4" pageNumber="69">Vietnam</collectingCountry>
|
||
[between Quy Nohon and Phan Rang]) are known to reach
|
||
<quantity box="[163,235,1429,1451]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="69" unit="mm" value="74.0">74 mm</quantity>
|
||
SL and to have 8 + 8 branched rays (
|
||
<bibRefCitation author="Freyhof" box="[105,342,1458,1480]" journalOrPublisher="Ichthyological Exploration of Freshwatersi" pageId="4" pageNumber="69" pagination="133 - 191" part="12" refString="Freyhofi J. & D. V. Serov. 2001. Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwatersi 12: 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof & Serov, 2001</bibRefCitation>
|
||
). In these two species, the air bladder does not have a posterior chamber in the abdominal cavity. Besides
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[462,584,1516,1537]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="antennata">
|
||
<emphasis box="[462,584,1516,1537]" italics="true" pageId="4" pageNumber="69">S. antennata</emphasis>
|
||
</taxonomicName>
|
||
is immediately distinguished by its more slender appearance, its unique colour pattern in which the black bars are broader with age and are joined at their upper and lower extremities and only leave narrow yellowish interspaces.
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[434,584,1660,1681]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[434,584,1660,1681]" italics="true" pageId="4" pageNumber="69">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[933,1071,824,846]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="namboensis">
|
||
<emphasis box="[933,1071,824,846]" italics="true" pageId="4" pageNumber="69">S. namboensis</emphasis>
|
||
</taxonomicName>
|
||
by having a complete lateral line (vs. reaching at most to anal-fin origin), and a very different appearance with 17 or 18 narrow and regular bars on the body (vs. 7-10, irregular, often dissociated into a saddle and a blotch), and no axial band of deep pigmentation (vs. presence of such a band) (data on
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[676,798,1026,1048]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="antennata">
|
||
<emphasis box="[676,798,1026,1048]" italics="true" pageId="4" pageNumber="69">S. antennata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Freyhof & Serov" authorityYear="2001" box="[848,985,1026,1047]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="namboensis">
|
||
<emphasis box="[848,985,1026,1047]" italics="true" pageId="4" pageNumber="69">S. namboensis</emphasis>
|
||
</taxonomicName>
|
||
from specimens and from
|
||
<bibRefCitation author="Freyhof" box="[750,989,1055,1077]" journalOrPublisher="Ichthyological Exploration of Freshwatersi" pageId="4" pageNumber="69" pagination="133 - 191" part="12" refString="Freyhofi J. & D. V. Serov. 2001. Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwatersi 12: 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof & Serov, 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[644,1157,824,1681]" pageId="4" pageNumber="69">
|
||
The presence of a posterior chamber of the air bladder in the abdominal cavity had been used to diagnose some genera in
|
||
<taxonomicName box="[937,1097,1141,1163]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
(e.g.,
|
||
<taxonomicName box="[644,791,1170,1191]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="genus">
|
||
<emphasis box="[644,791,1170,1191]" italics="true" pageId="4" pageNumber="69">Physoschistura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Rendahl" authorityYear="1933" box="[803,919,1170,1191]" class="Actinopterygii" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="genus">
|
||
<emphasis box="[803,919,1170,1191]" italics="true" pageId="4" pageNumber="69">Triplophysa</emphasis>
|
||
</taxonomicName>
|
||
) but even in these it is variously developed or present. Within
|
||
<taxonomicName authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[1064,1156,1199,1220]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="genus" sensu="lato" status="s. l.">
|
||
<emphasis box="[1064,1156,1199,1220]" italics="true" pageId="4" pageNumber="69">Schistura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[644,676,1227,1249]" pageId="4" pageNumber="69" sensu="lato">s. l.</taxonomicNameLabel>
|
||
a free posterior chamber has been reported only in
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[722,811,1256,1278]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis box="[722,811,1256,1278]" italics="true" pageId="4" pageNumber="69">S. similis</emphasis>
|
||
</taxonomicName>
|
||
from the Salween drainage (
|
||
<bibRefCitation author="Kottelat" bookContentInfo="262 pp." journalOrPublisher="Pfeili Muncheni" pageId="4" pageNumber="69" refString="Kottelati M. 1990. Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailandi Burmai Laosi Cambodia and southern Viet Nam. Pfeili Muncheni 262 pp." title="Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailandi Burmai Laosi Cambodia and southern Viet Nam" type="book" year="1990">Kottelat, 1990: 210</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[828,976,1285,1307]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[828,976,1285,1307]" italics="true" pageId="4" pageNumber="69">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[702,791,1314,1335]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="similis">
|
||
<emphasis box="[702,791,1314,1335]" italics="true" pageId="4" pageNumber="69">S. similis</emphasis>
|
||
</taxonomicName>
|
||
by having a very different appearance, 17 or 18 narrow and regular bars on the body (vs. 7 or 8 broad blackish brown regular and very contrasted bars), 8 +7 branched caudal rays (vs. 9 + 8), the intestine with a bend immediately behind the stomach (vs. at some distance behind the stomach), and a more slender caudal peduncle (depth 1.3-1.5 times in its length, vs. 0.8-1.3).
|
||
</paragraph>
|
||
<paragraph blockId="4.[644,1157,824,1681]" pageId="4" pageNumber="69">
|
||
Peculiar patterns of black markings at the base of the caudal fin are known to be diagnostic for some species or some lineages within
|
||
<taxonomicName authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[1030,1121,1602,1623]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="genus" sensu="lato" status="s. l.">
|
||
<emphasis box="[1030,1121,1602,1623]" italics="true" pageId="4" pageNumber="69">Schistura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1125,1157,1602,1624]" pageId="4" pageNumber="69" sensu="lato">s. l.</taxonomicNameLabel>
|
||
That of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[728,800,1631,1652]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="69" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[728,800,1631,1652]" italics="true" pageId="4" pageNumber="69">S. titan</emphasis>
|
||
</taxonomicName>
|
||
is faint and of a very generalised
|
||
<typeStatus box="[644,691,1660,1681]" pageId="4" pageNumber="69">type</typeStatus>
|
||
and, therefore, uninformative.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/886892/files/figure.png" pageId="5" pageNumber="70" targetBox="[130,1181,165,994]" targetPageId="5">
|
||
<paragraph blockId="5.[129,1187,1009,1054]" pageId="5" pageNumber="70">
|
||
<emphasis bold="true" box="[129,186,1009,1028]" pageId="5" pageNumber="70">Fig. 4.</emphasis>
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC88863FFFADFF4AFC19FEE8FBC3" box="[197,372,1009,1028]" country="Laos" county="South" municipality="North" name="Dakchung Plateau" pageId="5" pageNumber="70">Dakchung Plateau</location>
|
||
(
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC88863FFFADFE0EFC19FE70FBC2" box="[385,492,1009,1029]" country="Laos" county="South" municipality="North" name="North East" pageId="5" pageNumber="70">North East</location>
|
||
;
|
||
<emphasis bold="true" box="[502,519,1009,1028]" pageId="5" pageNumber="70">A</emphasis>
|
||
) and
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC88863FFFADFDB0FC19FD44FBC3" box="[575,728,1009,1028]" country="Laos" county="South" municipality="North" name="Bolaven Plateau" pageId="5" pageNumber="70">Bolaven Plateau</location>
|
||
(
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC88863FFFADFD69FC19FC84FBC3" box="[742,792,1009,1028]" country="Laos" county="South" municipality="North" name="West" pageId="5" pageNumber="70">West</location>
|
||
;
|
||
<emphasis bold="true" box="[802,816,1010,1029]" pageId="5" pageNumber="70">B</emphasis>
|
||
) in southern
|
||
<collectingCountry box="[947,994,1010,1029]" name="Laos" pageId="5" pageNumber="70">Laos</collectingCountry>
|
||
, and
|
||
<location LSID="urn:lsid:plazi:treatment:03BE5B458638FFAFFF46FD87FE98FE8F:8EC8BC88863FFFADFB99FC1AFB3FFBC3" box="[1046,1187,1009,1029]" country="Laos" county="South" municipality="North" name="Xe Kong River" pageId="5" pageNumber="70">Xe Kong River</location>
|
||
flowing from
|
||
<collectingMunicipality box="[259,316,1035,1054]" pageId="5" pageNumber="70">North</collectingMunicipality>
|
||
to
|
||
<collectingCounty box="[347,402,1035,1054]" pageId="5" pageNumber="70">South</collectingCounty>
|
||
between them.:
|
||
<typeStatus box="[589,631,1035,1054]" pageId="5" pageNumber="70">type</typeStatus>
|
||
locality of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[736,865,1035,1054]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[736,865,1035,1054]" italics="true" pageId="5" pageNumber="70">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[129,642,1108,1677]" pageId="5" pageNumber="70">
|
||
With a maximum known size of
|
||
<quantity box="[527,599,1108,1130]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="70" unit="mm" value="95.0">95 mm</quantity>
|
||
SL,
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[129,201,1137,1159]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[129,201,1137,1159]" italics="true" pageId="5" pageNumber="70">S. titan</emphasis>
|
||
</taxonomicName>
|
||
is a giant within
|
||
<taxonomicName authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[387,479,1137,1158]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="genus" sensu="lato" status="s. l.">
|
||
<emphasis box="[387,479,1137,1158]" italics="true" pageId="5" pageNumber="70">Schistura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[486,518,1137,1159]" pageId="5" pageNumber="70" sensu="lato">s. l.</taxonomicNameLabel>
|
||
Most other species have a maximum length below
|
||
<quantity box="[531,603,1166,1187]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="70" unit="mm" value="70.0">70 mm</quantity>
|
||
SL. Other large species in the Mekong drainage are:
|
||
<taxonomicName box="[129,231,1224,1245]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="sokolooi">
|
||
<emphasis box="[129,231,1224,1245]" italics="true" pageId="5" pageNumber="70">S. sokolooi</emphasis>
|
||
</taxonomicName>
|
||
which also reaches
|
||
<quantity box="[458,530,1223,1245]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="70" unit="mm" value="95.0">95 mm</quantity>
|
||
SL (
|
||
<bibRefCitation author="Freyhof" journalOrPublisher="Ichthyological Exploration of Freshwatersi" pageId="5" pageNumber="70" pagination="133 - 191" part="12" refString="Freyhofi J. & D. V. Serov. 2001. Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwatersi 12: 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof & Serov, 2001: 172</bibRefCitation>
|
||
) from headwaters of the Sesan and Srepok rivers in
|
||
<collectingCountry box="[409,498,1281,1303]" name="Vietnam" pageId="5" pageNumber="70">Vietnam</collectingCountry>
|
||
and
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[548,641,1281,1302]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="colossa">
|
||
<emphasis box="[548,641,1281,1302]" italics="true" pageId="5" pageNumber="70">S. colossa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity box="[136,209,1310,1331]" metricMagnitude="-2" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="70" unit="mm" value="98.0">98 mm</quantity>
|
||
SL) on Bolaven Plateau.
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[472,618,1310,1332]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[472,618,1310,1332]" italics="true" pageId="5" pageNumber="70">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from them by the presence of the posterior chamber of the air bladder (vs. absence), by having 8 +7 branched caudal rays (vs. 9 + 8), and by having the pelvic-fin origin behind the dorsal-fin origin (vs. below or slightly in front of it).
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[166,315,1483,1504]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[166,315,1483,1504]" italics="true" pageId="5" pageNumber="70">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from
|
||
<taxonomicName box="[129,231,1512,1533]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="sokolooi">
|
||
<emphasis box="[129,231,1512,1533]" italics="true" pageId="5" pageNumber="70">S. sokolooi</emphasis>
|
||
</taxonomicName>
|
||
by having more bars on the body (17 or 18, vs. 11-15, in both cases quite regularly set, continuous with contralaterals at least behind the dorsal fin), the more backward pelvic-fin origin (prepelvic length 53.8-55.3 % SL, vs. 50.4-53.2), a wider head (maximum width 16.5-18.0 % SL, vs.
|
||
</paragraph>
|
||
<paragraph blockId="5.[675,1187,1107,1678]" pageId="5" pageNumber="70">
|
||
13.4-16.9), and a shorter snout (41-44 % of lateral head length, vs. 44-54) (morphometric data from
|
||
<bibRefCitation author="Freyhof" box="[675,916,1166,1188]" journalOrPublisher="Ichthyological Exploration of Freshwatersi" pageId="5" pageNumber="70" pagination="133 - 191" part="12" refString="Freyhofi J. & D. V. Serov. 2001. Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwatersi 12: 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof & Serov, 2001</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[934,1081,1166,1188]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[934,1081,1166,1188]" italics="true" pageId="5" pageNumber="70">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[881,973,1195,1216]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="colossa">
|
||
<emphasis box="[881,973,1195,1216]" italics="true" pageId="5" pageNumber="70">S. colossa</emphasis>
|
||
</taxonomicName>
|
||
by its colour pattern (17 or 18 regular bars on the flank, vs. 16-21 bars quite regularly shaped in juveniles and with increasing size becoming more irregular and, on the caudal peduncle, broken up in irregular blotches), a rounded snout (vs. pointed), wider head (16.5- 18.0 % SL, vs. 11.8-14.7) and body (12.0-13.6 % SL, vs. 9.9-13.5), a greater interorbital distance (34-36 % of lateral head length, vs. 25-31), a longer head (lateral head length 24.7-25.2 % SL, vs. 21.1-23.8), a greater prepelvic length (53.8- 55.3 % SL, vs. 48.8-52.2), and a greater preanal length (76.2-77.9 % SL, vs. 73.9-76.4).
|
||
</paragraph>
|
||
<paragraph blockId="5.[675,1187,1107,1678]" lastBlockId="6.[98,611,846,1675]" lastPageId="6" lastPageNumber="71" pageId="5" pageNumber="70">
|
||
By its general appearance and large size,
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[675,747,1598,1620]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[675,747,1598,1620]" italics="true" pageId="5" pageNumber="70">S. titan</emphasis>
|
||
</taxonomicName>
|
||
is somewhat reminiscent of
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[1059,1181,1598,1619]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="klydonion">
|
||
<emphasis box="[1059,1181,1598,1619]" italics="true" pageId="5" pageNumber="70">S. klydonion</emphasis>
|
||
</taxonomicName>
|
||
, also from Bolaven Plateau, of which the largest known individual is
|
||
<quantity box="[891,963,1656,1678]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="70" unit="mm" value="76.0">76 mm</quantity>
|
||
SL. However,
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[1115,1187,1656,1677]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[1115,1187,1656,1677]" italics="true" pageId="5" pageNumber="70">S. titan</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[324,446,846,868]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="klydonion">
|
||
<emphasis box="[324,446,846,868]" italics="true" pageId="6" pageNumber="71">S. klydonion</emphasis>
|
||
</taxonomicName>
|
||
by the presence of the posterior chamber of the air bladder (vs. absence), 8+ 7 branched caudal-fin rays (vs. 9 + 8), the colour pattern (17 or 18 regular bars on the flank, continuous with contralaterals at least behind the dorsal fin, vs. a midlateral row of 12-21 bars that do not reach the dorsal midline and that alternate with a middorsal row of saddles or small blotches, leaving a pale zigzag line between the two rows), the lips without sparsely-set pointed papillae (vs. presence), a more backward pelvic-fin origin (prepelvic length 53.8-55.3 % SL, vs. 50.3-52.2), a wider head (maximum width 16.5-18.0 % SL, vs. 13.3-16.1), and a greater interorbital distance (34-36 % of lateral head length, vs. 26-33). Besides, the black pattern at the base of the caudal fin in
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[127,249,1307,1328]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="klydonion">
|
||
<emphasis box="[127,249,1307,1328]" italics="true" pageId="6" pageNumber="71">S. klydonion</emphasis>
|
||
</taxonomicName>
|
||
is made of a vertically elongated blotch occupying the middle half of the base of the fin, sometimes with constriction at the level of the lateral line; above and below, the blotch is continued by an arched band, along the base of the principal rays, not reaching dorsal and ventral midlines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption box="[98,983,773,792]" httpUri="https://zenodo.org/record/886894/files/figure.png" pageId="6" pageNumber="71" targetBox="[131,1132,177,737]" targetPageId="6">
|
||
<paragraph blockId="6.[98,983,773,792]" box="[98,983,773,792]" pageId="6" pageNumber="71">
|
||
<emphasis bold="true" box="[98,155,773,792]" pageId="6" pageNumber="71">Fig. 5.</emphasis>
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[166,320,773,792]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[166,320,773,792]" italics="true" pageId="6" pageNumber="71">Schistura clatrata</emphasis>
|
||
</taxonomicName>
|
||
, CMK 22329, 70.9 mm SL; Laos: Xekong Province: Dakchung Plateau.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71" type="distribution">
|
||
<paragraph blockId="6.[98,611,846,1675]" lastBlockId="6.[644,1158,846,1674]" pageId="6" pageNumber="71">
|
||
A single other species of
|
||
<taxonomicName authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[395,487,1509,1530]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus" sensu="lato" status="s. l.">
|
||
<emphasis box="[395,487,1509,1530]" italics="true" pageId="6" pageNumber="71">Schistura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[492,524,1509,1531]" pageId="6" pageNumber="71" sensu="lato">s. l.</taxonomicNameLabel>
|
||
was observed on Dakchung Plateau,
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[409,508,1538,1559]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[409,508,1538,1559]" italics="true" pageId="6" pageNumber="71">S. clatrata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[381,481,1566,1588]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[381,481,1566,1588]" italics="true" pageId="6" pageNumber="71">S. clatrata</emphasis>
|
||
</taxonomicName>
|
||
by the presence of the posterior chamber of the air bladder (vs. absence), 8 + 7 branched caudal rays (vs. 9+ 8), and the colour pattern (17 or 18 regular bars on the flank, continuous with contralaterals at least behind the dorsal fin, vs. 9-13, usually continuous with contralaterals also in front of the dorsal fin, with exceptions).
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[877,1051,933,954]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[877,1051,933,954]" italics="true" pageId="6" pageNumber="71">Schistura clatrata</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in the Xe Kong drainage; there is considerable variation in the colour pattern between different samples, and it cannot be excluded that more than one species are involved. Awaiting further analysis, they are herein recognised as a single species. In the samples from the upper Xe Kamam watershed on Dackchung Plateau, the bars are usually not very contrasted, short, not reaching much downwards beyond the lateral line (
|
||
<figureCitation box="[698,758,1220,1242]" captionStart="Fig. 5" captionStartId="6.[98,130,773,792]" captionTargetBox="[131,1132,177,737]" captionTargetId="figure@6.[98,1156,156,481]" captionTargetPageId="6" captionText="Fig. 5. Schistura clatrata, CMK 22329, 70.9 mm SL; Laos: Xekong Province: Dakchung Plateau." httpUri="https://zenodo.org/record/886894/files/figure.png" pageId="6" pageNumber="71">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="6.[644,1158,846,1674]" lastBlockId="7.[129,641,162,328]" lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71">
|
||
Only 11 fish species have been observed from a total of seven sites on Dakchung Plateau:
|
||
<emphasis box="[644,790,1307,1329]" italics="true" pageId="6" pageNumber="71">Deoario gibber</emphasis>
|
||
,
|
||
<taxonomicName baseAuthorityName="Pellegrin & Fang" baseAuthorityYear="1940" box="[803,1009,1307,1328]" class="Actinopterygii" family="Cyprinidae" genus="Neolissochilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="blanci">
|
||
<emphasis box="[803,1009,1307,1328]" italics="true" pageId="6" pageNumber="71">Neolissochilus blanci</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1998" class="Actinopterygii" family="Cyprinidae" genus="Onychostoma" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="meridionale">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Onychostoma meridionale</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Wu & Lin" baseAuthorityYear="1977" box="[771,1024,1336,1358]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="carinatus">
|
||
<emphasis box="[771,892,1336,1357]" italics="true" pageId="6" pageNumber="71">Poropuntius</emphasis>
|
||
cf.
|
||
<emphasis box="[933,1024,1336,1357]" italics="true" pageId="6" pageNumber="71">carinatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Vinciguerra" authorityYear="1890" class="Actinopterygii" family="Cyprinidae" genus="Scaphiodonichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Scaphiodonichthys</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName box="[758,877,1364,1386]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
|
||
),
|
||
<taxonomicName baseAuthorityName="Hora" baseAuthorityYear="1931" box="[894,1068,1365,1386]" class="Actinopterygii" family="Balitoridae" genus="Annamia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="normani">
|
||
<emphasis box="[894,1068,1365,1386]" italics="true" pageId="6" pageNumber="71">Annamia normani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Roberts" authorityYear="1998" class="Maxillopoda" family="Thalestridae" genus="Sewellia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="diardi">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Sewellia diardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[714,794,1393,1414]" class="Maxillopoda" family="Thalestridae" genus="Sewellia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[714,794,1393,1414]" italics="true" pageId="6" pageNumber="71">Sewellia</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName authorityName="Fowler" authorityYear="1905" box="[845,1044,1393,1415]" class="Actinopterygii" family="Gastromyzontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="family">Gastromyzontidae</taxonomicName>
|
||
),
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Schistura clatrata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[739,814,1422,1444]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[739,814,1422,1444]" italics="true" pageId="6" pageNumber="71">S. titan</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[833,999,1422,1444]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
), and
|
||
<taxonomicName baseAuthorityName="Hamilton" baseAuthorityYear="1822" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="gachua">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Channa gachua</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[729,849,1451,1473]" class="Actinopterygii" family="Channidae" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="71" phylum="Chordata" rank="family">Channidae</taxonomicName>
|
||
). Of them, three are new to science (
|
||
<taxonomicName authorityName="Maurice Kottelat" authorityYear="2017" box="[735,883,1480,1501]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="titan">
|
||
<emphasis box="[735,883,1480,1501]" italics="true" pageId="6" pageNumber="71">Schistura titan</emphasis>
|
||
</taxonomicName>
|
||
, described here, and
|
||
<taxonomicName authorityName="Vinciguerra" authorityYear="1890" class="Actinopterygii" family="Cyprinidae" genus="Scaphiodonichthys" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="6" pageNumber="71">Scaphiodonichthys</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName box="[875,955,1509,1530]" class="Maxillopoda" family="Thalestridae" genus="Sewellia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[875,955,1509,1530]" italics="true" pageId="6" pageNumber="71">Sewellia</emphasis>
|
||
</taxonomicName>
|
||
sp., to be described soon). The variability in
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[894,1062,1537,1558]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[894,1062,1537,1558]" italics="true" pageId="6" pageNumber="71">Schistura clatrata</emphasis>
|
||
</taxonomicName>
|
||
has been mentioned above. The status of
|
||
<taxonomicName baseAuthorityName="Wu & Lin" baseAuthorityYear="1977" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="carinatus">
|
||
<emphasis box="[1001,1122,1567,1588]" italics="true" pageId="6" pageNumber="71">Poropuntius</emphasis>
|
||
cf.
|
||
<emphasis box="[644,735,1596,1617]" italics="true" pageId="6" pageNumber="71">carinatus</emphasis>
|
||
</taxonomicName>
|
||
is not clear yet. It seems to be unnamed too but a number of nominal species of
|
||
<taxonomicName authorityName="Smith" authorityYear="1931" box="[1038,1156,1624,1645]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1038,1156,1624,1645]" italics="true" pageId="6" pageNumber="71">Poropuntius</emphasis>
|
||
</taxonomicName>
|
||
have been reported or described in recent years from adjacent areas of
|
||
<collectingCountry box="[370,461,162,184]" name="Vietnam" pageId="7" pageNumber="72">Vietnam</collectingCountry>
|
||
, including in the Xe Kong drainage (
|
||
<taxonomicName baseAuthorityName="Nguyen" baseAuthorityYear="1997" box="[346,469,191,213]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="aluoiensis">
|
||
<emphasis box="[346,469,191,213]" italics="true" pageId="7" pageNumber="72">P. aluoiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Chevey" baseAuthorityYear="1934" box="[485,635,191,213]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="kontumensis">
|
||
<emphasis box="[485,635,191,213]" italics="true" pageId="7" pageNumber="72">P. kontumensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[129,241,220,241]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="yalyensis">
|
||
<emphasis box="[129,241,220,241]" italics="true" pageId="7" pageNumber="72">P. yalyensis</emphasis>
|
||
</taxonomicName>
|
||
). Unfortunately these descriptions and the accompanying illustrations are of limited use and it is presently not possible to reach reasonable conclusions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |