223 lines
28 KiB
XML
223 lines
28 KiB
XML
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<mods:title id="E2BE7804BAFACB19B0F1F7385D22AE14">Phylogenetic and morphological resolution of the Helobdella stagnalis species-complex (Annelida: Clitellata: Hirudinea)</mods:title>
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<mods:namePart id="2C449BCB2BBAA700567402434F433422">Saglam, Naim</mods:namePart>
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<mods:namePart id="4D9630053547CFD164B8D3941A42A848">Kutschera, Ulrich</mods:namePart>
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<mods:namePart id="564F47472F947C0F76C1E55D62BC0479">Saunders, Ralph</mods:namePart>
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<mods:namePart id="17ACDDC8C8F68B28E2C990EE17F08AD5">Saidel, William M.</mods:namePart>
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<mods:namePart id="EEB9A945955C9D21BD05BABC7C9B6681">Balombini, Katherine L. W.</mods:namePart>
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<mods:namePart id="EB7C81DEC28CEB4F0772A75EA4CF1D77">Shain, Daniel H.</mods:namePart>
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<mods:title id="D51AE3FE715B62058E9F7992E8D05358">Zootaxa</mods:title>
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<mods:date id="2F7CBE5899557A2A26FAE3D02CD0AD66">2018</mods:date>
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<treatment id="277487D6FF8B4441E5C859E5FAFA3297" ID-DOI="http://doi.org/10.5281/zenodo.5673213" ID-GBIF-Taxon="143467318" ID-Zenodo-Dep="5673213" LSID="urn:lsid:zoobank.org:act:F55674B6-1767-4F64-ABEF-0347122944E0" httpUri="http://treatment.plazi.org/id/277487D6FF8B4441E5C859E5FAFA3297" lastPageNumber="71" pageId="10" pageNumber="71">
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<subSubSection id="E7C7654BFF8B4441E5C859E5FDA33000" box="[199,564,1269,1294]" pageId="10" pageNumber="71" type="nomenclature">
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<paragraph id="AF6236C0FF8B4441E5C859E5FDA33000" blockId="10.[151,1437,1269,2017]" box="[199,564,1269,1294]" pageId="10" pageNumber="71">
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<taxonomicName id="68DD4D43FF8B4441E5C859E5FE5D3000" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[199,458,1269,1294]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="echoensis" status="sp. nov.">
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<emphasis id="9DA9EAD2FF8B4441E5C859E5FE5D3000" box="[199,458,1269,1294]" italics="true" pageId="10" pageNumber="71">Helobdella echoensis</emphasis>
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</taxonomicName>
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<emphasis id="9DA9EAD2FF8B4441E4EC59E5FDA33000" bold="true" box="[483,564,1269,1294]" pageId="10" pageNumber="71">
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<taxonomicNameLabel id="869A57A9FF8B4441E4EC59E5FDA33000" box="[483,564,1269,1294]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="E7C7654BFF8B4441E74259E5FD4F3000" box="[589,728,1269,1294]" pageId="10" pageNumber="71" type="materials_examined">
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<paragraph id="AF6236C0FF8B4441E74259E5FD4F3000" blockId="10.[151,1437,1269,2017]" box="[589,728,1269,1294]" pageId="10" pageNumber="71">
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(
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<collectingRegion id="6D19F822FF8B4441E75A59E5FD073000" box="[597,656,1269,1294]" country="United Kingdom" name="York" pageId="10" pageNumber="71">York</collectingRegion>
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, PA)
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</paragraph>
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<subSubSection id="E7C7654BFF8B4441E7FE59E5FC363000" box="[753,929,1269,1294]" pageId="10" pageNumber="71" type="description">
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<paragraph id="AF6236C0FF8B4441E7FE59E5FC363000" blockId="10.[151,1437,1269,2017]" box="[753,929,1269,1294]" pageId="10" pageNumber="71">
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(
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<figureCitation id="37E62A45FF8B4441E7F559E5FCFC3000" box="[762,875,1269,1294]" captionStart="FIGURE 2" captionStartId="5.[151,250,1920,1942]" captionTargetBox="[165,1433,419,1907]" captionTargetId="figure@5.[156,1433,404,1907]" captionTargetPageId="5" captionText="FIGURE 2. Position of reproductive morphology of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. ac: atrial cornua; an, anus; e, eyes; fg, female gonopore; ic, intestinal caecum; mg, male gonopore; od, oviduct; os: ovisac; ps, posterior sucker; r, rectum; s, scute; sd, sperm duct; t, testisac, vd. vas deferens." httpUri="https://zenodo.org/record/1212242/files/figure.png" pageId="10" pageNumber="71">Figs. 2E</figureCitation>
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–
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<figureCitation id="37E62A45FF8B4441E67559E6FC0E3000" box="[890,921,1270,1294]" captionStart="FIGURE 7" captionStartId="10.[151,250,1172,1194]" captionTargetBox="[155,1432,382,1148]" captionTargetId="figure@10.[151,1436,373,1150]" captionTargetPageId="10" captionText="FIGURE 7. Eye position and shape of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitousn. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp." httpUri="https://zenodo.org/record/1212252/files/figure.png" pageId="10" pageNumber="71">7E</figureCitation>
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)
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</paragraph>
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<subSubSection id="E7C7654BFF8B4441E6B559E5FAA43000" box="[954,1331,1269,1294]" pageId="10" pageNumber="71" type="materials_examined">
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<paragraph id="AF6236C0FF8B4441E6B559E5FAA43000" blockId="10.[151,1437,1269,2017]" box="[954,1331,1269,1294]" pageId="10" pageNumber="71">
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(
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<typeStatus id="70668862FF8B4441E6CD59E5FBA73000" box="[962,1072,1269,1294]" pageId="10" pageNumber="71" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="C9CCAE05FF8B4441E14359E6FB043000" box="[1100,1171,1270,1294]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="71">ANSP</collectionCode>
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GI 19496)
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</paragraph>
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</subSubSection>
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<subSubSection id="E7C7654BFF8B4441E04359E6FC2A303C" pageId="10" pageNumber="71" type="nomenclature">
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<paragraph id="AF6236C0FF8B4441E04359E6FC2A303C" blockId="10.[151,1437,1269,2017]" pageId="10" pageNumber="71">
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(LSID:
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<emphasis id="9DA9EAD2FF8B4441E5985809FC21303C" box="[151,950,1305,1330]" italics="true" pageId="10" pageNumber="71">
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urn:lsid:zoobank.org:act:
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<accessionNumber id="B08EAB23FF8B4441E4A55809FD92303C" box="[426,517,1305,1330]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F55674" pageId="10" pageNumber="71" type="EnaNcbi">F55674</accessionNumber>
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B
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<date id="DB631000FF8B4441E7185809FDEA303C" box="[535,637,1305,1330]" pageId="10" pageNumber="71" value="1767-06-04">6-1767-4</date>
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F64-ABEF-0347122944E0
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)
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<subSubSection id="E7C7654BFF8B4441E6B25809FAFA3297" pageId="10" pageNumber="71" type="description">
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<paragraph id="AF6236C0FF8B4441E6B25809FA843378" blockId="10.[151,1437,1269,2017]" pageId="10" pageNumber="71">
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—Dorsal body light to yellowish-grey, sometimes with a greenish tint. Black dots on dorsal surface. Ventral coloration and annuli numbers same as
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<taxonomicName id="68DD4D43FF8B4441E08E582EFEA23074" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="serendipitious" status="sp. nov.">
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<emphasis id="9DA9EAD2FF8B4441E08E582EFEA23074" italics="true" pageId="10" pageNumber="71">H. serendipitious</emphasis>
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</taxonomicName>
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<emphasis id="9DA9EAD2FF8B4441E4335871FEEB3074" bold="true" box="[316,380,1377,1402]" pageId="10" pageNumber="71">
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<taxonomicNameLabel id="869A57A9FF8B4441E4335871FEEB3074" box="[316,380,1377,1402]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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One pair of eyes close to each other on segment III (
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<figureCitation id="37E62A45FF8B4441E6D75871FBB83074" box="[984,1071,1377,1402]" captionStart="FIGURE 7" captionStartId="10.[151,250,1172,1194]" captionTargetBox="[155,1432,382,1148]" captionTargetId="figure@10.[151,1436,373,1150]" captionTargetPageId="10" captionText="FIGURE 7. Eye position and shape of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitousn. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp." httpUri="https://zenodo.org/record/1212252/files/figure.png" pageId="10" pageNumber="71">Fig. 7E</figureCitation>
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). Male and female gonopores in position similar to
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<taxonomicName id="68DD4D43FF8B4441E4635896FDBB3090" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[364,556,1413,1438]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="serendipitious" status="sp. nov.">
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<emphasis id="9DA9EAD2FF8B4441E4635896FDBB3090" box="[364,556,1413,1438]" italics="true" pageId="10" pageNumber="71">H. serendipitious</emphasis>
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</taxonomicName>
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||
<emphasis id="9DA9EAD2FF8B4441E73B5895FDE43090" bold="true" box="[564,627,1413,1438]" pageId="10" pageNumber="71">
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<taxonomicNameLabel id="869A57A9FF8B4441E73B5895FDE43090" box="[564,627,1413,1438]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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Six pairs of testisacs intersegmentally arranged from XII/XIII to XVII/ XVIII; shape similar to
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<taxonomicName id="68DD4D43FF8B4441E4B758BAFDD230CC" box="[440,581,1449,1474]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="stagnalis">
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<emphasis id="9DA9EAD2FF8B4441E4B758BAFDD230CC" box="[440,581,1449,1474]" italics="true" pageId="10" pageNumber="71">H. stagnalis</emphasis>
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</taxonomicName>
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but smaller. Ejaculatory bulbs (atrial cornua) moderately longer, ovoidcylindrical, lying at a much lower level by the sides of the atrium, connected by slender ejaculatory ducts with sharp turn backwards into atrium in XIth segment. Ovisacs large, connected with long and curled common oviduct. Oviduct length almost equal to ovisacs (
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<figureCitation id="37E62A45FF8B4441E7625B05FD463320" box="[621,721,1557,1582]" captionStart="FIGURE 2" captionStartId="5.[151,250,1920,1942]" captionTargetBox="[165,1433,419,1907]" captionTargetId="figure@5.[156,1433,404,1907]" captionTargetPageId="5" captionText="FIGURE 2. Position of reproductive morphology of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. ac: atrial cornua; an, anus; e, eyes; fg, female gonopore; ic, intestinal caecum; mg, male gonopore; od, oviduct; os: ovisac; ps, posterior sucker; r, rectum; s, scute; sd, sperm duct; t, testisac, vd. vas deferens." httpUri="https://zenodo.org/record/1212242/files/figure.png" pageId="10" pageNumber="71">Figs. 2E</figureCitation>
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,
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<figureCitation id="37E62A45FF8B4441E7EF5B06FC943320" box="[736,771,1558,1582]" captionStart="FIGURE 4" captionStartId="7.[151,250,1938,1960]" captionTargetBox="[158,1424,688,1887]" captionTargetId="figure@7.[151,1437,681,1892]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of reproductive system of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. a, atrium; ac: atrial cornua; os: ovisac; od, oviduct; sd, sperm duct; t, testisac, vd. vas deferens." httpUri="https://zenodo.org/record/1212246/files/figure.png" pageId="10" pageNumber="71">4E</figureCitation>
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). Six pairs crop caeca, 6th directed backwards and short, extending up to fourth intestinal caecum. Four pairs intestinal caecum, first two pairs slightly curved upward, third pair flat and last pair curved downward. Rectum short, “S” shaped, similar
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<taxonomicName id="68DD4D43FF8B4441E6EE5B4EFBFF3378" box="[993,1128,1629,1654]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="stagnalis">
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<emphasis id="9DA9EAD2FF8B4441E6EE5B4EFBFF3378" box="[993,1128,1629,1654]" italics="true" pageId="10" pageNumber="71">H. stagnalis</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="37E62A45FF8B4441E1785B4DFB4F3378" box="[1143,1240,1629,1654]" captionStart="FIGURE 3" captionStartId="6.[151,250,1785,1807]" captionTargetBox="[151,1410,201,1764]" captionTargetId="figure@6.[151,1436,193,1764]" captionTargetPageId="6" captionText="FIGURE 3. Position of digestive system of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. ac: atrial cornua; an, anus; cc, corp caecum; e, eyes; ic, intestinal caecum; p, proboscis;ps, posterior sucker; r, rectum; s, scute." httpUri="https://zenodo.org/record/1212244/files/figure.png" pageId="10" pageNumber="71">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="37E62A45FF8B4441E1EB5B4DFA913378" box="[1252,1286,1629,1654]" captionStart="FIGURE 5" captionStartId="8.[151,250,1521,1543]" captionTargetBox="[151,1426,194,1507]" captionTargetId="figure@8.[151,1436,185,1507]" captionTargetPageId="8" captionText="FIGURE 5. Digestive system of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. an, anus; cc, corp caecum; ic, intestinal caecum, r, rectum." httpUri="https://zenodo.org/record/1212248/files/figure.png" pageId="10" pageNumber="71">5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="AF6236C0FF8B4441E5C85B91FAFA3297" blockId="10.[151,1437,1269,2017]" pageId="10" pageNumber="71">
|
||
The dorsal nuchal scute of
|
||
<taxonomicName id="68DD4D43FF8B4441E7055B92FD013394" box="[522,662,1665,1690]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="stagnalis">
|
||
<emphasis id="9DA9EAD2FF8B4441E7055B92FD013394" box="[522,662,1665,1690]" italics="true" pageId="10" pageNumber="71">H. stagnalis</emphasis>
|
||
</taxonomicName>
|
||
from Europe has been depicted in a SEM-micrograph by
|
||
<bibRefCitation id="CB4C4B31FF8B4441E0475B92FEAD33B0" pageId="10" pageNumber="71" refId="ref9288" refString="Sawyer, R. T. (1986) Leech Biology and Behavior. Vol. I - III. Clarendon Press, Oxford, xiv + 1065 pp." type="book">Sawyer (1986, p. 538)</bibRefCitation>
|
||
. Additional SEM identified differences in scute morphology and position between species (
|
||
<figureCitation id="37E62A45FF8B4441E0465BB5FA1933B0" box="[1353,1422,1701,1726]" captionStart="FIGURE 8" captionStartId="11.[151,250,1300,1322]" captionTargetBox="[161,1423,311,1275]" captionTargetId="figure@11.[151,1435,301,1286]" captionTargetPageId="11" captionText="FIGURE 8. Representative scanning electron micrographs of the chitonous plate (scute) of Helobdella stagnalis-like species. A, of Helobdella modesta. B, Helobdella serendipitous n.sp. C, Helobdella n. sp. (Montana).D, Helobdella n. sp. (Oregon)." httpUri="https://zenodo.org/record/1212254/files/figure.png" pageId="10" pageNumber="71">Fig. 8</figureCitation>
|
||
). Specifically, the
|
||
<emphasis id="9DA9EAD2FF8B4441E45E5BDDFECB33EA" box="[337,348,1741,1764]" italics="true" pageId="10" pageNumber="71">s</emphasis>
|
||
cute
|
||
<emphasis id="9DA9EAD2FF8B4441E49E5BDDFDBD33EB" box="[401,554,1740,1765]" italics="true" pageId="10" pageNumber="71">
|
||
of
|
||
<taxonomicName id="68DD4D43FF8B4441E4A25BDCFDBD33EB" box="[429,554,1740,1765]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="genus">Helobdella</taxonomicName>
|
||
</emphasis>
|
||
spp. ranged from rhomboidal to hexagonal, settled on the12th and extending to the 13th annulus The
|
||
<taxonomicName id="68DD4D43FF8B4441E4AF5BE1FDB33207" box="[416,548,1776,1801]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="modesta">
|
||
<emphasis id="9DA9EAD2FF8B4441E4AF5BE1FDB33207" box="[416,548,1776,1801]" italics="true" pageId="10" pageNumber="71">H. modesta</emphasis>
|
||
</taxonomicName>
|
||
scute was square-shaped, settled on the 12th annulus (on the VII-a1) and sometimes extending up to half the 13th annulus (
|
||
<figureCitation id="37E62A45FF8B4441E7C25A04FCB13223" box="[717,806,1812,1837]" captionStart="FIGURE 6" captionStartId="9.[151,250,1224,1246]" captionTargetBox="[151,1437,196,1202]" captionTargetId="figure@9.[151,1437,193,1202]" captionTargetPageId="9" captionText="FIGURE 6. View of the scute (Nuchal plate) of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdellaserendipitous n. sp.; D, Helobdellaeriensis n. sp.; E, Helobdella echoensisn. sp. s, scute; sb: scute base." httpUri="https://zenodo.org/record/1212250/files/figure.png" pageId="10" pageNumber="71">Fig. 6B</figureCitation>
|
||
). The
|
||
<taxonomicName id="68DD4D43FF8B4441E6615A05FBB93223" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[878,1070,1812,1837]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="serendipitious">
|
||
<emphasis id="9DA9EAD2FF8B4441E6615A05FBB93223" box="[878,1070,1812,1837]" italics="true" pageId="10" pageNumber="71">H. serendipitious</emphasis>
|
||
</taxonomicName>
|
||
scute was pentagonal, settled in the dorsal surface of segment VII-a2 (on annulus 12) (
|
||
<figureCitation id="37E62A45FF8B4441E7F05A28FCCE325F" box="[767,857,1848,1873]" captionStart="FIGURE 6" captionStartId="9.[151,250,1224,1246]" captionTargetBox="[151,1437,196,1202]" captionTargetId="figure@9.[151,1437,193,1202]" captionTargetPageId="9" captionText="FIGURE 6. View of the scute (Nuchal plate) of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdellaserendipitous n. sp.; D, Helobdellaeriensis n. sp.; E, Helobdella echoensisn. sp. s, scute; sb: scute base." httpUri="https://zenodo.org/record/1212250/files/figure.png" pageId="10" pageNumber="71">Fig. 6C</figureCitation>
|
||
). The
|
||
<emphasis id="9DA9EAD2FF8B4441E6AE5A28FB543241" box="[929,1219,1848,1873]" italics="true" pageId="10" pageNumber="71">
|
||
<taxonomicName id="68DD4D43FF8B4441E6AE5A28FBEB325E" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[929,1148,1848,1873]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="eriensis" status="sp. nov.">Helobdella eriensis</taxonomicName>
|
||
<emphasis id="9DA9EAD2FF8B4441E18B5A28FB543241" bold="true" box="[1156,1219,1848,1871]" italics="true" pageId="10" pageNumber="71">
|
||
<taxonomicNameLabel id="869A57A9FF8B4441E18B5A28FB543241" box="[1156,1219,1848,1871]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</emphasis>
|
||
(Farm Pond) scute was similar to
|
||
<taxonomicName id="68DD4D43FF8B4441E4445A4DFD87327B" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[331,528,1884,1909]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="serendipitious" status="sp. nov.">
|
||
<emphasis id="9DA9EAD2FF8B4441E4445A4DFD87327B" box="[331,528,1884,1909]" italics="true" pageId="10" pageNumber="71">H. serendipitious</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DA9EAD2FF8B4441E7125A4CFDF6327B" bold="true" box="[541,609,1884,1909]" pageId="10" pageNumber="71">
|
||
<taxonomicNameLabel id="869A57A9FF8B4441E7125A4CFDF6327B" box="[541,609,1884,1909]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but settled in the 12th and up to all of 13th annulus (
|
||
<figureCitation id="37E62A45FF8B4441E1F75A4CFAC0327B" box="[1272,1367,1884,1909]" captionStart="FIGURE 6" captionStartId="9.[151,250,1224,1246]" captionTargetBox="[151,1437,196,1202]" captionTargetId="figure@9.[151,1437,193,1202]" captionTargetPageId="9" captionText="FIGURE 6. View of the scute (Nuchal plate) of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdellaserendipitous n. sp.; D, Helobdellaeriensis n. sp.; E, Helobdella echoensisn. sp. s, scute; sb: scute base." httpUri="https://zenodo.org/record/1212250/files/figure.png" pageId="10" pageNumber="71">Fig. 6D</figureCitation>
|
||
). The
|
||
<emphasis id="9DA9EAD2FF8B4441E5985A90FE5B3299" box="[151,460,1920,1945]" italics="true" pageId="10" pageNumber="71">
|
||
<taxonomicName id="68DD4D43FF8B4441E5985A90FE103297" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[151,391,1920,1945]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="echoensis" status="sp. nov.">Helobdella echoensis</taxonomicName>
|
||
<emphasis id="9DA9EAD2FF8B4441E4815A90FE5B3299" bold="true" box="[398,460,1920,1943]" italics="true" pageId="10" pageNumber="71">
|
||
<taxonomicNameLabel id="869A57A9FF8B4441E4815A90FE5B3299" box="[398,460,1920,1943]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="6D19F822FF8B4441E4D55A90FD823297" box="[474,533,1920,1945]" country="United Kingdom" name="York" pageId="10" pageNumber="71">York</collectingRegion>
|
||
, PA) was similar to
|
||
<taxonomicName id="68DD4D43FF8B4441E7F95A91FC213297" authority="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain, 2018" authorityName="Saglam & Kutschera & Saunders & Saidel & Balombini & Shain" authorityYear="2018" box="[758,950,1920,1945]" class="Clitellata" family="Glossiphoniidae" genus="Helobdella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchobdellida" pageId="10" pageNumber="71" phylum="Annelida" rank="species" species="serendipitious" status="sp. nov.">
|
||
<emphasis id="9DA9EAD2FF8B4441E7F95A91FC213297" box="[758,950,1920,1945]" italics="true" pageId="10" pageNumber="71">H. serendipitious</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DA9EAD2FF8B4441E6B25A90FC6C3297" bold="true" box="[957,1019,1920,1945]" pageId="10" pageNumber="71">
|
||
<taxonomicNameLabel id="869A57A9FF8B4441E6B25A90FC6C3297" box="[957,1019,1920,1945]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but mushroom shaped (
|
||
<figureCitation id="37E62A45FF8B4441E0065A90FAF73297" box="[1289,1376,1920,1945]" captionStart="FIGURE 5" captionStartId="8.[151,250,1521,1543]" captionTargetBox="[151,1426,194,1507]" captionTargetId="figure@8.[151,1436,185,1507]" captionTargetPageId="8" captionText="FIGURE 5. Digestive system of Helobdella species. A, Helobdella stagnalis; B, Helobdella modesta; C, Helobdella serendipitous n. sp.; D, Helobdella eriensis n. sp.; E, Helobdella echoensis n. sp. an, anus; cc, corp caecum; ic, intestinal caecum, r, rectum." httpUri="https://zenodo.org/record/1212248/files/figure.png" pageId="10" pageNumber="71">Fig. 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |