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154 lines
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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.32.38161" ID-GBIF-Dataset="23cdd843-223d-4b59-b7c6-307e35e71c07" ID-GBIF-Taxon="159168994" ID-Pensoft-Pub="1314-2615-32-15" ID-Pensoft-UUID="6291936E45B856A1AA18C2B24BDDDC76" ID-ZooBank="731EF530E43D4FB8AAA39B4A0AD1AE09" LSID="urn:lsid:zoobank.org:act:9737DDFE-2F14-4F5A-96FF-8E47BB053387" ModsDocID="1314-2615-32-15" checkinTime="1566303228586" checkinUser="pensoft" docAuthor="Shear, William A. & Steinmann, David B." docDate="2019" docId="2714317BCE3B5179B5E58DB57959DA37" docLanguage="en" docName="SubterBiol 32: 15-32" docOrigin="Subterranean Biology 32" docSource="http://dx.doi.org/10.3897/subtbiol.32.38161" docTitle="Coloradesmus beckleyi Shear & Steinmann, sp. nov." docType="treatment" docUuid="9737DDFE-2F14-4F5A-96FF-8E47BB053387" docUuidSource="ZooBank" docVersion="4" id="6291936E45B856A1AA18C2B24BDDDC76" lastPageId="10" lastPageNumber="25" masterDocId="6291936E45B856A1AA18C2B24BDDDC76" masterDocTitle="Cave millipedes of the United States. XV. Coloradesmus gen. nov. (Diplopoda, Polydesmida, Macrosternodesmidae), and four new species from caves in Colorado, USA" masterLastPageNumber="32" masterPageNumber="15" pageId="9" pageNumber="24" updateTime="1643504485831" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cave millipedes of the United States. XV. Coloradesmus gen. nov. (Diplopoda, Polydesmida, Macrosternodesmidae), and four new species from caves in Colorado, USA</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shear, William A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Steinmann, David B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Subterranean Biology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>32</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>32</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/subtbiol.32.38161</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.32.38161</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2615-32-15</mods:identifier>
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<mods:identifier type="ZooBank">731EF530E43D4FB8AAA39B4A0AD1AE09</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159168994" LSID="urn:lsid:zoobank.org:act:9737DDFE-2F14-4F5A-96FF-8E47BB053387" httpUri="http://treatment.plazi.org/id/2714317BCE3B5179B5E58DB57959DA37" lastPageId="10" lastPageNumber="25" pageId="9" pageNumber="24">
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<subSubSection pageId="9" pageNumber="24" type="nomenclature">
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<paragraph pageId="9" pageNumber="24">
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<taxonomicName LSID="2714317b-ce3b-5179-b5e5-8db57959da37" authority="Shear & Steinmann" authorityName="Shear & Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus beckleyi" order="Polydesmida" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="beckleyi">Coloradesmus beckleyi Shear & Steinmann</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="24">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 21–25" captionStartId="F6" captionText="Figures 21 - 25. Coloradesmus species males. 21 C. manitou gonopods, posterior view 22 - 24 C. lasunda male 22 head, collum and anterior four rings, dorsal view 23 collum, dorsal view 24 midbody ring, dorsal view 25 telson, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures21-25" httpUri="https://binary.pensoft.net/fig/326949" pageId="9" pageNumber="24">Figs 22-25</figureCitation>
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,
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<figureCitation captionStart="Figures 26, 27" captionStartId="F7" captionText="Figures 26, 27. Coloradesmus lasunda male. 26 Right gonopod, mesal view 27 right gonopod, lateral view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures26-27" httpUri="https://binary.pensoft.net/fig/326950" pageId="9" pageNumber="24">26, 27</figureCitation>
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,
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<figureCitation captionStart="Figures 28, 29" captionStartId="F8" captionText="Figures 28, 29. Coloradesmus species gonopods, posterior views. 28 C. lasunda 29 C. warneri. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="9" pageNumber="24">28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="types">
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<paragraph pageId="10" pageNumber="25">
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<pageBreakToken pageId="10" pageNumber="25" start="start">Types</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="10" pageNumber="25">
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Male holotype and male and female paratypes from La Sunder Cave, Garfield Co.: Colorado, 7500' (2287 m) asl,
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<geoCoordinate degrees="39" direction="north" minutes="42" orientation="latitude" precision="925" value="39.7">39°42'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="west" minutes="10" orientation="longitude" precision="925" value="-107.166664">107°10'W</geoCoordinate>
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, in organic matter and under cardboard along the first 1000' of passage, collected 31 July 1999 by D. Steinmann.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="diagnosis">
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<paragraph pageId="10" pageNumber="25">Diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="25">
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A small species of
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<taxonomicName authority="Shear & Steinmann, 2019" authorityName="Shear & Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus" order="Polydesmida" pageId="10" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="25">Coloradesmus</emphasis>
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</taxonomicName>
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, as is the following species, but differing from
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<taxonomicName genus="C." lsidName="C. warneri" pageId="10" pageNumber="25" rank="species" species="warneri">
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<emphasis italics="true" pageId="10" pageNumber="25">C. warneri</emphasis>
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</taxonomicName>
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in details of the gonopods, as shown in
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<figureCitation captionStart="Figures 28, 29" captionStartId="F8" captionText="Figures 28, 29. Coloradesmus species gonopods, posterior views. 28 C. lasunda 29 C. warneri. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="10" pageNumber="25">Figs 28-29</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="etymology">
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<paragraph pageId="10" pageNumber="25">Etymology.</paragraph>
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<paragraph pageId="10" pageNumber="25">The species epithet honors Steve Beckley, for his dedication to cave conservation and education as the owner of Glenwood Caverns in nearby Glenwood Springs.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="description">
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<paragraph pageId="10" pageNumber="25">Description.</paragraph>
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<paragraph pageId="10" pageNumber="25">
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<emphasis bold="true" italics="true" pageId="10" pageNumber="25">Male holotype.</emphasis>
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Length, about 4 mm, greatest width 0.4 mm. Head with short, sparse setae; basal joint of mandible exceeding width of collum (
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<figureCitation captionStart="Figures 21–25" captionStartId="F6" captionText="Figures 21 - 25. Coloradesmus species males. 21 C. manitou gonopods, posterior view 22 - 24 C. lasunda male 22 head, collum and anterior four rings, dorsal view 23 collum, dorsal view 24 midbody ring, dorsal view 25 telson, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures21-25" httpUri="https://binary.pensoft.net/fig/326949" pageId="10" pageNumber="25">Fig. 22</figureCitation>
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). Collum relatively short, posterior margin evenly, concavely curved (
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<figureCitation captionStart="Figures 21–25" captionStartId="F6" captionText="Figures 21 - 25. Coloradesmus species males. 21 C. manitou gonopods, posterior view 22 - 24 C. lasunda male 22 head, collum and anterior four rings, dorsal view 23 collum, dorsal view 24 midbody ring, dorsal view 25 telson, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures21-25" httpUri="https://binary.pensoft.net/fig/326949" pageId="10" pageNumber="25">Fig. 23</figureCitation>
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); collum setae short, clavate. Metazonites with somewhat suppressed but distinct paranota bearing obsolete teeth (
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<figureCitation captionStart="Figures 21–25" captionStartId="F6" captionText="Figures 21 - 25. Coloradesmus species males. 21 C. manitou gonopods, posterior view 22 - 24 C. lasunda male 22 head, collum and anterior four rings, dorsal view 23 collum, dorsal view 24 midbody ring, dorsal view 25 telson, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures21-25" httpUri="https://binary.pensoft.net/fig/326949" pageId="10" pageNumber="25">Figs 22-25</figureCitation>
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); setae distinctly clavate, short, set on prominent tubercles, rows even, uninterrupted. Telson (
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<figureCitation captionStart="Figures 21–25" captionStartId="F6" captionText="Figures 21 - 25. Coloradesmus species males. 21 C. manitou gonopods, posterior view 22 - 24 C. lasunda male 22 head, collum and anterior four rings, dorsal view 23 collum, dorsal view 24 midbody ring, dorsal view 25 telson, dorsal view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures21-25" httpUri="https://binary.pensoft.net/fig/326949" pageId="10" pageNumber="25">Fig. 25</figureCitation>
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) with few setae, epiproct indistinct. Gonopods (
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<figureCitation captionStart="Figures 26, 27" captionStartId="F7" captionText="Figures 26, 27. Coloradesmus lasunda male. 26 Right gonopod, mesal view 27 right gonopod, lateral view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures26-27" httpUri="https://binary.pensoft.net/fig/326950" pageId="10" pageNumber="25">Figs 26</figureCitation>
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-
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<figureCitation captionStart="Figures 28, 29" captionStartId="F8" captionText="Figures 28, 29. Coloradesmus species gonopods, posterior views. 28 C. lasunda 29 C. warneri. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="10" pageNumber="25">28</figureCitation>
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) with coxae rather oval, immovable; acropodite (a,
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<figureCitation captionStart="Figures 26, 27" captionStartId="F7" captionText="Figures 26, 27. Coloradesmus lasunda male. 26 Right gonopod, mesal view 27 right gonopod, lateral view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures26-27" httpUri="https://binary.pensoft.net/fig/326950" pageId="10" pageNumber="25">Figs 26</figureCitation>
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-
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<figureCitation captionStart="Figures 28, 29" captionStartId="F8" captionText="Figures 28, 29. Coloradesmus species gonopods, posterior views. 28 C. lasunda 29 C. warneri. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="10" pageNumber="25">28</figureCitation>
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) robust, blocky, pore without cuticular fimbriae, endomerite evidently absent. Prefemoral process (pfp,
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<figureCitation captionStart="Figures 26, 27" captionStartId="F7" captionText="Figures 26, 27. Coloradesmus lasunda male. 26 Right gonopod, mesal view 27 right gonopod, lateral view. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures26-27" httpUri="https://binary.pensoft.net/fig/326950" pageId="10" pageNumber="25">Figs 26</figureCitation>
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-
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<figureCitation captionStart="Figures 28, 29" captionStartId="F8" captionText="Figures 28, 29. Coloradesmus species gonopods, posterior views. 28 C. lasunda 29 C. warneri. See text for explanation of labels." figureDoi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="10" pageNumber="25">28</figureCitation>
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) exceeding acropodite, narrow, with distinct acute hook at tip.
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</paragraph>
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<caption doi="10.3897/subtbiol.32.38161.figures26-27" httpUri="https://binary.pensoft.net/fig/326950" pageId="10" pageNumber="25" start="Figures 26, 27" startId="F7">
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<paragraph pageId="10" pageNumber="25">
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Figures 26, 27.
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<taxonomicName authority="Shear & Steinmann, 2019" authorityName="Shear & Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus lasunda" order="Polydesmida" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="lasunda">
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<emphasis italics="true" pageId="10" pageNumber="25">Coloradesmus lasunda</emphasis>
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</taxonomicName>
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male.
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<emphasis bold="true" pageId="10" pageNumber="25">26</emphasis>
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Right gonopod, mesal view
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<emphasis bold="true" pageId="10" pageNumber="25">27</emphasis>
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right gonopod, lateral view. See text for explanation of labels.
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</paragraph>
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</caption>
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<caption doi="10.3897/subtbiol.32.38161.figures28-29" httpUri="https://binary.pensoft.net/fig/326951" pageId="10" pageNumber="25" start="Figures 28, 29" startId="F8">
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<paragraph pageId="10" pageNumber="25">
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Figures 28, 29.
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<taxonomicName authority="Shear & Steinmann, 2019" authorityName="Shear & Steinmann" authorityYear="2019" class="Diplopoda" family="Macrosternodesmidae" genus="Coloradesmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coloradesmus" order="Polydesmida" pageId="10" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="25">Coloradesmus</emphasis>
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</taxonomicName>
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species gonopods, posterior views.
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<emphasis bold="true" pageId="10" pageNumber="25">28</emphasis>
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<taxonomicName genus="C." lsidName="C. lasunda" pageId="10" pageNumber="25" rank="species" species="lasunda">
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<emphasis italics="true" pageId="10" pageNumber="25">C. lasunda</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="10" pageNumber="25">29</emphasis>
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<taxonomicName genus="C." lsidName="C. warneri" pageId="10" pageNumber="25" rank="species" species="warneri">
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<emphasis italics="true" pageId="10" pageNumber="25">C. warneri</emphasis>
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</taxonomicName>
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. See text for explanation of labels.
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</paragraph>
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</caption>
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<paragraph pageId="10" pageNumber="25">
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<emphasis bold="true" italics="true" pageId="10" pageNumber="25">Female paratype.</emphasis>
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Similar in all nonsexual characters to male.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="25" type="distribution">
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<paragraph pageId="10" pageNumber="25">Distribution.</paragraph>
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<paragraph pageId="10" pageNumber="25">Known only from the type locality.</paragraph>
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</subSubSection>
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</treatment>
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</document> |