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<document id="33A824225D30F73390C20F3B066E6F8F" ID-DOI="10.5281/zenodo.187264" ID-GBIF-Dataset="9b9b4d51-711d-4aad-b397-df9b04b465e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="187264" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461066418959" checkinUser="plazi" docAuthor="Grimaldi, David A., Cumming, Jeffrey M. &amp; Arillo, Antonio" docDate="2009" docId="270087AD5227FFBDFF2268F238B4FDAC" docLanguage="en" docName="zt02078p054.pdf" docOrigin="Zootaxa 2078" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Chimeromyina concilia Arillo &amp; Grimaldi, sp. n." docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="DB39FFD55237FFAFFFB56E573B0AFFB8" masterDocTitle="Chimeromyiidae, a new family of Eremoneuran Diptera from the Cretaceous" masterLastPageNumber="54" masterPageNumber="34" pageNumber="50" updateTime="1698583433394" updateUser="plazi">
<mods:mods id="7B41A746336C66268F2CDEEC41535C07" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="788EE613229A4743E4614374FA8F020A">
<mods:title id="FFF8898FB973B4134F719DD82C3516B6">Chimeromyiidae, a new family of Eremoneuran Diptera from the Cretaceous</mods:title>
</mods:titleInfo>
<mods:name id="FDB661CEF8A58642E139B66CAA3B5D75" type="personal">
<mods:role id="3AD45C4A898F168C975F14FED2CEA057">
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<mods:namePart id="BB4C9FCF55FDCBA0C38B0271BE838626">Grimaldi, David A.</mods:namePart>
</mods:name>
<mods:name id="472A0D5DD85F4D34808EA9A39D2BF70A" type="personal">
<mods:role id="A88BAA074B9035634E8F7BE4A46DCC61">
<mods:roleTerm id="17CF2454D6C58AA9C1B28F4DE3A2DB1E">Author</mods:roleTerm>
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<mods:namePart id="AFD253BBA4BD05F2822E87D58D31A395">Cumming, Jeffrey M.</mods:namePart>
</mods:name>
<mods:name id="376F5E36D08C546DB8EEB52C8AFB7CDF" type="personal">
<mods:role id="8DCBD7CC52B6CA0AFEB1A2E7E02B60C0">
<mods:roleTerm id="88B4683F25245280E9BD0726BC94D33F">Author</mods:roleTerm>
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<mods:namePart id="D7D72541699C7B1D0D718257CEE9F129">Arillo, Antonio</mods:namePart>
</mods:name>
<mods:typeOfResource id="6AAD79C15BBFC3C79E35B0B3F46F0923">text</mods:typeOfResource>
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<mods:title id="208BB61D581AD29DBA32811688465BEC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FEEDCA48A52C594583AE224463327F75">
<mods:date id="BC635BB6C37C0B6B201D6814391E15E0">2009</mods:date>
<mods:detail id="77DE552CCEB30CD1C61300CF95BA4555" type="volume">
<mods:number id="20EBE7823D4EBEC0ABA463BA1B75BC5F">2078</mods:number>
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<mods:classification id="8DA570B7AEDC19205BD57934ACC8080A">journal article</mods:classification>
<mods:identifier id="26114426D310ECB0D76AD89E0DC7FDE7" type="DOI">10.5281/zenodo.187264</mods:identifier>
<mods:identifier id="5235CD7E89809A86A6D35607D4CF7071" type="GBIF-Dataset">9b9b4d51-711d-4aad-b397-df9b04b465e8</mods:identifier>
<mods:identifier id="BFD7B02CB4EC51A7C94F4BE52A78655A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EA0C6EBCBEF32010313B2788DC55B413" type="Zenodo-Dep">187264</mods:identifier>
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<treatment id="270087AD5227FFBDFF2268F238B4FDAC" ID-DOI="http://doi.org/10.5281/zenodo.6222210" ID-GBIF-Taxon="119615002" ID-Zenodo-Dep="6222210" LSID="urn:lsid:plazi:treatment:270087AD5227FFBDFF2268F238B4FDAC" httpUri="http://treatment.plazi.org/id/270087AD5227FFBDFF2268F238B4FDAC" lastPageId="18" lastPageNumber="52" pageId="16" pageNumber="50">
<subSubSection id="E7B365305227FFBFFF2268F23BF7F95B" pageId="16" pageNumber="50" type="nomenclature">
<paragraph id="AF1636BB5227FFBFFF2268F239E5F907" blockId="16.[151,751,1701,1763]" box="[151,751,1701,1727]" pageId="16" pageNumber="50">
<heading id="F45E81D75227FFBFFF2268F239E5F907" bold="true" box="[151,751,1701,1727]" fontSize="11" level="1" pageId="16" pageNumber="50" reason="1">
<emphasis id="9DDDEAA95227FFBFFF2268F239E5F907" bold="true" box="[151,751,1701,1727]" pageId="16" pageNumber="50">
<taxonomicName id="68A94D385227FFBFFF2268F239AAF907" authority="Arillo &amp; Grimaldi" authorityName="Arillo &amp; Grimaldi" box="[151,672,1701,1727]" class="Insecta" family="Opetiidae" genus="Chimeromyina" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="50" phylum="Arthropoda" rank="species" species="concilia" status="sp. nov.">
<emphasis id="9DDDEAA95227FFBFFF2268F23AB9F907" bold="true" box="[151,435,1701,1727]" italics="true" pageId="16" pageNumber="50">Chimeromyina concilia</emphasis>
Arillo &amp; Grimaldi
</taxonomicName>
,
<taxonomicNameLabel id="86EE57D25227FFBFFD1868F239E5F907" box="[685,751,1701,1727]" pageId="16" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="AF1636BB5227FFBFFF22689E3BF7F95B" blockId="16.[151,751,1701,1763]" box="[151,253,1737,1763]" pageId="16" pageNumber="50">
(
<figureCitation id="37922A3E5227FFBFFF2A689E3BFEF95B" box="[159,244,1737,1763]" captionStart="FIGURE 13" captionStartId="17.[151,257,1758,1782]" captionTargetBox="[276,1296,203,1729]" captionTargetId="figure@17.[276,1312,197,1733]" captionTargetPageId="17" captionText="FIGURE 13. Chimeromyina concilia, n. gen., n. sp., in Spanish amber. Holotype, MCNA 8882. A. Dorsal view of anterior half of body. B. Detail of antennae and vertex of head. C. Left habitus (terminalia lost at surface of amber)." httpUri="https://zenodo.org/record/187277/files/figure.png" pageId="16" pageNumber="50">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E7B365305227FFBFFF2269413AB1F888" box="[151,443,1814,1840]" pageId="16" pageNumber="50" type="diagnosis">
<paragraph id="AF1636BB5227FFBFFF2269413AB1F888" blockId="16.[151,1437,1814,2033]" box="[151,443,1814,1840]" pageId="16" pageNumber="50">
<emphasis id="9DDDEAA95227FFBFFF2269413A1AF888" bold="true" box="[151,272,1814,1840]" pageId="16" pageNumber="50">Diagnosis</emphasis>
. As for genus.
</paragraph>
</subSubSection>
<subSubSection id="E7B365305227FFBDFF70696B3802FEEA" lastPageId="18" lastPageNumber="52" pageId="16" pageNumber="50" type="description">
<paragraph id="AF1636BB5227FFBDFF70696B3802FEEA" blockId="16.[151,1437,1814,2033]" lastBlockId="18.[151,1437,152,532]" lastPageId="18" lastPageNumber="52" pageId="16" pageNumber="50">
<emphasis id="9DDDEAA95227FFBFFF70696B3A69F8EE" bold="true" box="[197,355,1852,1878]" pageId="16" pageNumber="50">Description.</emphasis>
HEAD: Fairly broad, width more than 1.5x width of notum; eyes large, bare, facets not differentiated. Ocelli on small tubercle, tubercle with 2 pairs long setae. Antenna with apical arista; 3 aristomeres present. Vibrissae not present. THORAX:
<quantity id="68519B5E5227FFBFFCA869DD388DF81C" box="[797,903,1930,1956]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="16" pageNumber="50" unit="mm" value="0.44">0.44 mm</quantity>
long, bristly, each side of notum with 2 long notopleural setae and supra-alar seta; 2 pairs of dorsocentrals, row of long setulae anterior to dorsocentrals. Acrostichal setulae in one, extensive row, extended to posterior margin of notum. Scutellum with 2 pairs setae, apical pair long (ca.
<date id="DB17107B5226FFBEFE65691D3AE7F8DC" box="[464,493,1866,1892]" pageId="17" pageNumber="51">2x</date>
length of other pair), cruciate. Wing
<quantity id="68519B5E5226FFBEFC15691D3F03F8DC" box="[928,1033,1866,1892]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="17" pageNumber="51" unit="mm" value="0.93">0.93 mm</quantity>
long; fairly broad, width less than half the length. Vein C extended to apex of M1, though obscured because tip of wing is folded; Sc present but very incomplete and faint; R1 virtually straight; R2+3 parallels R1, curved costad only at very apex; fork of R4 and R5 widely divergent, nearly at right angle; crossveins bm-cu slightly oblique to r-m (not parallel) and proximal to r-m by distance equal to length of r-m; cell dm and crossvein dm-cu present, cell dm large; vein M2 complete, closing cell dm with crossvein dm-cu; CuA1 complete; CuA2 present, complete, forming cell cu
<emphasis id="9DDDEAA95225FFBDFF006EBF3BCEFEB9" box="[181,196,232,257]" italics="true" pageId="18" pageNumber="52">p</emphasis>
, with incomplete “spur” of veins A1[+CuA2?] originating from cell cu
<emphasis id="9DDDEAA95225FFBDFBE46EBF3F6AFEB9" box="[1105,1120,232,257]" italics="true" pageId="18" pageNumber="52">p</emphasis>
. Anal lobe of wing well developed. Legs long, bristly, with row of stiff setae on ventral margin of hind tibia. ABDOMEN: Very folded and not visible in dorsal view, unclear in lateral view.
</paragraph>
</subSubSection>
<caption id="FBD666335226FFBEFF2268893E6CF8A0" httpUri="https://zenodo.org/record/187277/files/figure.png" pageId="17" pageNumber="51" targetBox="[276,1296,203,1729]" targetPageId="17">
<paragraph id="AF1636BB5226FFBEFF2268893E6CF8A0" blockId="17.[151,1436,1758,1816]" pageId="17" pageNumber="51">
<emphasis id="9DDDEAA95226FFBEFF2268893A21F94E" bold="true" box="[151,299,1758,1782]" pageId="17" pageNumber="51">FIGURE 13.</emphasis>
<taxonomicName id="68A94D385226FFBEFE866888393CF94E" box="[307,566,1759,1782]" class="Insecta" family="Opetiidae" genus="Chimeromyina" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="51" phylum="Arthropoda" rank="species" species="concilia" status="gen. nov.">
<emphasis id="9DDDEAA95226FFBEFE866888393CF94E" box="[307,566,1759,1782]" italics="true" pageId="17" pageNumber="51">Chimeromyina concilia</emphasis>
</taxonomicName>
,
<emphasis id="9DDDEAA95226FFBEFDF06889399EF94E" bold="true" box="[581,660,1758,1782]" pageId="17" pageNumber="51">
<taxonomicNameLabel id="86EE57D25226FFBEFDF06889399EF94E" box="[581,660,1758,1782]" pageId="17" pageNumber="51" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="9DDDEAA95226FFBEFD16688939EBF94E" bold="true" box="[675,737,1758,1782]" pageId="17" pageNumber="51">n. sp.</emphasis>
, in Spanish amber. Holotype, MCNA 8882. A. Dorsal view of anterior half of body. B. Detail of antennae and vertex of head. C. Left habitus (terminalia lost at surface of amber).
</paragraph>
</caption>
<subSubSection id="E7B365305225FFBDFF706F0839ADFE27" pageId="18" pageNumber="52" type="materials_examined">
<paragraph id="AF1636BB5225FFBDFF706F0839ADFE27" blockId="18.[151,1437,152,532]" pageId="18" pageNumber="52">
<emphasis id="9DDDEAA95225FFBDFF706F083A09FEC1" bold="true" box="[197,259,351,377]" pageId="18" pageNumber="52">
<typeStatus id="701288195225FFBDFF706F083A09FEC1" box="[197,259,351,377]" pageId="18" pageNumber="52">Type</typeStatus>
</emphasis>
. Sex? (tip of abdomen not preserved),
<typeStatus id="701288195225FFBDFD456F083862FEC1" box="[752,872,351,377]" pageId="18" pageNumber="52" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="D7BE762B5225FFBDFCC16F0838C5FEC1" box="[884,975,351,377]" name="Spain" pageId="18" pageNumber="52">SPAIN</collectingCountry>
:
<taxonomicName id="68A94D385225FFBDFC696F083F49FEC1" box="[988,1091,351,377]" class="Insecta" family="Opetiidae" genus="Chimeromyia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="50" phylum="Arthropoda" rank="species" species="alava" status="sp. nov.">
<collectionCode id="C9B8AE7E5225FFBDFC696F083F49FEC1" box="[988,1091,351,377]" pageId="18" pageNumber="52">ÁLAVA</collectionCode>
</taxonomicName>
, near Peñacerrada, Escucha Formation, Lower Cretaceous;
<collectionCode id="C9B8AE7E5225FFBDFDB06FD23957FE27" box="[517,605,389,415]" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="18" pageNumber="52">MCNA</collectionCode>
8882.
</paragraph>
</subSubSection>
<subSubSection id="E7B365305225FFBDFF706FFB3FA4FE55" pageId="18" pageNumber="52" type="etymology">
<paragraph id="AF1636BB5225FFBDFF706FFB3FA4FE55" blockId="18.[151,1437,152,532]" pageId="18" pageNumber="52">
<emphasis id="9DDDEAA95225FFBDFF706FFB3A5BFE7E" bold="true" box="[197,337,428,454]" pageId="18" pageNumber="52">Etymology</emphasis>
. From the Latin,
<emphasis id="9DDDEAA95225FFBDFD986FFB399BFE7D" box="[557,657,428,453]" italics="true" pageId="18" pageNumber="52">concilio</emphasis>
, unite separate parts into a whole, connect; in reference to the plesiomorphic features of this genus that link
<emphasis id="9DDDEAA95225FFBDFD1A6F843847FE54" box="[687,845,467,492]" italics="true" pageId="18" pageNumber="52">Chimeromyia</emphasis>
with other basal Eremoneura.
</paragraph>
</subSubSection>
<subSubSection id="E7B365305225FFBDFF706FAE38B4FDAC" box="[197,958,505,532]" pageId="18" pageNumber="52" type="discussion">
<paragraph id="AF1636BB5225FFBDFF706FAE38B4FDAC" blockId="18.[151,1437,152,532]" box="[197,958,505,532]" pageId="18" pageNumber="52">
<emphasis id="9DDDEAA95225FFBDFF706FAE3A59FDAB" bold="true" box="[197,339,505,531]" pageId="18" pageNumber="52">Comments.</emphasis>
See discussion of phylogenetic relationships, below.
</paragraph>
</subSubSection>
</treatment>
</document>
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