treatments-xml/data/1D/57/B2/1D57B212FFA8FFF07EB9FC581183FEEC.xml
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<document id="4623DB9B6E34B556F488DED09BEB063E" ID-CLB-Dataset="9025" ID-DOI="10.11646/zootaxa.4861.4.6" ID-GBIF-Dataset="59dc47c8-01b5-4f14-baa6-95b57158ca96" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416789" ID-ZooBank="1A9329DD-74BA-4B44-9008-383D4C88C9D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609801562384" checkinUser="plazi" docAuthor="Ma, Zijun, Huang, Sunbin &amp; Tian, Mingyi" docDate="2020" docId="1D57B212FFA8FFF07EB9FC581183FEEC" docLanguage="en" docName="zootaxa.4861.4.6.pdf" docOrigin="Zootaxa 4861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Giraffaphaenops brevicephalus Ma &amp; Huang &amp; Tian 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="591" masterDocId="E16ECA6AFFAEFFFA7E2EFFAD151DFF41" masterDocTitle="Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="593" masterPageNumber="581" pageNumber="587" updateTime="1698852245223" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="89BCAFE940005B460461C44A95199BA1">Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="32FC1580F0447333DB943BBE3ADCA76E">Ma, Zijun</mods:namePart>
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<mods:affiliation id="70B78E2740137DAF2A9B839856F19C9F">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; 3275124967 @ qq. com; https: // orcid. org / 0000 - 0001 - 8884 - 3376</mods:affiliation>
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<mods:namePart id="F3A7A90316B12C9C62B2349E15F40D31">Huang, Sunbin</mods:namePart>
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<mods:affiliation id="3DCD7CC82109986143279E8A2F09AEB5">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; Mécanismes adaptatifs et évolution (MECADEV), UMR 7179 CNRS-MNHN, Muséum National dHistoire Naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France. huangsunbin @ 163. com; https: // orcid. org / 0000 - 0001 - 8357 - 6651</mods:affiliation>
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<mods:namePart id="3A2145C04F2FEA3FB3B7575A128BE39C">Tian, Mingyi</mods:namePart>
<mods:affiliation id="EEC0935757297E4B688E23769311E180">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA</mods:affiliation>
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<mods:date id="5D40B4EF98084A7ECA5D6628ECF5288A">2020</mods:date>
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<treatment id="1D57B212FFA8FFF07EB9FC581183FEEC" ID-DOI="http://doi.org/10.5281/zenodo.4416961" ID-GBIF-Taxon="176440759" ID-Zenodo-Dep="4416961" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC" httpUri="http://treatment.plazi.org/id/1D57B212FFA8FFF07EB9FC581183FEEC" lastPageId="10" lastPageNumber="591" pageId="6" pageNumber="587">
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<heading id="CE09B468FFA8FFFC7EB9FC581740FB4E" bold="true" box="[151,605,1013,1040]" fontSize="11" level="1" pageId="6" pageNumber="587" reason="1">
<emphasis id="A78ADF16FFA8FFFC7EB9FC581740FB4E" bold="true" box="[151,605,1013,1040]" pageId="6" pageNumber="587">
<taxonomicName id="52FE7887FFA8FFFC7EB9FC58170EFB51" ID-CoL="8MGW5" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[151,531,1013,1040]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA8FFFC7EB9FC58170EFB51" bold="true" box="[151,531,1013,1040]" italics="true" pageId="6" pageNumber="587">Giraffaphaenops brevicephalus</emphasis>
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<taxonomicNameLabel id="BCB9626DFFA8FFFC7C35FC581740FB4E" box="[539,605,1013,1039]" pageId="6" pageNumber="587" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="0DC51F81FFA8FFFC7EB1FBB41405FB72" box="[159,280,1049,1075]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStart-5="FIGURE 12" captionStartId-0="7.[151,250,1107,1131]" captionStartId-1="7.[151,250,2009,2033]" captionStartId-2="8.[151,250,1974,1998]" captionStartId-3="9.[151,250,1962,1986]" captionStartId-4="10.[151,250,1385,1409]" captionStartId-5="11.[151,250,1967,1991]" captionTargetBox-1="[435,1158,1165,1984]" captionTargetBox-2="[198,1383,567,1927]" captionTargetBox-3="[189,1396,526,1916]" captionTargetBox-4="[157,1430,454,1360]" captionTargetBox-5="[171,1417,181,1943]" captionTargetId-0="figure-20@7.[592,996,181,1083]" captionTargetId-1="figure-31@7.[428,1160,1155,1985]" captionTargetId-2="figure-194@8.[192,1396,561,1951]" captionTargetId-3="figure-185@9.[165,1421,489,1938]" captionTargetId-4="figure-156@10.[151,1436,453,1361]" captionTargetId-5="figure-20@11.[171,1417,181,1943]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="FIGURE 7. Habitus of Giraffaphaenops brevicephalus n. sp." captionText-1="FIGURE 8. Head of Giraffaphaenops species, ventral view (A. G. brevicephalus n. sp., paratype, female; B. G. yangi, male)." captionText-2="FIGURE 9. Elytra chaetotaxy of Giraffaphaenops species (A. G. brevicephalus n. sp., male, holotype; B. G. yangi, male; C. G. clarkei, male)." captionText-3="FIGURE 10. Male genitalia of Giraffaphaenops brevicephalus n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." captionText-4="FIGURE 11. Distribution map of the genus Giraffaphaenops." captionText-5="FIGURE 12. Cave Dalongguan Dong, type locality of Giraffaphaenops brevicephalus n. sp. (A. entrance; B. cave environment; C. Giraffaphaenops brevicephalus n. sp.)." figureDoi-0="http://doi.org/10.5281/zenodo.4426436" figureDoi-1="http://doi.org/10.5281/zenodo.4416799" figureDoi-2="http://doi.org/10.5281/zenodo.4416801" figureDoi-3="http://doi.org/10.5281/zenodo.4416803" figureDoi-4="http://doi.org/10.5281/zenodo.4416805" figureDoi-5="http://doi.org/10.5281/zenodo.4416807" httpUri-0="https://zenodo.org/record/4426436/files/figure.png" httpUri-1="https://zenodo.org/record/4416799/files/figure.png" httpUri-2="https://zenodo.org/record/4416801/files/figure.png" httpUri-3="https://zenodo.org/record/4416803/files/figure.png" httpUri-4="https://zenodo.org/record/4416805/files/figure.png" httpUri-5="https://zenodo.org/record/4416807/files/figure.png" pageId="6" pageNumber="587">Figs. 712</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDE4508FFFA8FFFC7EB9FBCC1154FBA6" pageId="6" pageNumber="587" type="materials_examined">
<paragraph id="95410304FFA8FFFC7EB9FBCC1154FBA6" blockId="6.[151,1437,1121,1976]" pageId="6" pageNumber="587">
<emphasis id="A78ADF16FFA8FFFC7EB9FBCC141BFB3A" bold="true" box="[151,262,1121,1147]" pageId="6" pageNumber="587">Material.</emphasis>
<materialsCitation id="25960959FFA8FFFC7F23FBCC175DFB82" ID-GBIF-Occurrence="3012566303" collectingDate="2020-05-15" collectionCode="SCAU" collectorName="Chenghai Fu" country="China" county="Autonomous Region" elevation="1238" latitude="24.573185" location="cave Dalongguan Dong" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" pageId="6" pageNumber="587" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="4A45BDA6FFA8FFFC7F23FBCC1468FB3A" box="[269,373,1121,1147]" pageId="6" pageNumber="587" type="holotype">Holotype</typeStatus>
male,
<location id="902155DFFFA8FFFC7FEFFBCF17D6FB3D" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC:902155DFFFA8FFFC7FEFFBCF17D6FB3D" box="[449,715,1121,1148]" country="China" county="Autonomous Region" latitude="24.573185" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" name="cave Dalongguan Dong" pageId="6" pageNumber="587" stateProvince="Guangxi">cave Dalongguan Dong</location>
,
<location id="902155DFFFA8FFFC7CF9FBCC1688FB3D" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC:902155DFFFA8FFFC7CF9FBCC1688FB3D" box="[727,917,1121,1148]" country="China" county="Autonomous Region" latitude="24.573185" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" name="Dalongguan Tun" pageId="6" pageNumber="587" stateProvince="Guangxi">Dalongguan Tun</location>
,
<location id="902155DFFFA8FFFC7D8EFBCC1135FB3A" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC:902155DFFFA8FFFC7D8EFBCC1135FB3A" box="[928,1064,1121,1147]" country="China" county="Autonomous Region" latitude="24.573185" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" name="Jialong Cun" pageId="6" pageNumber="587" stateProvince="Guangxi">Jialong Cun</location>
,
<location id="902155DFFFA8FFFC7A1AFBCC11CDFB3A" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC:902155DFFFA8FFFC7A1AFBCC11CDFB3A" box="[1076,1232,1121,1147]" country="China" county="Autonomous Region" latitude="24.573185" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" name="Gantian Zhen" pageId="6" pageNumber="587" stateProvince="Guangxi">Gantian Zhen</location>
,
<location id="902155DFFFA8FFFC7AF5FBCF104EFB3A" LSID="urn:lsid:plazi:treatment:1D57B212FFA8FFF07EB9FC581183FEEC:902155DFFFA8FFFC7AF5FBCF104EFB3A" box="[1243,1363,1121,1148]" country="China" county="Autonomous Region" latitude="24.573185" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" name="Leye Xian" pageId="6" pageNumber="587" stateProvince="Guangxi">Leye Xian</location>
,
<collectingMunicipality id="7525997EFFA8FFFC7B70FBCC15A2FBDE" pageId="6" pageNumber="587">Baise Shi</collectingMunicipality>
,
<collectingRegion id="573ACDE6FFA8FFFC7EE4FB28149AFBDE" box="[202,391,1157,1183]" country="China" name="Guangxi" pageId="6" pageNumber="587">Guangxi Zhuang</collectingRegion>
<collectingCounty id="7C207B88FFA8FFFC7FA5FB2B176BFBDE" box="[395,630,1157,1184]" pageId="6" pageNumber="587">Autonomous Region</collectingCounty>
,
<collectingCountry id="EDE94394FFA8FFFC7CAFFB2817DAFBDE" box="[641,711,1157,1183]" name="China" pageId="6" pageNumber="587">China</collectingCountry>
,
<geoCoordinate id="F0CA65C3FFA8FFFC7CFFFB281666FBDE" box="[721,891,1157,1183]" degrees="24" direction="north" minutes="34" orientation="latitude" pageId="6" pageNumber="587" precision="1" seconds="23.46" value="24.573185">24°34'23.46&quot;N</geoCoordinate>
,
<geoCoordinate id="F0CA65C3FFA8FFFC7DABFB281127FBE1" box="[901,1082,1157,1184]" degrees="106" direction="east" minutes="25" orientation="longitude" pageId="6" pageNumber="587" precision="1" seconds="27.96" value="106.42443">106°25'27.96&quot;E</geoCoordinate>
,
<quantity id="5206AEE1FFA8FFFC7A6BFB281184FBE1" box="[1093,1177,1157,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.238" pageId="6" pageNumber="587" unit="m" value="1238.0">
<elevation id="1ED3E437FFA8FFFC7A6BFB281184FBE1" box="[1093,1177,1157,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.238" pageId="6" pageNumber="587" unit="m" value="1238.0">1238 m</elevation>
</quantity>
in altitude,
<date id="E14025C4FFA8FFFC7B33FB281085FBE1" box="[1309,1432,1157,1184]" pageId="6" pageNumber="587" value="2020-05-15">
<collectingDate id="F104DC2CFFA8FFFC7B33FB281085FBE1" box="[1309,1432,1157,1184]" pageId="6" pageNumber="587" value="2020-05-15">15-V-2020</collectingDate>
</date>
, leg.
<collectorName id="380B66D2FFA8FFFC7EE9FB041444FB85" box="[199,345,1193,1220]" pageId="6" pageNumber="587">Chenghai Fu</collectorName>
, deposited in
<collectionCode id="F3EF9BC1FFA8FFFC7FDCFB04175DFB82" box="[498,576,1193,1219]" pageId="6" pageNumber="587">SCAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="25960959FFA8FFFC7C65FB07109EFB82" ID-GBIF-Occurrence="3012566304" box="[587,1411,1193,1220]" collectingDate="2020-05-15" collectionCode="SCAU" collectorName="Chenghai Fu" country="China" county="Autonomous Region" elevation="1238" latitude="24.573185" location="cave Dalongguan Dong" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" pageId="6" pageNumber="587" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus id="4A45BDA6FFA8FFFC7C65FB0717A1FB85" box="[587,700,1194,1220]" pageId="6" pageNumber="587" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="83F8C88DFFA8FFFC7CE8FB04160CFB82" box="[710,785,1193,1219]" pageId="6" pageNumber="587" type="male">1 male</specimenCount>
&amp;
<specimenCount id="83F8C88DFFA8FFFC7D1AFB041684FB82" box="[820,921,1193,1219]" pageId="6" pageNumber="587" type="female">1 female</specimenCount>
; same data as holotype; deposited in
<collectionCode id="F3EF9BC1FFA8FFFC7B1BFB04109EFB82" box="[1333,1411,1193,1219]" pageId="6" pageNumber="587">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="25960959FFA8FFFC7BA0FB041158FBA6" ID-GBIF-Occurrence="3012566305" collectingDate="2020-06-29" collectionCode="SCAU" collectorName="Mingyi Tian" country="China" county="Autonomous Region" elevation="1238" latitude="24.573185" location="cave Dalongguan Dong" longLatPrecision="1" longitude="106.42443" municipality="Baise Shi" pageId="6" pageNumber="587" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="83F8C88DFFA8FFFC7BA0FB0415D0FBA6" pageId="6" pageNumber="587" type="male">1 male</specimenCount>
&amp;
<specimenCount id="83F8C88DFFA8FFFC7EDFFB60147DFBA6" box="[241,352,1229,1255]" pageId="6" pageNumber="587" type="female">2 females</specimenCount>
, same locality,
<date id="E14025C4FFA8FFFC7C22FB60178EFBA6" box="[524,659,1229,1255]" pageId="6" pageNumber="587" value="2020-06-29">
<collectingDate id="F104DC2CFFA8FFFC7C22FB60178EFBA6" box="[524,659,1229,1255]" pageId="6" pageNumber="587" value="2020-06-29">29-VI-2020</collectingDate>
</date>
, leg.
<collectorName id="380B66D2FFA8FFFC7CE1FB601641FBA6" box="[719,860,1229,1255]" pageId="6" pageNumber="587">Mingyi Tian</collectorName>
, deposited in
<collectionCode id="F3EF9BC1FFA8FFFC7DD9FB601158FBA6" box="[1015,1093,1229,1255]" pageId="6" pageNumber="587">SCAU</collectionCode>
</materialsCitation>
.
</paragraph>
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<subSubSection id="DDE4508FFFA8FFF37EE9FB5C105BFE90" lastPageId="9" lastPageNumber="590" pageId="6" pageNumber="587" type="description">
<paragraph id="95410304FFA8FFFC7EE9FB5C1089FA71" blockId="6.[151,1437,1121,1976]" pageId="6" pageNumber="587">
<emphasis id="A78ADF16FFA8FFFC7EE9FB5C1444FA4A" bold="true" box="[199,345,1265,1291]" pageId="6" pageNumber="587">Description.</emphasis>
Length:
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including mandibles, or
<quantity id="5206AEE1FFA8FFFC7D54FB5C1112FA4D" box="[890,1039,1265,1292]" metricMagnitude="-3" metricUnit="m" metricValue="8.17" metricValueMax="8.4" metricValueMin="7.94" pageId="6" pageNumber="587" unit="mm" value="8.17" valueMax="8.4" valueMin="7.94">7.948.4 mm</quantity>
excluding mandibles; width:
<quantity id="5206AEE1FFA8FFFC7B73FB5C15E3FA71" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.69" metricValueMin="1.33" pageId="6" pageNumber="587" unit="mm" value="1.51" valueMax="1.69" valueMin="1.33">1.33 1.69 mm</quantity>
; fore body, head (excluding mandibles) plus pronotum, distinctly longer than elytra. Habitus as in
<figureCitation id="0DC51F81FFA8FFFC7B62FAB8108CFA71" box="[1356,1425,1301,1328]" captionStart="FIGURE 7" captionStartId="7.[151,250,1107,1131]" captionTargetId="figure-20@7.[592,996,181,1083]" captionTargetPageId="7" captionText="FIGURE 7. Habitus of Giraffaphaenops brevicephalus n. sp." figureDoi="http://doi.org/10.5281/zenodo.4426436" httpUri="https://zenodo.org/record/4426436/files/figure.png" pageId="6" pageNumber="587">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="95410304FFA8FFFC7EE9FA941469FADA" blockId="6.[151,1437,1121,1976]" pageId="6" pageNumber="587">Body yellowish brown to reddish brown, elytra slightly lighter, palps, antennae and tarsi pale; shiny, wholly pubescent including appendages, covered with short and erected setae, except propleura, mesosternum and epipleura, which are glabrous.</paragraph>
<paragraph id="95410304FFA8FFFC7EE9FA0811CCF8F9" blockId="6.[151,1437,1121,1976]" pageId="6" pageNumber="587">
Head elongate, nearly triangular in form, widest at 2/5 from the base (including mandibles), a little behind the level of antennal articulation, temples slightly and gradually narrowed posteriad towards the distinct neck constriction, base semicolumn-shaped, constriction about 2/5 as wide as head; head excluding mandibles distinctly shorter than pronotum, but as long as pronotum including mandibles; frontal furrows short but distinct, nearly parallel-sided though slightly divergent posteriorly; only anterior pair of supraorbital pores present, at about the widest level; a pair of suborbital pores present, close to the base of head; clypeus transverse, 6-setose; labrum transverse, distinctly protruded medially, 6-setose; mandibles thin and elongated, right one faintly bidentate; palps thin and very elongated, penultimate palpomeres much longer than the apical ones (1.3 times); the 2
<superScript id="628BAE4CFFA8FFFC7A11F90C1152F9EE" attach="left" box="[1087,1103,1697,1711]" fontSize="6" pageId="6" pageNumber="587">nd</superScript>
labial palpomere bisetose on inner margin, with two additional short setae on outer margin, one at middle and the other at subapex; mentum and submentum completely fused; mentum widely concave basally, mental tooth bifid, with two setae on each side; submentum 13-setose (
<figureCitation id="0DC51F81FFA8FFFC7F5DF8A014D0F866" box="[371,461,1805,1831]" captionStart="FIGURE 8" captionStartId="7.[151,250,2009,2033]" captionTargetBox="[435,1158,1165,1984]" captionTargetId="figure-31@7.[428,1160,1155,1985]" captionTargetPageId="7" captionText="FIGURE 8. Head of Giraffaphaenops species, ventral view (A. G. brevicephalus n. sp., paratype, female; B. G. yangi, male)." figureDoi="http://doi.org/10.5281/zenodo.4416799" httpUri="https://zenodo.org/record/4416799/files/figure.png" pageId="6" pageNumber="587">Fig. 8A</figureCitation>
); antennae filiform, extraordinary long and slender, 10
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and 11
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antennomeres exceeding over elytral apices; 1
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antennomere stout, 2.7 times as long as wide; 1
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subequal to 2
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in length, about a third as long as 3
<superScript id="628BAE4CFFA8FFFC7ED3F8F81416F822" attach="left" box="[253,267,1877,1891]" fontSize="6" pageId="6" pageNumber="587">rd</superScript>
which is subequal in length to 4
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and 5
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respectively; length of antennomeres gradually shortened from 6
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to 10
<superScript id="628BAE4CFFA8FFFC7EDDF8D4141DF8C6" attach="none" box="[243,256,1913,1927]" fontSize="6" pageId="6" pageNumber="587">th</superScript>
, and 7
<superScript id="628BAE4CFFA8FFFC7F62F8D41444F8C6" attach="left" box="[332,345,1913,1927]" fontSize="6" pageId="6" pageNumber="587">th</superScript>
as long as the 11
<superScript id="628BAE4CFFA8FFFC7C32F8D41734F8C6" attach="none" box="[540,553,1913,1927]" fontSize="6" pageId="6" pageNumber="587">th</superScript>
. The comparative length ratio of antennomere as follows: 1
<superScript id="628BAE4CFFA8FFFC7AE4F8D411C8F8C6" attach="left" box="[1226,1237,1913,1927]" fontSize="6" pageId="6" pageNumber="587">st</superScript>
(1.00), 2
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(1.17), 3
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(3.17), 4
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(3.25), 5
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(3.08), 6
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(2.92), 7
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(2.67), 8
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(2.25), 9
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(2.08), 10
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(1.83) and 11
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(2.58).
</paragraph>
<caption id="C181538CFFA9FFFD7EB9FBFE1614FB2A" ID-DOI="http://doi.org/10.5281/zenodo.4426436" ID-Zenodo-Dep="4426436" box="[151,777,1107,1131]" httpUri="https://zenodo.org/record/4426436/files/figure.png" pageId="7" pageNumber="588" targetBox="[600,987,190,1077]" targetPageId="7">
<paragraph id="95410304FFA9FFFD7EB9FBFE1614FB2A" blockId="7.[151,777,1107,1131]" box="[151,777,1107,1131]" pageId="7" pageNumber="588">
<emphasis id="A78ADF16FFA9FFFD7EB9FBFE140EFB2A" bold="true" box="[151,275,1107,1131]" pageId="7" pageNumber="588">FIGURE 7.</emphasis>
Habitus of
<taxonomicName id="52FE7887FFA9FFFD7FA4FBFE17D4FB2A" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[394,713,1107,1131]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA9FFFD7FA4FBFE17D4FB2A" box="[394,713,1107,1131]" italics="true" pageId="7" pageNumber="588">Giraffaphaenops brevicephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCB9626DFFA9FFFD7CE1FBFE1614FB2A" box="[719,777,1107,1131]" pageId="7" pageNumber="588" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="C181538CFFA9FFFD7EB9F8741088F8B0" ID-DOI="http://doi.org/10.5281/zenodo.4416799" ID-Zenodo-Dep="4416799" box="[151,1429,2009,2034]" httpUri="https://zenodo.org/record/4416799/files/figure.png" pageId="7" pageNumber="588" startId="7.[151,250,2009,2033]" targetBox="[435,1158,1165,1984]" targetPageId="7">
<paragraph id="95410304FFA9FFFD7EB9F8741088F8B0" blockId="7.[151,1429,2009,2034]" box="[151,1429,2009,2034]" pageId="7" pageNumber="588">
<emphasis id="A78ADF16FFA9FFFD7EB9F874140EF8B0" bold="true" box="[151,275,2009,2033]" pageId="7" pageNumber="588">FIGURE 8.</emphasis>
Head of
<taxonomicName id="52FE7887FFA9FFFD7F5FF8741700F8B0" authorityName="Deuve" authorityYear="2002" box="[369,541,2009,2033]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA9FFFD7F5FF8741700F8B0" box="[369,541,2009,2033]" italics="true" pageId="7" pageNumber="588">Giraffaphaenops</emphasis>
</taxonomicName>
species, ventral view (A.
<taxonomicName id="52FE7887FFA9FFFD7D0AF87716CDF8B0" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[804,976,2009,2033]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA9FFFD7D0AF87716CDF8B0" box="[804,976,2009,2033]" italics="true" pageId="7" pageNumber="588">G. brevicephalus</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFA9FFFD7DF9F874110DF8B0" bold="true" box="[983,1040,2009,2033]" pageId="7" pageNumber="588">
<taxonomicNameLabel id="BCB9626DFFA9FFFD7DF9F874110DF8B0" box="[983,1040,2009,2033]" pageId="7" pageNumber="588" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, female; B.
<emphasis id="A78ADF16FFA9FFFD7ADEF8771053F8B0" box="[1264,1358,2010,2033]" italics="true" pageId="7" pageNumber="588">
<taxonomicName id="52FE7887FFA9FFFD7ADEF8771057F8B0" authorityName="Tian &amp; Luo" authorityYear="2015" box="[1264,1354,2010,2033]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="yangi">G. yangi</taxonomicName>
,
</emphasis>
male).
</paragraph>
</caption>
<paragraph id="95410304FFA6FFF27EE9FF3A16E3FE00" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="589">Prothorax extremely elongate, nearly as long as head including mandibles, PrL/HL=0.911.02; fore half rodlike prolonged, evenly widened, hind half form oval-like; propleura strongly tumid, making widest at about basal fourth; pronotum narrow and markedly elongate, nearly parallel-sided throughout, basal part distinctly widened; base much wider than front, PbW/PfW=1.391.45; pronotum much narrower than prothorax, PtW/PnW=1.331.48, and narrower than head, HW/PnW=1.231.33, without lateral-marginal setae.</paragraph>
<paragraph id="95410304FFA6FFF27EE9FEE61750FD58" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="589">
Elytra similar to
<taxonomicName id="52FE7887FFA6FFF27FADFEE6179AFE24" authority="Deuve, 2002" authorityName="Deuve" authorityYear="2002" box="[387,647,331,357]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="clarkei">
<emphasis id="A78ADF16FFA6FFF27FADFEE614EFFE24" box="[387,498,331,357]" italics="true" pageId="8" pageNumber="589">G. clarkei</emphasis>
<bibRefCitation id="F16F7EF5FFA6FFF27FD7FEE6179AFE24" author="Deuve, T." box="[505,647,331,357]" pageId="8" pageNumber="589" pagination="515 - 523" refId="ref4798" refString="Deuve, T. (2002) Deux remarquables Trechinae anophtalmes des cavites souterraines du Guangxi nord-occidental, Chine (Coleoptera, Trechidae). Bulletin de la Societe entomologique de France, 107 (5), 515 - 523." type="journal article" year="2002">Deuve, 2002</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="52FE7887FFA6FFF27C93FEE616F7FE24" authority="Tian &amp; Luo, 2015" authorityName="Tian &amp; Luo" authorityYear="2015" box="[701,1002,330,357]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA6FFF27C93FEE61601FE25" box="[701,796,331,357]" italics="true" pageId="8" pageNumber="589">G. yangi</emphasis>
<bibRefCitation id="F16F7EF5FFA6FFF27D0DFEE616F7FE24" author="Tian, M. Y. &amp; Luo, X. Z." box="[803,1002,330,357]" pageId="8" pageNumber="589" pagination="581 - 588" refId="ref4840" refString="Tian, M. Y. &amp; Luo, X. Z. (2015) A new species of the highly modified hypogean genus Giraffaphaenops Deuve, 2002 (Coleoptera: Carabidae: Trechinae). Zootaxa, 3911 (4), 581 - 588. https: // doi. org / 10.11646 / zootaxa. 3911.4.7" type="journal article" year="2015">Tian &amp; Luo, 2015</bibRefCitation>
</taxonomicName>
(
<figureCitation id="0DC51F81FFA6FFF27DD7FEE61121FE24" box="[1017,1084,331,357]" captionStart="FIGURE 9" captionStartId="8.[151,250,1974,1998]" captionTargetBox="[198,1383,567,1927]" captionTargetId="figure-194@8.[192,1396,561,1951]" captionTargetPageId="8" captionText="FIGURE 9. Elytra chaetotaxy of Giraffaphaenops species (A. G. brevicephalus n. sp., male, holotype; B. G. yangi, male; C. G. clarkei, male)." figureDoi="http://doi.org/10.5281/zenodo.4416801" httpUri="https://zenodo.org/record/4416801/files/figure.png" pageId="8" pageNumber="589">Fig. 9</figureCitation>
) elongate-ovate, about twice as wide as prothorax, much longer than pronotum, EW/PnW=2.213.07; widest at about 3/5 from base, strongly convex, but basal part nearly flat, shoulders not well marked; discal setiferous pores absent, preapical pore present, seta short and indistinct; chaetotaxal pattern of the marginal umbilicate pores similar to that of
<taxonomicName id="52FE7887FFA6FFF27AA0FE1A11E3FE90" authorityName="Deuve" authorityYear="2002" box="[1166,1278,439,465]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="clarkei">
<emphasis id="A78ADF16FFA6FFF27AA0FE1A11BFFE90" box="[1166,1186,439,465]" italics="true" pageId="8" pageNumber="589">G</emphasis>
.
<emphasis id="A78ADF16FFA6FFF27A9EFE1A11E3FE90" box="[1200,1278,439,465]" italics="true" pageId="8" pageNumber="589">clarkei</emphasis>
</taxonomicName>
and
<taxonomicName id="52FE7887FFA6FFF27B1BFE1A1088FE91" authorityName="Tian &amp; Luo" authorityYear="2015" box="[1333,1429,439,465]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA6FFF27B1BFE1A1054FE90" box="[1333,1353,439,465]" italics="true" pageId="8" pageNumber="589">G</emphasis>
.
<emphasis id="A78ADF16FFA6FFF27B79FE1A1088FE91" box="[1367,1429,439,464]" italics="true" pageId="8" pageNumber="589">yangi</emphasis>
</taxonomicName>
, but the 2
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pore obviously more closer to 3
<superScript id="628BAE4CFFA6FFF27CADFE77178CFEA9" attach="left" box="[643,657,474,488]" fontSize="6" pageId="8" pageNumber="589">rd</superScript>
than to 1
<superScript id="628BAE4CFFA6FFF27D2EFE761616FEA8" attach="none" box="[768,779,475,489]" fontSize="6" pageId="8" pageNumber="589">st</superScript>
, preapical pore 1.5 times closer to suture than to margin; scutellum small, scutellar pore present.
</paragraph>
<caption id="C181538CFFA6FFF27EB9F81B1457F8B3" ID-DOI="http://doi.org/10.5281/zenodo.4416801" ID-Zenodo-Dep="4416801" httpUri="https://zenodo.org/record/4416801/files/figure.png" pageId="8" pageNumber="589" startId="8.[151,250,1974,1998]" targetBox="[198,1383,567,1927]" targetPageId="8">
<paragraph id="95410304FFA6FFF27EB9F81B1457F8B3" blockId="8.[151,1437,1974,2034]" pageId="8" pageNumber="589">
<emphasis id="A78ADF16FFA6FFF27EB9F81B1409F88F" bold="true" box="[151,276,1974,1998]" pageId="8" pageNumber="589">FIGURE 9.</emphasis>
Elytra chaetotaxy of
<taxonomicName id="52FE7887FFA6FFF27FDCF81B1783F88F" authorityName="Deuve" authorityYear="2002" box="[498,670,1974,1998]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFA6FFF27FDCF81B1783F88F" box="[498,670,1974,1998]" italics="true" pageId="8" pageNumber="589">Giraffaphaenops</emphasis>
</taxonomicName>
species (A.
<taxonomicName id="52FE7887FFA6FFF27D30F81B16D6F88F" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[798,971,1974,1998]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA6FFF27D30F81B16D6F88F" box="[798,971,1974,1998]" italics="true" pageId="8" pageNumber="589">G. brevicephalus</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFA6FFF27DFDF81B1110F88F" bold="true" box="[979,1037,1974,1998]" pageId="8" pageNumber="589">
<taxonomicNameLabel id="BCB9626DFFA6FFF27DFDF81B1110F88F" box="[979,1037,1974,1998]" pageId="8" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, holotype; B.
<taxonomicName id="52FE7887FFA6FFF27AF1F81B102AF88F" authorityName="Tian &amp; Luo" authorityYear="2015" box="[1247,1335,1974,1998]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA6FFF27AF1F81B11ECF88C" box="[1247,1265,1974,1997]" italics="true" pageId="8" pageNumber="589">G</emphasis>
.
<emphasis id="A78ADF16FFA6FFF27AD1F81A102AF88F" box="[1279,1335,1975,1998]" italics="true" pageId="8" pageNumber="589">yangi</emphasis>
</taxonomicName>
, male; C.
<taxonomicName id="52FE7887FFA6FFF27EB9F87715E1F8B3" authorityName="Deuve" authorityYear="2002" box="[151,252,2010,2034]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="clarkei">
<emphasis id="A78ADF16FFA6FFF27EB9F87715B4F8B0" box="[151,169,2010,2033]" italics="true" pageId="8" pageNumber="589">G</emphasis>
.
<emphasis id="A78ADF16FFA6FFF27E98F87715E1F8B3" box="[182,252,2010,2034]" italics="true" pageId="8" pageNumber="589">clarkei</emphasis>
</taxonomicName>
, male).
</paragraph>
</caption>
<paragraph id="95410304FFA7FFF37EE9FF3A104EFF94" blockId="9.[151,1436,151,465]" pageId="9" pageNumber="590">
Legs slender and long, hind femora shorter than elytra; fore tarsomeres not modified in male; the 1
<superScript id="628BAE4CFFA7FFF37B32FF3A103AFFE4" attach="left" box="[1308,1319,151,165]" fontSize="6" pageId="9" pageNumber="590">st</superScript>
tarsomere almost as long as 2
<superScript id="628BAE4CFFA7FFF37F42FF171491FF94" attach="none" box="[364,396,186,213]" fontSize="6" pageId="9" pageNumber="590">nd</superScript>
4
<superScript id="628BAE4CFFA7FFF37FB7FF1714BBFF89" attach="left" box="[409,422,186,200]" fontSize="6" pageId="9" pageNumber="590">th</superScript>
combined in fore legs, while distinctly longer in middle and hind legs, respectively.
</paragraph>
<paragraph id="95410304FFA7FFF37EE9FF721458FE5C" blockId="9.[151,1436,151,465]" pageId="9" pageNumber="590">Ventrite IVVI each with a pair of paramedian setae, VII bearing a pair of marginal setae in male, while two pairs in female.</paragraph>
<paragraph id="95410304FFA7FFF37EE9FE8A105BFE90" blockId="9.[151,1436,151,465]" pageId="9" pageNumber="590">
Male genitalia (
<figureCitation id="0DC51F81FFA7FFF37F5BFE8A14D5FE00" box="[373,456,295,321]" captionStart="FIGURE 10" captionStartId="9.[151,250,1962,1986]" captionTargetBox="[189,1396,526,1916]" captionTargetId="figure-185@9.[165,1421,489,1938]" captionTargetPageId="9" captionText="FIGURE 10. Male genitalia of Giraffaphaenops brevicephalus n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416803" httpUri="https://zenodo.org/record/4416803/files/figure.png" pageId="9" pageNumber="590">Fig. 10</figureCitation>
): median lobe of aedeagus very similar to that of
<taxonomicName id="52FE7887FFA7FFF37DC8FE8A1158FE01" authorityName="Tian &amp; Luo" authorityYear="2015" box="[998,1093,295,321]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA7FFF37DC8FE8A16E7FE00" box="[998,1018,295,321]" italics="true" pageId="9" pageNumber="590">G</emphasis>
.
<emphasis id="A78ADF16FFA7FFF37A29FE8A1158FE01" box="[1031,1093,295,320]" italics="true" pageId="9" pageNumber="590">yangi</emphasis>
</taxonomicName>
, slender and arcuate, basal orifice large, but apex slightly broader and more blunt, not distinctly constrict (
<figureCitation id="0DC51F81FFA7FFF37DCBFEE61150FE24" box="[997,1101,331,357]" captionStart="FIGURE 10" captionStartId="9.[151,250,1962,1986]" captionTargetBox="[189,1396,526,1916]" captionTargetId="figure-185@9.[165,1421,489,1938]" captionTargetPageId="9" captionText="FIGURE 10. Male genitalia of Giraffaphaenops brevicephalus n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416803" httpUri="https://zenodo.org/record/4416803/files/figure.png" pageId="9" pageNumber="590">Fig. 10A</figureCitation>
); sagittal aileron small, nearly rounded and semitransparent; inner sac weakly marked, armed with a broad copulatory piece which is about 2/7 as long as the median lobe; in dorsal view, apical lobe slenderer, rounded at apex (
<figureCitation id="0DC51F81FFA7FFF37A2EFE3E117BFEEC" box="[1024,1126,403,429]" captionStart="FIGURE 10" captionStartId="9.[151,250,1962,1986]" captionTargetBox="[189,1396,526,1916]" captionTargetId="figure-185@9.[165,1421,489,1938]" captionTargetPageId="9" captionText="FIGURE 10. Male genitalia of Giraffaphaenops brevicephalus n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416803" httpUri="https://zenodo.org/record/4416803/files/figure.png" pageId="9" pageNumber="590">Fig. 10B</figureCitation>
); parameres similar to those of
<taxonomicName id="52FE7887FFA7FFF37E98FE1A140BFE91" authorityName="Tian &amp; Luo" authorityYear="2015" box="[182,278,439,465]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA7FFF37E98FE1A15D7FE90" box="[182,202,439,465]" italics="true" pageId="9" pageNumber="590">G</emphasis>
.
<emphasis id="A78ADF16FFA7FFF37EF6FE1A140BFE91" box="[216,278,439,464]" italics="true" pageId="9" pageNumber="590">yangi</emphasis>
</taxonomicName>
, the right bearing three long setae at apex, while the left only two; genital ring wide (
<figureCitation id="0DC51F81FFA7FFF37AFCFE1A1024FE90" box="[1234,1337,439,465]" captionStart="FIGURE 10" captionStartId="9.[151,250,1962,1986]" captionTargetBox="[189,1396,526,1916]" captionTargetId="figure-185@9.[165,1421,489,1938]" captionTargetPageId="9" captionText="FIGURE 10. Male genitalia of Giraffaphaenops brevicephalus n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416803" httpUri="https://zenodo.org/record/4416803/files/figure.png" pageId="9" pageNumber="590">Fig. 10C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="C181538CFFA7FFF37EB9F807163CF8A7" ID-DOI="http://doi.org/10.5281/zenodo.4416803" ID-Zenodo-Dep="4416803" httpUri="https://zenodo.org/record/4416803/files/figure.png" pageId="9" pageNumber="590" startId="9.[151,250,1962,1986]" targetBox="[189,1396,526,1916]" targetPageId="9">
<paragraph id="95410304FFA7FFF37EB9F807163CF8A7" blockId="9.[151,1436,1962,2022]" pageId="9" pageNumber="590">
<emphasis id="A78ADF16FFA7FFF37EB9F8071402F883" bold="true" box="[151,287,1962,1986]" pageId="9" pageNumber="590">FIGURE 10.</emphasis>
Male genitalia of
<taxonomicName id="52FE7887FFA7FFF37FF8F8071609F883" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[470,788,1962,1986]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA7FFF37FF8F8071609F883" box="[470,788,1962,1986]" italics="true" pageId="9" pageNumber="590">Giraffaphaenops brevicephalus</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFA7FFF37D34F807164FF883" bold="true" box="[794,850,1962,1986]" pageId="9" pageNumber="590">
<taxonomicNameLabel id="BCB9626DFFA7FFF37D34F807164FF883" box="[794,850,1962,1986]" pageId="9" pageNumber="590" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view).
</paragraph>
</caption>
<subSubSection id="DDE4508FFFA4FFF07EE9FF3A1493FFB8" pageId="10" pageNumber="591" type="discussion">
<paragraph id="95410304FFA4FFF07EE9FF3A1493FFB8" blockId="10.[151,1437,151,429]" pageId="10" pageNumber="591">
<emphasis id="A78ADF16FFA4FFF07EE9FF3A1420FFF0" bold="true" box="[199,317,151,177]" pageId="10" pageNumber="591">Remarks:</emphasis>
<taxonomicName id="52FE7887FFA4FFF07F6BFF3A171EFFF0" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[325,515,151,177]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="591" phylum="Arthropoda" rank="species" species="brevicephalus" status="sp. nov.">
<emphasis id="A78ADF16FFA4FFF07F6BFF3A1444FFF0" box="[325,345,151,177]" italics="true" pageId="10" pageNumber="591">G</emphasis>
.
<emphasis id="A78ADF16FFA4FFF07F49FF3A171EFFF0" box="[359,515,151,177]" italics="true" pageId="10" pageNumber="591">brevicephalus</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFA4FFF07C24FF3A1754FFF0" bold="true" box="[522,585,151,177]" pageId="10" pageNumber="591">
<taxonomicNameLabel id="BCB9626DFFA4FFF07C24FF3A1754FFF0" box="[522,585,151,177]" pageId="10" pageNumber="591" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is very similar in prothorax and elytra shape to
<taxonomicName id="52FE7887FFA4FFF07A4DFF3A11CEFFF0" authorityName="Deuve" authorityYear="2002" box="[1123,1235,151,177]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="591" phylum="Arthropoda" rank="species" species="clarkei">
<emphasis id="A78ADF16FFA4FFF07A4DFF3A116AFFF0" box="[1123,1143,151,177]" italics="true" pageId="10" pageNumber="591">G</emphasis>
.
<emphasis id="A78ADF16FFA4FFF07AABFF3A11CEFFF0" box="[1157,1235,151,177]" italics="true" pageId="10" pageNumber="591">clarkei</emphasis>
</taxonomicName>
and
<taxonomicName id="52FE7887FFA4FFF07B24FF3A1077FFF1" authorityName="Tian &amp; Luo" authorityYear="2015" box="[1290,1386,151,177]" class="Insecta" family="Carabidae" genus="Giraffaphaenops" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="591" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="A78ADF16FFA4FFF07B24FF3A1003FFF0" box="[1290,1310,151,177]" italics="true" pageId="10" pageNumber="591">G</emphasis>
.
<emphasis id="A78ADF16FFA4FFF07B02FF3A1077FFF1" box="[1324,1386,151,176]" italics="true" pageId="10" pageNumber="591">yangi</emphasis>
</taxonomicName>
, but it is easily separated from them by its shorter and broader head (
<figureCitation id="0DC51F81FFA4FFF07D5EFF1616D7FF94" box="[880,970,187,213]" captionStart="FIGURE 8" captionStartId="7.[151,250,2009,2033]" captionTargetBox="[435,1158,1165,1984]" captionTargetId="figure-31@7.[428,1160,1155,1985]" captionTargetPageId="7" captionText="FIGURE 8. Head of Giraffaphaenops species, ventral view (A. G. brevicephalus n. sp., paratype, female; B. G. yangi, male)." figureDoi="http://doi.org/10.5281/zenodo.4416799" httpUri="https://zenodo.org/record/4416799/files/figure.png" pageId="10" pageNumber="591">Fig. 8A</figureCitation>
), versus longer and narrower in the latter two species (
<figureCitation id="0DC51F81FFA4FFF07F06FF72149DFFB8" box="[296,384,223,249]" captionStart="FIGURE 8" captionStartId="7.[151,250,2009,2033]" captionTargetBox="[435,1158,1165,1984]" captionTargetId="figure-31@7.[428,1160,1155,1985]" captionTargetPageId="7" captionText="FIGURE 8. Head of Giraffaphaenops species, ventral view (A. G. brevicephalus n. sp., paratype, female; B. G. yangi, male)." figureDoi="http://doi.org/10.5281/zenodo.4416799" httpUri="https://zenodo.org/record/4416799/files/figure.png" pageId="10" pageNumber="591">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDE4508FFFA4FFF07EE9FEAE115FFE5C" box="[199,1090,259,285]" pageId="10" pageNumber="591" type="etymology">
<paragraph id="95410304FFA4FFF07EE9FEAE115FFE5C" blockId="10.[151,1437,151,429]" box="[199,1090,259,285]" pageId="10" pageNumber="591">
<emphasis id="A78ADF16FFA4FFF07EE9FEAE144FFE5C" bold="true" box="[199,338,259,285]" pageId="10" pageNumber="591">Etymology:</emphasis>
“brevi-”+“cephalus”, to indicate its short head of this new species.
</paragraph>
</subSubSection>
<subSubSection id="DDE4508FFFA4FFF07EE9FE8B1183FEEC" pageId="10" pageNumber="591" type="distribution">
<paragraph id="95410304FFA4FFF07EE9FE8B1183FEEC" blockId="10.[151,1437,151,429]" pageId="10" pageNumber="591">
<emphasis id="A78ADF16FFA4FFF07EE9FE8B147EFE01" bold="true" box="[199,355,294,320]" pageId="10" pageNumber="591">Distribution:</emphasis>
<collectingCountry id="EDE94394FFA4FFF07F43FE8A14ADFE00" box="[365,432,295,321]" name="China" pageId="10" pageNumber="591">China</collectingCountry>
(
<collectingRegion id="573ACDE6FFA4FFF07FECFE8A1734FE03" box="[450,553,295,322]" country="China" name="Guangxi" pageId="10" pageNumber="591">Guangxi</collectingRegion>
). Known only from the cave Dalongguan Dong (
<figureCitation id="0DC51F81FFA4FFF07A49FE8A11A1FE00" box="[1127,1212,295,321]" captionStart="FIGURE 11" captionStartId="10.[151,250,1385,1409]" captionTargetBox="[157,1430,454,1360]" captionTargetId="figure-156@10.[151,1436,453,1361]" captionTargetPageId="10" captionText="FIGURE 11. Distribution map of the genus Giraffaphaenops." figureDoi="http://doi.org/10.5281/zenodo.4416805" httpUri="https://zenodo.org/record/4416805/files/figure.png" pageId="10" pageNumber="591">Fig. 11</figureCitation>
). This cave is very close to the village Dalongguan. With a very small and narrow entrance (
<figureCitation id="0DC51F81FFA4FFF07D91FEE6113AFE24" box="[959,1063,331,357]" captionStart="FIGURE 12" captionStartId="11.[151,250,1967,1991]" captionTargetBox="[171,1417,181,1943]" captionTargetId="figure-20@11.[171,1417,181,1943]" captionTargetPageId="11" captionText="FIGURE 12. Cave Dalongguan Dong, type locality of Giraffaphaenops brevicephalus n. sp. (A. entrance; B. cave environment; C. Giraffaphaenops brevicephalus n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4416807" httpUri="https://zenodo.org/record/4416807/files/figure.png" pageId="10" pageNumber="591">Fig. 12A</figureCitation>
), then the passage becomes much wider. It is about two hundred meters long before joining an underground river. All of the beetles were discovered and collected in a wet area about four to five dozen meters from the entrance (
<figureCitation id="0DC51F81FFA4FFF07A2CFE3E118DFEEC" box="[1026,1168,403,429]" captionStart="FIGURE 12" captionStartId="11.[151,250,1967,1991]" captionTargetBox="[171,1417,181,1943]" captionTargetId="figure-20@11.[171,1417,181,1943]" captionTargetPageId="11" captionText="FIGURE 12. Cave Dalongguan Dong, type locality of Giraffaphaenops brevicephalus n. sp. (A. entrance; B. cave environment; C. Giraffaphaenops brevicephalus n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4416807" httpUri="https://zenodo.org/record/4416807/files/figure.png" pageId="10" pageNumber="591">Fig. 12 B, C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>