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<document id="8C95744EC3E2FA0BACC335428AB1A188" ID-CLB-Dataset="24882" ID-DOI="10.11646/zootaxa.4698.1.1" ID-GBIF-Dataset="1c94057f-64de-4f19-b047-2223e4d664e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3542876" ID-ZooBank="0B4482B9-F89F-45CA-B929-E9B098605CD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573811688707" checkinUser="plazi" docAuthor="Calvanese, Victor C., Brescovit, Antonio D. &amp; Bonato, Lucio" docDate="2019" docId="06130E72A523FFCAFF34FBFEFDD2AE9E" docLanguage="en" docName="zootaxa.4698.1.1.pdf" docOrigin="Zootaxa 4698 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mecophilus tupiniquim Calvanese &amp; Brescovit &amp; Bonato 2019, new species" docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="FA2A760AA504FFE2FFA3FFDCFFCBAC35" masterDocTitle="Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily" masterLastPageNumber="72" masterPageNumber="1" pageNumber="40" updateTime="1698767377376" updateUser="ExternalLinkService">
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<mods:title id="6854402B6DB5B2E903302313F2B9DB71">Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily</mods:title>
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<mods:namePart id="A26AA9EF95A80362377363DD57829DA2">Calvanese, Victor C.</mods:namePart>
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<mods:namePart id="B1D27F7B0D8FA97A68DA808A2F04ACDA">Brescovit, Antonio D.</mods:namePart>
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<mods:namePart id="06C78D61161F88018E243E369C4AD2C2">Bonato, Lucio</mods:namePart>
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<mods:title id="7531275E3944F6EE33B8860F1DABE594">Zootaxa</mods:title>
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<mods:date id="8707A58C9F6A56EA4A63A659201DE9B2">2019</mods:date>
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<mods:number id="DC887C27A6875886871DE44B05D54D5B">2019-11-15</mods:number>
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<mods:number id="2A4B08E7DC39FD15DB1BD44006F3A4CF">4698</mods:number>
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<mods:identifier id="32587FE32CBF1AFB32C3C588C81706E3" type="DOI">10.11646/zootaxa.4698.1.1</mods:identifier>
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<treatment id="06130E72A523FFCAFF34FBFEFDD2AE9E" ID-DOI="http://doi.org/10.5281/zenodo.5623817" ID-GBIF-Taxon="160406584" ID-Zenodo-Dep="5623817" LSID="urn:lsid:plazi:treatment:06130E72A523FFCAFF34FBFEFDD2AE9E" httpUri="http://treatment.plazi.org/id/06130E72A523FFCAFF34FBFEFDD2AE9E" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="C6A0ECEFA523FFC5FF34FBFEFD82A809" box="[151,585,1058,1085]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8E05BF64A523FFC5FF34FBFEFD82A809" blockId="39.[151,585,1058,1120]" box="[151,585,1058,1085]" pageId="39" pageNumber="40">
<heading id="D54D0808A523FFC5FF34FBFEFD82A809" bold="true" box="[151,585,1058,1085]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="BCCE6376A523FFC5FF34FBFEFD82A809" bold="true" box="[151,585,1058,1085]" pageId="39" pageNumber="40">
<taxonomicName id="49BAC4E7A523FFC5FF34FBFEFE79A809" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[151,434,1058,1085]" class="Chilopoda" family="Aphilodontidae" genus="Mecophilus" kingdom="Animalia" order="Geophilomorpha" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="tupiniquim" status="sp. nov.">
<emphasis id="BCCE6376A523FFC5FF34FBFEFE79A809" bold="true" box="[151,434,1058,1085]" italics="true" pageId="39" pageNumber="40">Mecophilus tupiniquim</emphasis>
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<taxonomicNameLabel id="A7FDDE0DA523FFC5FE1AFBFFFD82A809" box="[441,585,1058,1085]" pageId="39" pageNumber="40" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C6A0ECEFA523FFC5FF34FB9AFDF2A855" box="[151,569,1094,1120]" pageId="39" pageNumber="40" type="description">
<paragraph id="8E05BF64A523FFC5FF34FB9AFDF2A855" blockId="39.[151,585,1058,1120]" box="[151,569,1094,1120]" pageId="39" pageNumber="40">
(
<figureCitation id="1681A3E1A523FFC5FF3CFB9AFF3DA855" box="[159,246,1094,1120]" captionStart="FIGURE 30" captionStartId="37.[151,250,1709,1734]" captionTargetBox="[157,1427,1361,1686]" captionTargetId="figure@37.[151,1435,1354,1690]" captionTargetPageId="37" captionText="FIGURE 30. AC, Mecophilus diagnostic characters. Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, forcipules, ventral; B, first leg-bearing segment, ventral; C, intermediate part of trunk, ventral. Abbreviations: DTA: denticle of tarsungulum; FD1: distal denticle of trocantheroprefemur; FD2: denticle of femur; FD3: denticle of tibia; ST: sternite; STF: sternite of first leg-bearing segment. Scale bars: 0.04 mm (AC)." figureDoi="http://doi.org/10.5281/zenodo.3542939" httpUri="https://zenodo.org/record/3542939/files/figure.png" pageId="39" pageNumber="40">Figs 30</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FEA1FB9AFEEBA855" box="[258,288,1094,1120]" captionStart="FIGURE 32" captionStartId="40.[151,250,1380,1405]" captionTargetBox="[157,1429,701,1354]" captionTargetId="figure@40.[151,1436,697,1361]" captionTargetPageId="40" captionText="FIGURE 32. AG, Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, denticles of left forcipule, ventral; E, terminal part of body, ventral. Female holotype (IBSP 6172): F, terminal part of body, ventral; G, pretarsus of ultimate leg, lateral. Abbreviations: FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; DTA: denticle of tarsungulum; FPT: forcipular pretergite; FTE: forcipular tergite; PT: pretarsus. Scale bars: 1 (A), 0.1 (BC, EF), 0.02 (D, G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542943" httpUri="https://zenodo.org/record/3542943/files/figure.png" pageId="39" pageNumber="40">32</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FE8FFB9AFE93A855" box="[300,344,1094,1120]" captionStart="FIGURE 40" captionStartId="50.[151,250,1910,1935]" captionTargetBox="[214,1373,1179,1883]" captionTargetId="figure@50.[208,1379,1175,1890]" captionTargetPageId="50" captionText="FIGURE 40. AF, anterior part of body, dorsal: A, Dicellophilus carniolensis (Koch, 1847) (PD); B, Aphilodon pereirai n. sp. (IBSP); C, Philacroterium cribellatum Attems, 1926 (SAM); D, Mairata itatiaiensis n. sp. (IBSP); E, Mecophilus neotropicus (MSNG); F, Mecophilus tupiniquim n. sp. (IBSP). Abbreviations: FPT: forcipular pretergite; FTE: forcipular tergite. Scale bars: 0.5 mm (AF)." figureDoi="http://doi.org/10.5281/zenodo.3542961" httpUri="https://zenodo.org/record/3542961/files/figure.png" pageId="39" pageNumber="40">40F</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FEC0FB9AFE5DA855" box="[355,406,1094,1120]" captionStart="FIGURE 42" captionStartId="52.[151,250,1344,1369]" captionTargetBox="[171,1395,181,1319]" captionTargetId="figure@52.[160,1402,174,1325]" captionTargetPageId="52" captionText="FIGURE 42. AI, forcipule, ventral: A, Ribautia centralis (Silvestri, 1907) (INPA); B, Geoperingueyia dentata Verhoeff, 1938 (MNRJ); C, Mairata butantan n. sp. (IBSP); D, Aphilodon indespectus n. sp. (IBSP); E, Aphilodon silvestrii n. sp. (IBSP); F, Aphilodon micronyx Brölemann, 1902 (IBSP); G, Mecophilus tupiniquim n. sp. (IBSP); H, Philacroterium weberi (Silvestri, 1909) n. comb. (KZN); I, Philacroterium maritimum (Lawrence, 1963) n. comb. (KZN). Abbreviations: FE: femur; TA: tarsungulum; TI: tibia; TFF: trochanteroprefemur+femur; TR: trocantheroprefemur. Scale bars: 0.2 (AF, HI), 0.1 (G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542965" httpUri="https://zenodo.org/record/3542965/files/figure.png" pageId="39" pageNumber="40">42G</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FE02FB9AFE1FA855" box="[417,468,1094,1120]" captionStart="FIGURE 43" captionStartId="53.[151,250,1556,1581]" captionTargetBox="[195,1392,415,1531]" captionTargetId="figure@53.[189,1398,409,1536]" captionTargetPageId="53" captionText="FIGURE 43. AD, forcipular denticles, ventral: A, Ribautia centralis (Silvestri, 1907) (INPA); B, Geoperingueyia dentata Verhoeff, 1938 (MNRJ); C, Aphilodon pereirai n. sp. (IBSP); D, Mecophilus tupiniquim n. sp. (IBSP). Abbreviations: DTA: denticle of tarsungulum; FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; PDT: proximal denticle of trochanteroprefemur. Scale bars: 0.2 (AC), 0.02 (D) mm." figureDoi="http://doi.org/10.5281/zenodo.3542967" httpUri="https://zenodo.org/record/3542967/files/figure.png" pageId="39" pageNumber="40">43D</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FE43FB9AFE37A855" box="[480,508,1094,1120]" captionStart="FIGURE 44" captionStartId="54.[151,250,1724,1749]" captionTargetBox="[159,1428,524,1696]" captionTargetId="figure@54.[151,1437,517,1704]" captionTargetPageId="54" captionText="FIGURE 44. AP, ultimate leg-bearing segment, ventral: A, Dicellophilus carniolensis (Koch, 1847) (PD), male; B, Pachymerium ferrugineum (Koch, 1835) (PD), female; C, Macronicophilus abbreviatus Pereira, Foddai &amp; Minelli, 2000 (INPA), male; D, Henia vesuviana (Newport, 1844) (PD), male; E, Strigamia crassipes (Koch, 1835) (PD), male; F, Geoperingueyia dentata Verhoeff, 1938 (MNRJ), female; G, Geoperingueyia dentata Verhoeff, 1938 (MNRJ), male; H, Philacroterium maritimum (Lawrence, 1963) n. comb (KZN), female; I, Philacroterium brevipes (Verhoeff, 1938) n. comb. (KZN), male; J, Philacroterium cribellatum Attems, 1926 (NHMW), male, (photo by N.Akkari); K, Mecophilus tupiniquim n. sp. (IBSP), female; L, Mecophilus tupiniquim n. sp. (IBSP), male; M, Mairata itatiaiensis n. sp. (IBSP), female; N, Mairata itatiaiensis n. sp. (IBSP), male; O, Aphilodon micronyx Brölemann, 1902 (IBSP), female; P, Aphilodon pereirai n. sp. (IBSP), male. Scale bars: 0.2 mm (AP)." figureDoi="http://doi.org/10.5281/zenodo.3542973" httpUri="https://zenodo.org/record/3542973/files/figure.png" pageId="39" pageNumber="40">44</figureCitation>
KL)
</paragraph>
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<emphasis id="BCCE6376A523FFC5FF34FB53FE91A89C" bold="true" box="[151,346,1166,1193]" pageId="39" pageNumber="40">Type specimens.</emphasis>
<materialsCitation id="3ED2B539A523FFC5FEC1FB52FE95A8F8" ID-GBIF-Occurrence="2451575491" collectingDate="2017-11-15" collectionCode="IBSP" collectorName="V. Calvanese" country="Brazil" location="Morro do Cruzeiro" municipality="Sao Roque" pageId="39" pageNumber="40" specimenCode="IBSP 6172" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="510101C6A523FFC5FEC1FB52FE04A89D" box="[354,463,1166,1192]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
:
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, from
<location id="8B65E9BFA523FFC5FD9FFB53FCDFA89D" LSID="urn:lsid:plazi:treatment:06130E72A523FFCAFF34FBFEFDD2AE9E:8B65E9BFA523FFC5FD9FFB53FCDFA89D" box="[572,788,1166,1193]" country="Brazil" municipality="Sao Roque" name="Morro do Cruzeiro" pageId="39" pageNumber="40" stateProvince="Sao Paulo">Morro do Cruzeiro</location>
,
<collectingMunicipality id="6E61251EA523FFC5FC83FB52FC56A89C" box="[800,925,1166,1193]" pageId="39" pageNumber="40">São Roque</collectingMunicipality>
,
<collectingRegion id="4C7E7186A523FFC5FC0AFB52FBD5A89D" box="[937,1054,1166,1192]" country="Brazil" name="Sao Paulo" pageId="39" pageNumber="40">São Paulo</collectingRegion>
,
<collectingCountry id="F6ADFFF4A523FFC5FB89FB52FBBAA89D" box="[1066,1137,1166,1192]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
,
<date id="FA0499A4A523FFC5FBDEFB52FB35A89C" box="[1149,1278,1166,1193]" pageId="39" pageNumber="40" value="2017-11-15">
<collectingDate id="EA40604CA523FFC5FBDEFB52FB35A89C" box="[1149,1278,1166,1193]" pageId="39" pageNumber="40" value="2017-11-15">15/11/2017</collectingDate>
</date>
,
<collectorName id="234FDAB2A523FFC5FAA9FB53FA57A89D" box="[1290,1436,1166,1193]" pageId="39" pageNumber="40">V. Calvanese</collectorName>
col., (
<specimenCode id="DE1C171FA523FFC5FF75FB6EFE9DA8F8" box="[214,342,1202,1229]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="39" pageNumber="40">IBSP 6172</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3ED2B539A523FFC5FECBFB6FFC69A8F9" ID-GBIF-Occurrence="2451575496" box="[360,930,1202,1229]" collectingDate="2017-11-15" collectionCode="IBSP" collectorName="V. Calvanese" country="Brazil" location="Morro do Cruzeiro" municipality="Sao Roque" pageId="39" pageNumber="40" specimenCode="IBSP 6173" specimenCount="4" specimenCount-male="4" stateProvince="Sao Paulo" typeStatus="paratype">
<typeStatus id="510101C6A523FFC5FECBFB6FFE12A8F8" box="[360,473,1203,1229]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="98BC74EDA523FFC5FE47FB6EFDCCA8F9" box="[484,519,1202,1228]" pageId="39" pageNumber="40" type="male">4♂</specimenCount>
(
<specimenCode id="DE1C171FA523FFC5FDB6FB6EFD5FA8F8" box="[533,660,1202,1229]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="39" pageNumber="40">IBSP 6173</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
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<materialsCitation id="3ED2B539A523FFC5FF64FB0BFB74A8C5" ID-GBIF-Occurrence="2451575494" box="[199,1215,1238,1265]" country="Brazil" latitude="-23.538332" location="Morro do Cruzeiro" longLatPrecision="21" longitude="-47.14528" municipality="Sao Roque" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="BCCE6376A523FFC5FF64FB0BFEADA8C4" bold="true" box="[199,358,1238,1265]" pageId="39" pageNumber="40">
<typeStatus id="510101C6A523FFC5FF64FB0BFEC9A8C4" box="[199,258,1239,1265]" pageId="39" pageNumber="40">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F6ADFFF4A523FFC5FECFFB0AFE7EA8C5" box="[364,437,1238,1264]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
:
<collectingRegion id="4C7E7186A523FFC5FE63FB0AFDFEA8C5" box="[448,565,1238,1264]" country="Brazil" name="Sao Paulo" pageId="39" pageNumber="40">São Paulo</collectingRegion>
:
<collectingMunicipality id="6E61251EA523FFC5FD9CFB0AFD75A8C4" box="[575,702,1238,1265]" pageId="39" pageNumber="40">São Roque</collectingMunicipality>
:
<location id="8B65E9BFA523FFC5FD6BFB0BFC6BA8C5" LSID="urn:lsid:plazi:treatment:06130E72A523FFCAFF34FBFEFDD2AE9E:8B65E9BFA523FFC5FD6BFB0BFC6BA8C5" box="[712,928,1238,1265]" country="Brazil" latitude="-23.538332" longLatPrecision="21" longitude="-47.14528" municipality="Sao Roque" name="Morro do Cruzeiro" pageId="39" pageNumber="40" stateProvince="Sao Paulo">Morro do Cruzeiro</location>
:
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,
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.
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<paragraph id="8E05BF64A523FFC5FF64FB26FBBEA90C" blockId="39.[151,1437,1166,2021]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64FB26FE73A920" bold="true" box="[199,440,1274,1301]" pageId="39" pageNumber="40">Material examined.</emphasis>
<materialsCitation id="3ED2B539A523FFC5FE64FB26FBBBA90C" ID-GBIF-Occurrence="2451575498" collectingDate="2017-11-15" collectionCode="IBSP" collectorName="V. Calvanese" country="Brazil" latitude="-23.538332" location="Morro do Cruzeiro" longLatPrecision="21" longitude="-47.14528" municipality="Sao Roque" pageId="39" pageNumber="40" specimenCode="IBSP 6172, IBSP 6173" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="BCCE6376A523FFC5FE64FB26FDF9A920" bold="true" box="[455,562,1274,1301]" pageId="39" pageNumber="40">
<collectingCountry id="F6ADFFF4A523FFC5FE64FB26FDF9A920" box="[455,562,1274,1301]" name="Brazil" pageId="39" pageNumber="40">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="BCCE6376A523FFC5FDEAFB26FD0CA920" bold="true" box="[585,711,1274,1301]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="4C7E7186A523FFC5FDEAFB26FD0CA920" box="[585,711,1274,1301]" country="Brazil" name="Sao Paulo" pageId="39" pageNumber="40">São Paulo</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6E61251EA523FFC5FD7DFB26FCA8A920" box="[734,867,1274,1301]" pageId="39" pageNumber="40">São Roque</collectingMunicipality>
,
<location id="8B65E9BFA523FFC5FCD6FB27FB93A921" LSID="urn:lsid:plazi:treatment:06130E72A523FFCAFF34FBFEFDD2AE9E:8B65E9BFA523FFC5FCD6FB27FB93A921" box="[885,1112,1274,1301]" country="Brazil" latitude="-23.538332" longLatPrecision="21" longitude="-47.14528" municipality="Sao Roque" name="Morro do Cruzeiro" pageId="39" pageNumber="40" stateProvince="Sao Paulo">Morro do Cruzeiro</location>
(
<geoCoordinate id="EB8ED9A3A523FFC5FBD3FB26FB3CA921" box="[1136,1271,1274,1300]" degrees="23" direction="south" minutes="32" orientation="latitude" pageId="39" pageNumber="40" precision="15" seconds="18" value="-23.538332">23°3218”S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A523FFC5FAA8FB26FA44A921" box="[1291,1423,1274,1300]" degrees="47" direction="west" minutes="8" orientation="longitude" pageId="39" pageNumber="40" precision="15" seconds="43" value="-47.14528">47°843”W</geoCoordinate>
),
<date id="FA0499A4A523FFC5FF34FAC2FED3A90C" box="[151,280,1310,1337]" pageId="39" pageNumber="40" value="2017-11-15">
<collectingDate id="EA40604CA523FFC5FF34FAC2FED3A90C" box="[151,280,1310,1337]" pageId="39" pageNumber="40" value="2017-11-15">15/11/2017</collectingDate>
</date>
,
<collectorName id="234FDAB2A523FFC5FE80FAC3FE7EA90D" box="[291,437,1310,1337]" pageId="39" pageNumber="40">V. Calvanese</collectorName>
col.,
<specimenCount id="98BC74EDA523FFC5FE50FAC2FDDDA90D" box="[499,534,1310,1336]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
[
<typeStatus id="510101C6A523FFC5FD87FAC2FD47A90C" box="[548,652,1310,1337]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
] (
<specimenCode id="DE1C171FA523FFC5FD03FAC2FCD4A90C" box="[672,799,1310,1337]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="39" pageNumber="40">IBSP 6172</specimenCode>
),
<specimenCount id="98BC74EDA523FFC5FC91FAC2FC9EA90D" box="[818,853,1310,1336]" pageId="39" pageNumber="40" type="male">4♂</specimenCount>
[
<typeStatus id="510101C6A523FFC5FCC0FAC2FC1FA90C" box="[867,980,1310,1337]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
] (
<specimenCode id="DE1C171FA523FFC5FC4AFAC2FBA2A90C" box="[1001,1129,1310,1337]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="39" pageNumber="40">IBSP 6173</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA523FFC5FF64FA9FFA98A969" box="[199,1363,1346,1373]" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8E05BF64A523FFC5FF64FA9FFA98A969" blockId="39.[151,1437,1166,2021]" box="[199,1363,1346,1373]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64FA9FFE85A968" bold="true" box="[199,334,1347,1373]" pageId="39" pageNumber="40">Etymology.</emphasis>
The epithet is an adjective and means “originating in
<collectingCountry id="F6ADFFF4A523FFC5FC08FA9EFC39A969" box="[939,1010,1346,1372]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
” in the Tupi-Guarani language.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA523FFC5FF64FABAFC1EAA25" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8E05BF64A523FFC5FF64FABAFD5CA9D9" blockId="39.[151,1437,1166,2021]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64FABAFE89A9B4" bold="true" box="[199,322,1382,1409]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
A species of
<taxonomicName id="49BAC4E7A523FFC5FE79FABAFD96A9B5" authorityName=": Silvestri" authorityYear="1909" box="[474,605,1382,1408]" class="Chilopoda" family="Aphilodontidae" genus="Mecophilus" kingdom="Animalia" order="Geophilomorpha" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="BCCE6376A523FFC5FE79FABAFD96A9B5" box="[474,605,1382,1408]" italics="true" pageId="39" pageNumber="40">Mecophilus</emphasis>
</taxonomicName>
with: known body length in adults 7.0
<quantity id="49421281A523FFC5FB83FABAFBB2A9B4" box="[1056,1145,1382,1409]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="39" pageNumber="40" unit="mm" value="7.5">7.5 mm</quantity>
; cephalic plate subrectangular (
<figureCitation id="1681A3E1A523FFC5FF42FA56FE86A991" box="[225,333,1418,1444]" captionStart="FIGURE 32" captionStartId="40.[151,250,1380,1405]" captionTargetBox="[157,1429,701,1354]" captionTargetId="figure@40.[151,1436,697,1361]" captionTargetPageId="40" captionText="FIGURE 32. AG, Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, denticles of left forcipule, ventral; E, terminal part of body, ventral. Female holotype (IBSP 6172): F, terminal part of body, ventral; G, pretarsus of ultimate leg, lateral. Abbreviations: FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; DTA: denticle of tarsungulum; FPT: forcipular pretergite; FTE: forcipular tergite; PT: pretarsus. Scale bars: 1 (A), 0.1 (BC, EF), 0.02 (D, G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542943" httpUri="https://zenodo.org/record/3542943/files/figure.png" pageId="39" pageNumber="40">Figs 32B</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FEF8FA56FE40A991" box="[347,395,1418,1444]" captionStart="FIGURE 40" captionStartId="50.[151,250,1910,1935]" captionTargetBox="[214,1373,1179,1883]" captionTargetId="figure@50.[208,1379,1175,1890]" captionTargetPageId="50" captionText="FIGURE 40. AF, anterior part of body, dorsal: A, Dicellophilus carniolensis (Koch, 1847) (PD); B, Aphilodon pereirai n. sp. (IBSP); C, Philacroterium cribellatum Attems, 1926 (SAM); D, Mairata itatiaiensis n. sp. (IBSP); E, Mecophilus neotropicus (MSNG); F, Mecophilus tupiniquim n. sp. (IBSP). Abbreviations: FPT: forcipular pretergite; FTE: forcipular tergite. Scale bars: 0.5 mm (AF)." figureDoi="http://doi.org/10.5281/zenodo.3542961" httpUri="https://zenodo.org/record/3542961/files/figure.png" pageId="39" pageNumber="40">40F</figureCitation>
); forcipular tergite subrectangular (
<figureCitation id="1681A3E1A523FFC5FCBEFA56FC42A991" box="[797,905,1418,1444]" captionStart="FIGURE 32" captionStartId="40.[151,250,1380,1405]" captionTargetBox="[157,1429,701,1354]" captionTargetId="figure@40.[151,1436,697,1361]" captionTargetPageId="40" captionText="FIGURE 32. AG, Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, denticles of left forcipule, ventral; E, terminal part of body, ventral. Female holotype (IBSP 6172): F, terminal part of body, ventral; G, pretarsus of ultimate leg, lateral. Abbreviations: FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; DTA: denticle of tarsungulum; FPT: forcipular pretergite; FTE: forcipular tergite; PT: pretarsus. Scale bars: 1 (A), 0.1 (BC, EF), 0.02 (D, G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542943" httpUri="https://zenodo.org/record/3542943/files/figure.png" pageId="39" pageNumber="40">Figs 32B</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FC35FA56FC0DA991" box="[918,966,1418,1444]" captionStart="FIGURE 40" captionStartId="50.[151,250,1910,1935]" captionTargetBox="[214,1373,1179,1883]" captionTargetId="figure@50.[208,1379,1175,1890]" captionTargetPageId="50" captionText="FIGURE 40. AF, anterior part of body, dorsal: A, Dicellophilus carniolensis (Koch, 1847) (PD); B, Aphilodon pereirai n. sp. (IBSP); C, Philacroterium cribellatum Attems, 1926 (SAM); D, Mairata itatiaiensis n. sp. (IBSP); E, Mecophilus neotropicus (MSNG); F, Mecophilus tupiniquim n. sp. (IBSP). Abbreviations: FPT: forcipular pretergite; FTE: forcipular tergite. Scale bars: 0.5 mm (AF)." figureDoi="http://doi.org/10.5281/zenodo.3542961" httpUri="https://zenodo.org/record/3542961/files/figure.png" pageId="39" pageNumber="40">40F</figureCitation>
); denticle of tarsungulum relatively large and positioned close to basis of tarsungulum (
<figureCitation id="1681A3E1A523FFC5FD3DFA72FCC0A9FD" box="[670,779,1454,1480]" captionStart="FIGURE 30" captionStartId="37.[151,250,1709,1734]" captionTargetBox="[157,1427,1361,1686]" captionTargetId="figure@37.[151,1435,1354,1690]" captionTargetPageId="37" captionText="FIGURE 30. AC, Mecophilus diagnostic characters. Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, forcipules, ventral; B, first leg-bearing segment, ventral; C, intermediate part of trunk, ventral. Abbreviations: DTA: denticle of tarsungulum; FD1: distal denticle of trocantheroprefemur; FD2: denticle of femur; FD3: denticle of tibia; ST: sternite; STF: sternite of first leg-bearing segment. Scale bars: 0.04 mm (AC)." figureDoi="http://doi.org/10.5281/zenodo.3542939" httpUri="https://zenodo.org/record/3542939/files/figure.png" pageId="39" pageNumber="40">Figs 30A</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FCB4FA72FC87A9FD" box="[791,844,1454,1480]" captionStart="FIGURE 32" captionStartId="40.[151,250,1380,1405]" captionTargetBox="[157,1429,701,1354]" captionTargetId="figure@40.[151,1436,697,1361]" captionTargetPageId="40" captionText="FIGURE 32. AG, Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, denticles of left forcipule, ventral; E, terminal part of body, ventral. Female holotype (IBSP 6172): F, terminal part of body, ventral; G, pretarsus of ultimate leg, lateral. Abbreviations: FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; DTA: denticle of tarsungulum; FPT: forcipular pretergite; FTE: forcipular tergite; PT: pretarsus. Scale bars: 1 (A), 0.1 (BC, EF), 0.02 (D, G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542943" httpUri="https://zenodo.org/record/3542943/files/figure.png" pageId="39" pageNumber="40">32D</figureCitation>
); distal denticle of trochanteroprefemur and denticle of femur close to each other (
<figureCitation id="1681A3E1A523FFC5FE7CFA0EFD80A9D9" box="[479,587,1490,1516]" captionStart="FIGURE 30" captionStartId="37.[151,250,1709,1734]" captionTargetBox="[157,1427,1361,1686]" captionTargetId="figure@37.[151,1435,1354,1690]" captionTargetPageId="37" captionText="FIGURE 30. AC, Mecophilus diagnostic characters. Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, forcipules, ventral; B, first leg-bearing segment, ventral; C, intermediate part of trunk, ventral. Abbreviations: DTA: denticle of tarsungulum; FD1: distal denticle of trocantheroprefemur; FD2: denticle of femur; FD3: denticle of tibia; ST: sternite; STF: sternite of first leg-bearing segment. Scale bars: 0.04 mm (AC)." figureDoi="http://doi.org/10.5281/zenodo.3542939" httpUri="https://zenodo.org/record/3542939/files/figure.png" pageId="39" pageNumber="40">Figs 30A</figureCitation>
,
<figureCitation id="1681A3E1A523FFC5FDF5FA0EFD40A9D9" box="[598,651,1490,1516]" captionStart="FIGURE 32" captionStartId="40.[151,250,1380,1405]" captionTargetBox="[157,1429,701,1354]" captionTargetId="figure@40.[151,1436,697,1361]" captionTargetPageId="40" captionText="FIGURE 32. AG, Mecophilus tupiniquim n. sp. Male paratype (IBSP 6173): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, denticles of left forcipule, ventral; E, terminal part of body, ventral. Female holotype (IBSP 6172): F, terminal part of body, ventral; G, pretarsus of ultimate leg, lateral. Abbreviations: FD1: distal denticle of trochanteroprefemur; FD2: denticle of femur; FD3: denticle of tibia; DTA: denticle of tarsungulum; FPT: forcipular pretergite; FTE: forcipular tergite; PT: pretarsus. Scale bars: 1 (A), 0.1 (BC, EF), 0.02 (D, G) mm." figureDoi="http://doi.org/10.5281/zenodo.3542943" httpUri="https://zenodo.org/record/3542943/files/figure.png" pageId="39" pageNumber="40">32D</figureCitation>
).
</paragraph>
<paragraph id="8E05BF64A523FFC5FF64FA2BFC1EAA25" blockId="39.[151,1437,1166,2021]" box="[199,981,1526,1553]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64FA2BFE71AA24" bold="true" box="[199,442,1526,1553]" pageId="39" pageNumber="40">Recorded specimens</emphasis>
: 5, including
<specimenCount id="98BC74EDA523FFC5FDF5FA2AFDB2AA25" box="[598,633,1526,1552]" pageId="39" pageNumber="40" type="female">1♀</specimenCount>
and
<specimenCount id="98BC74EDA523FFC5FD13FA2AFD1FAA25" box="[688,724,1526,1552]" pageId="39" pageNumber="40" type="male">4♂</specimenCount>
, from a single locality.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA523FFC5FF64F9C6FD02AA00" box="[199,713,1562,1589]" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8E05BF64A523FFC5FF64F9C6FD02AA00" blockId="39.[151,1437,1166,2021]" box="[199,713,1562,1589]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64F9C6FEAAAA00" bold="true" box="[199,353,1562,1589]" pageId="39" pageNumber="40">Distribution.</emphasis>
Brazil—São Paulo: São Roque.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA523FFCAFF64F9E2FDD2AE9E" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8E05BF64A523FFC5FF64F9E2FD13AA6C" blockId="39.[151,1437,1166,2021]" box="[199,728,1598,1625]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64F9E2FE2EAA6C" bold="true" box="[199,485,1598,1625]" pageId="39" pageNumber="40">
Description of
<typeStatus id="510101C6A523FFC5FED4F9E3FE2BAA6C" box="[375,480,1599,1625]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
.
</emphasis>
Female (IBSP 6172).
</paragraph>
<paragraph id="8E05BF64A523FFC5FF64F9BFFBD3AAF0" blockId="39.[151,1437,1166,2021]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64F9BFFE83AA49" box="[199,328,1634,1660]" italics="true" pageId="39" pageNumber="40">Entire body</emphasis>
: length 7.5, maximum width 0.17.
<emphasis id="BCCE6376A523FFC5FD68F9BEFCA5AA49" box="[715,878,1634,1660]" italics="true" pageId="39" pageNumber="40">Cephalic plate</emphasis>
: sub-rectangular, length 0.20 and maximum width (at about mid-length) 0.13 (length/width ratio 1.53).
<emphasis id="BCCE6376A523FFC5FD4FF95BFC99AA94" box="[748,850,1671,1697]" italics="true" pageId="39" pageNumber="40">Antenna:</emphasis>
length 0.48, ca. 2.4 times as long as cephalic plate; AA IIXIII length/width ratio 0.550.70; LAA ca. 4.2 times as long as AA XIII.
</paragraph>
<paragraph id="8E05BF64A523FFC5FF64F912FAE6AB61" blockId="39.[151,1437,1166,2021]" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64F912FEE1AADD" box="[199,298,1742,1768]" italics="true" pageId="39" pageNumber="40">Clypeus:</emphasis>
2+2 LS, 1+1 PS, 5+5 SCS.
<emphasis id="BCCE6376A523FFC5FDCBF912FD18AADD" box="[616,723,1742,1768]" italics="true" pageId="39" pageNumber="40">Mandible</emphasis>
: PL with ca. 20 denticles.
<emphasis id="BCCE6376A523FFC5FBA3F913FB56AADD" box="[1024,1181,1742,1768]" italics="true" pageId="39" pageNumber="40">First maxillae</emphasis>
: TFM length 0.02 and width 0.02; DA with 3 apical and 1 subapical SL; MPC with 2 apical and 1 subapical SL.
<emphasis id="BCCE6376A523FFC5FBD0F92EFAE1AB39" box="[1139,1322,1778,1804]" italics="true" pageId="39" pageNumber="40">Second maxillae</emphasis>
: left TSM length 0.02 (1.15 times as long as TFM) and maximum width (basal margin) 0.01; left T3 length 0.01 (0.75 times as long as T2) and width 0.01 (0.70 times as wide as T2); both TSM with 1 apical and 1 subapical large SE.
</paragraph>
<paragraph id="8E05BF64A523FFCAFF64F882FACDACE2" blockId="39.[151,1437,1166,2021]" lastBlockId="40.[151,1436,153,683]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="BCCE6376A523FFC5FF64F882FE65AB4D" box="[199,430,1886,1912]" italics="true" pageId="39" pageNumber="40">Forcipular segment:</emphasis>
FTE length 0.07 (0.36 times as long as cephalic plate) and width 0.15 (length/width ratio 0.48); FPT length 0.05 (0.25 times as long as cephalic plate) and width 0.09; FCX length 0.25 and width 0.20 (length/width ratio 1.25); FRL length 0.22, surpassing anterior margin of cephalic plate, and maximum width (basal margin) 0.08; FRL with two denticles in TFF and one in TI; FD1 relatively large, pointed and without SE, positioned relatively close to FD2; FD2 bilobed, with 1 large apical and 1 large subapical SE; FD3 bilobed, with 1 large apical and 3 smaller subapical SE; DTA relatively large, near the basis of TA; TA 0.48 times as long as TFF.
</paragraph>
<paragraph id="8E05BF64A52CFFCAFF64FF3DFBA7ADBE" blockId="40.[151,1436,153,683]" pageId="40" pageNumber="41">
<emphasis id="BCCE6376A52CFFCAFF64FF3DFE9EACCE" box="[199,341,225,251]" italics="true" pageId="40" pageNumber="41">Walking legs</emphasis>
: 35 pairs; left FL length 0.17, 0.83 times as long as second leg; left leg 17 length 0.21.
<emphasis id="BCCE6376A52CFFCAFAACFF3EFD4EAD2A" italics="true" pageId="40" pageNumber="41">Tergum from the first to penultimate leg-bearing segment</emphasis>
: TE 17 length 0.09 and width 0.17 (length/width ratio 0.52), with one anterior row of 8 SE and one posterior row of little smaller SE; PT 0.57 times as long as TE and with two parallel rows of 6 SE.
<emphasis id="BCCE6376A52CFFCAFE9FFE91FC04AD52" box="[316,975,333,359]" italics="true" pageId="40" pageNumber="41">Sternum from the first to penultimate leg-bearing segment</emphasis>
: STF rectangular, length 0.10 and width 0.08 (length/width ratio 1.2); ST 17 length 0.13 and width 0.09 (length/width ratio 1.4).
</paragraph>
<paragraph id="8E05BF64A52CFFCAFF64FE49FEA5AE0A" blockId="40.[151,1436,153,683]" pageId="40" pageNumber="41">
<emphasis id="BCCE6376A52CFFCAFF64FE49FDD5AD9A" box="[199,542,405,431]" italics="true" pageId="40" pageNumber="41">Ultimate leg-bearing segment</emphasis>
: MTL trapezoidal, length 0.07 and maximum width 0.10 (length/width ratio 0.66); UPS exposed and not divided, length 0.01 and width 0.10.
<emphasis id="BCCE6376A52CFFCAFCC9FE65FBC2ADE6" box="[874,1033,441,467]" italics="true" pageId="40" pageNumber="41">Ultimate legs:</emphasis>
with two tarsal articles and terminal claw; slightly thickened when compared to penultimate leg; length 0.43, 1.83 times as long as penultimate leg; each CXP with 5or 6 little COP that are weakly grouped near MTL and 1 COP further away.
<emphasis id="BCCE6376A52CFFCAFB2AFDDDFAA3AE2E" box="[1161,1384,513,539]" italics="true" pageId="40" pageNumber="41">Postpedal segments</emphasis>
: GF with 3+3 distal SE.
</paragraph>
<paragraph id="8E05BF64A52CFFCAFF64FD95FDD2AE9E" blockId="40.[151,1436,153,683]" pageId="40" pageNumber="41">
<emphasis id="BCCE6376A52CFFCAFF64FD95FE52AE56" bold="true" box="[199,409,584,611]" pageId="40" pageNumber="41">Known variation.</emphasis>
<emphasis id="BCCE6376A52CFFCAFE3CFD95FDFEAE56" box="[415,565,585,611]" italics="true" pageId="40" pageNumber="41">Ultimate legs</emphasis>
: each coxopleuron with 57 COP in adults (
<specimenCount id="98BC74EDA52CFFCAFB86FD95FB8DAE56" box="[1061,1094,585,611]" pageId="40" pageNumber="41" type="female">1♀</specimenCount>
,
<specimenCount id="98BC74EDA52CFFCAFBF0FD95FBB2AE56" box="[1107,1145,585,611]" pageId="40" pageNumber="41" type="male">4♂</specimenCount>
).
<emphasis id="BCCE6376A52CFFCAFB2EFD95FB0CAE56" box="[1165,1223,585,611]" italics="true" pageId="40" pageNumber="41">Male</emphasis>
: ultimate legs with a single tarsal article, little more thickened than in female, without pretarsus; GM biarticulated; GM1 with 4 or 5 SE in one row; GM2 with 7 distal SE.
</paragraph>
</subSubSection>
</treatment>
</document>