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<document id="7A446EC901F9B1F17C7A668E95516129" ID-CLB-Dataset="24882" ID-DOI="10.11646/zootaxa.4698.1.1" ID-GBIF-Dataset="1c94057f-64de-4f19-b047-2223e4d664e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3542876" ID-ZooBank="0B4482B9-F89F-45CA-B929-E9B098605CD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573811688707" checkinUser="plazi" docAuthor="Calvanese, Victor C., Brescovit, Antonio D. &amp; Bonato, Lucio" docDate="2019" docId="06130E72A51AFFC2FF34F92BFD5BAE07" docLanguage="en" docName="zootaxa.4698.1.1.pdf" docOrigin="Zootaxa 4698 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aphilodon meganae Calvanese &amp; Brescovit &amp; Bonato 2019, new species" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FA2A760AA504FFE2FFA3FFDCFFCBAC35" masterDocTitle="Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily" masterLastPageNumber="72" masterPageNumber="1" pageNumber="31" updateTime="1698767377376" updateUser="ExternalLinkService">
<mods:mods id="9E6B85864A646574DBDFA1AA927B4E38" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="52D4D27D94470B99AED301B3191062F2">
<mods:title id="F7C234914F6C6F41F109468F6ECDBF75">Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily</mods:title>
</mods:titleInfo>
<mods:name id="3A69554CE3F3A83E4AD7860E8957036C" type="personal">
<mods:role id="1B019C7135430846E19031D3CABEE57D">
<mods:roleTerm id="3790FD49560AA8DF4548090A60B2889E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B7EA744316D530988CDA43B3874C9136">Calvanese, Victor C.</mods:namePart>
</mods:name>
<mods:name id="A05E1853E514402121347DD75A0192C5" type="personal">
<mods:role id="E193FB9A83C8AA2B1E24856DE4830524">
<mods:roleTerm id="F57E371C623A532DFC1811D4D4960E8A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A8265ABB86A58766B97981D853015EF">Brescovit, Antonio D.</mods:namePart>
</mods:name>
<mods:name id="8718824C1C16E823E861E38F1A603F78" type="personal">
<mods:role id="A19C96ABF49905EF8F79D61F0CCEAB04">
<mods:roleTerm id="610EE66F0EEE52E9308AB8C7D2D78B0D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CEE4C84969CD07C686ECFF44CA0F05F">Bonato, Lucio</mods:namePart>
</mods:name>
<mods:typeOfResource id="FB884FA4E366816ECFC27C76378E7576">text</mods:typeOfResource>
<mods:relatedItem id="FF430BC0DEEDBDB3EDCE0DAD3509CA72" type="host">
<mods:titleInfo id="6CDB6E0B40AE083CAD308DDCDF8280F0">
<mods:title id="059069C182013813850E3DD8000E5D3E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="19B9BC013715D08266C3298B361BE0C4">
<mods:date id="69378D3A6CF6760EC53F3AEF4A12B77F">2019</mods:date>
<mods:detail id="20C4BF18C8218875B89C7581F2E030F3" type="pubDate">
<mods:number id="641BFDE762410C09559E90530DC4F306">2019-11-15</mods:number>
</mods:detail>
<mods:detail id="FE66BCD5D10A3719CE0CF93A26DCA37D" type="volume">
<mods:number id="E3871C92534A2A31D43C67BFB8415B38">4698</mods:number>
</mods:detail>
<mods:detail id="26095C34DCB6946E091F8ACEB4C0544F" type="issue">
<mods:number id="E6EEDC579BFC4739F1B6084BCA6C6AAC">1</mods:number>
</mods:detail>
<mods:extent id="8D99ECEDA22407050DC3FA1048AEB4D5" unit="page">
<mods:start id="477B962A854D39042F064FEBBC76B0EF">1</mods:start>
<mods:end id="68624D81344B67ED1D7E02B224C24033">72</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C5730FA7B3B6FC27EA5ED075189EF16B">journal article</mods:classification>
<mods:identifier id="8E3EA906286CEE1510BC4E6C5746E91B" type="CLB-Dataset">24882</mods:identifier>
<mods:identifier id="78720E1F669EAB51A827792B02200C76" type="DOI">10.11646/zootaxa.4698.1.1</mods:identifier>
<mods:identifier id="4C1985BA37DBE2027AAE902A2FA1F24C" type="GBIF-Dataset">1c94057f-64de-4f19-b047-2223e4d664e8</mods:identifier>
<mods:identifier id="FF740EEFADC067A4C503F5C69D62837F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9AC6DD5C2239FAF6C4BC013506F622B7" type="Zenodo-Dep">3542876</mods:identifier>
<mods:identifier id="0DB8FE19F97C32DDFDEE2C0D0E47A965" type="ZooBank">0B4482B9-F89F-45CA-B929-E9B098605CD5</mods:identifier>
</mods:mods>
<treatment id="06130E72A51AFFC2FF34F92BFD5BAE07" ID-DOI="http://doi.org/10.5281/zenodo.5623809" ID-GBIF-Taxon="160406605" ID-Zenodo-Dep="5623809" LSID="urn:lsid:plazi:treatment:06130E72A51AFFC2FF34F92BFD5BAE07" httpUri="http://treatment.plazi.org/id/06130E72A51AFFC2FF34F92BFD5BAE07" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C6A0ECEFA51AFFFCFF34F92BFDE9AB24" box="[151,546,1783,1810]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8E05BF64A51AFFFCFF34F92BFDE9AB24" blockId="30.[151,546,1783,1845]" box="[151,546,1783,1810]" pageId="30" pageNumber="31">
<heading id="D54D0808A51AFFFCFF34F92BFDE9AB24" bold="true" box="[151,546,1783,1810]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="BCCE6376A51AFFFCFF34F92BFDE9AB24" bold="true" box="[151,546,1783,1810]" pageId="30" pageNumber="31">
<taxonomicName id="49BAC4E7A51AFFFCFF34F92BFE40AB27" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[151,395,1783,1810]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="meganae" status="sp. nov.">
<emphasis id="BCCE6376A51AFFFCFF34F92BFE40AB27" bold="true" box="[151,395,1783,1810]" italics="true" pageId="30" pageNumber="31">Aphilodon meganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A7FDDE0DA51AFFFCFE31F92BFDE9AB24" box="[402,546,1783,1809]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51AFFFCFF34F8C7FE59AB00" box="[151,402,1818,1845]" pageId="30" pageNumber="31" type="description">
<paragraph id="8E05BF64A51AFFFCFF34F8C7FE59AB00" blockId="30.[151,546,1783,1845]" box="[151,402,1818,1845]" pageId="30" pageNumber="31">
(
<figureCitation id="1681A3E1A51AFFFCFF3CF8C7FF31AB00" box="[159,250,1818,1845]" captionStart="FIGURE 5" captionStartId="13.[151,250,975,1000]" captionTargetBox="[155,1432,181,918]" captionTargetId="figure@13.[151,1436,174,922]" captionTargetPageId="13" captionText="FIGURE 5. A–D, Aphilodontinae clypeus and labrum. Aphilodon indespectus n. sp. (IBSP 3653): A, clypeus, ventral. Aphilodon meganae n. sp. (IBSP 4070): B, middle part of clypeus, ventral. Aphilodon silvestrii n. sp. (IBSP 3665): C, labrum, ventral. Philacroterium weberi (Silvestri, 1909) n. comb. (KZN 8142): D, labrum, ventral. Abbreviations: LP: lateral pieces of labrum; LS: lateral setae of clypeus; PS: post-antennal setae of clypeus; SCS: sub-clypeal setae. Scale bars: 0.1 (A, C–D), 0.02 (B) mm." figureDoi="http://doi.org/10.5281/zenodo.3542886" httpUri="https://zenodo.org/record/3542886/files/figure.png" pageId="30" pageNumber="31">Figs 5B</figureCitation>
,
<figureCitation id="1681A3E1A51AFFFCFEA5F8C7FE84AB00" box="[262,335,1819,1845]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStartId-0="31.[151,250,1886,1911]" captionStartId-1="32.[151,250,1737,1762]" captionTargetBox-0="[158,1428,1389,1859]" captionTargetBox-1="[181,1412,584,1709]" captionTargetId-0="figure@31.[151,1436,1381,1866]" captionTargetId-1="figure@32.[174,1414,576,1718]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 23. A–C, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, right telopodite of second maxillae, ventro-lateral; B, forcipular segment, ventral; C, right forcipule, ventral. Abbreviations: PLM: posterior lateral margin of forcipule; T2: article 2 of telopodite of second maxillae; T3: article 3 of telopodite of second maxillae. Scale bars: 0.02 (A), 0.1 (B–C) mm." captionText-1="FIGURE 24. A–E, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, terminal part of body, dorsal; E, same, ventral. Scale bars: 2.2 (A), 0.4 (B–C), 0.2 (D–E) mm." figureDoi-0="http://doi.org/10.5281/zenodo.3542925" figureDoi-1="http://doi.org/10.5281/zenodo.3542927" httpUri-0="https://zenodo.org/record/3542925/files/figure.png" httpUri-1="https://zenodo.org/record/3542927/files/figure.png" pageId="30" pageNumber="31">23–24</figureCitation>
,
<figureCitation id="1681A3E1A51AFFFCFEF9F8C6FE41AB00" box="[346,394,1818,1845]" captionStart="FIGURE 45" captionStartId="55.[151,250,1014,1039]" captionTargetBox="[289,1302,183,984]" captionTargetId="figure@55.[279,1308,175,988]" captionTargetPageId="55" captionText="FIGURE 45. A–D, posterior part of body, male, ventral: A, Dicellophilus carniolensis (Koch, 1847) (PD); B, Aphilodon micronyx Brölemann, 1902 (IBSP); C, Aphilodon meganae n. sp. (IBSP); D, Mairata butantan n. sp. (IBSP). Abbreviations: COP: coxal pores; MTL: metasternite of ultimate leg-bearing segment. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.3542976" httpUri="https://zenodo.org/record/3542976/files/figure.png" pageId="30" pageNumber="31">45C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51AFFFDFF34F8BFFA8BACE2" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8E05BF64A51AFFFCFF34F8BFFB72AB94" blockId="30.[151,1436,1890,2026]" pageId="30" pageNumber="31">
<emphasis id="BCCE6376A51AFFFCFF34F8BFFE97AB48" bold="true" box="[151,348,1890,1917]" pageId="30" pageNumber="31">Type specimens.</emphasis>
<materialsCitation id="3ED2B539A51AFFFCFEC6F8BFFDD0AB94" ID-GBIF-Occurrence="2451575471" collectorName="V. Calvanese &amp; A. Silva" country="Brazil" location="Sao Paulo" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="510101C6A51AFFFCFEC6F8BFFE19AB48" box="[357,466,1891,1917]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
:
<specimenCount id="98BC74EDA51AFFFCFE7CF8B8FE3FAB4B" box="[479,500,1892,1918]" pageId="30" pageNumber="31" type="female">♀</specimenCount>
, from Reserva Bertary, Iporanga,
<collectingRegion id="4C7E7186A51AFFFCFC23F8BFFC3CAB48" box="[896,1015,1891,1917]" country="Brazil" name="Sao Paulo" pageId="30" pageNumber="31">São Paulo</collectingRegion>
,
<collectingCountry id="F6ADFFF4A51AFFFCFBA7F8BFFB80AB48" box="[1028,1099,1891,1917]" name="Brazil" pageId="30" pageNumber="31">Brazil</collectingCountry>
, 07/2016,
<collectorName id="234FDAB2A51AFFFCFB66F8B8FA92AB48" box="[1221,1369,1891,1918]" pageId="30" pageNumber="31">V. Calvanese</collectorName>
&amp;
<collectorName id="234FDAB2A51AFFFCFA22F8BFFF1AAB94" pageId="30" pageNumber="31">A. Silva</collectorName>
col., deposited in IBSP 4068
</materialsCitation>
.
<materialsCitation id="3ED2B539A51AFFFCFD86F854FB7EAB94" ID-GBIF-Occurrence="2451575475" box="[549,1205,1927,1954]" collectionCode="IBSP" collectorName="V. Calvanese &amp; A. Silva" country="Brazil" location="Sao Paulo" pageId="30" pageNumber="31" specimenCode="IBSP 4070" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Sao Paulo" typeStatus="paratype">
<typeStatus id="510101C6A51AFFFCFD86F854FD5DAB97" box="[549,662,1928,1954]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="98BC74EDA51AFFFCFD02F85BFD0DAB94" box="[673,710,1927,1953]" pageId="30" pageNumber="31" type="female">4♀</specimenCount>
,
<specimenCount id="98BC74EDA51AFFFCFD71F85BFD3FAB94" box="[722,756,1927,1953]" pageId="30" pageNumber="31" type="male">2♂</specimenCount>
(
<specimenCode id="DE1C171FA51AFFFCFCA0F85BFC49AB94" box="[771,898,1927,1953]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="30" pageNumber="31">IBSP 4070</specimenCode>
), same locality as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8E05BF64A51AFFFCFF64F877FB6FABF0" blockId="30.[151,1436,1890,2026]" box="[199,1188,1962,1990]" pageId="30" pageNumber="31">
<materialsCitation id="3ED2B539A51AFFFCFF64F877FB6BABF0" ID-GBIF-Occurrence="2451575473" box="[199,1184,1962,1990]" country="Brazil" latitude="-24.587223" location="Reserva Bertary" longLatPrecision="21" longitude="-48.594723" municipality="Iporanga" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="BCCE6376A51AFFFCFF64F877FEADABF0" bold="true" box="[199,358,1962,1989]" pageId="30" pageNumber="31">
<typeStatus id="510101C6A51AFFFCFF64F877FEC9ABF0" box="[199,258,1963,1989]" pageId="30" pageNumber="31">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F6ADFFF4A51AFFFCFECFF877FE7EABF0" box="[364,437,1963,1989]" name="Brazil" pageId="30" pageNumber="31">Brazil</collectingCountry>
:
<collectingRegion id="4C7E7186A51AFFFCFE63F877FDFEABF0" box="[448,565,1963,1989]" country="Brazil" name="Sao Paulo" pageId="30" pageNumber="31">São Paulo</collectingRegion>
:
<collectingMunicipality id="6E61251EA51AFFFCFD9CF870FD6CABF3" box="[575,679,1964,1990]" pageId="30" pageNumber="31">Iporanga</collectingMunicipality>
:
<location id="8B65E9BFA51AFFFCFD12F870FCA0ABF3" LSID="urn:lsid:plazi:treatment:06130E72A51AFFC2FF34F92BFD5BAE07:8B65E9BFA51AFFFCFD12F870FCA0ABF3" box="[689,875,1964,1990]" country="Brazil" latitude="-24.587223" longLatPrecision="21" longitude="-48.594723" municipality="Iporanga" name="Reserva Bertary" pageId="30" pageNumber="31" stateProvince="Sao Paulo">Reserva Bertary</location>
:
<geoCoordinate id="EB8ED9A3A51AFFFCFCD6F876FBCBABF0" box="[885,1024,1962,1989]" degrees="24" direction="south" minutes="35" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="14" value="-24.587223">24°35’14’’S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A51AFFFCFBAFF876FB6BABF0" box="[1036,1184,1962,1989]" degrees="48" direction="west" minutes="35" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="41" value="-48.594723">48°35’41’’W</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8E05BF64A51AFFFDFF64F812FA8BACE2" blockId="30.[151,1436,1890,2026]" lastBlockId="31.[151,1437,152,1367]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="BCCE6376A51AFFFCFF64F812FE79ABDC" bold="true" box="[199,434,1998,2025]" pageId="30" pageNumber="31">Material examined.</emphasis>
<materialsCitation id="3ED2B539A51AFFFCFE1AF812FA53ABDC" ID-GBIF-Occurrence="2451575470" box="[441,1432,1998,2026]" country="Brazil" latitude="-24.587223" location="Reserva Bertary" longLatPrecision="21" longitude="-48.594723" municipality="Iporanga" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Sao Paulo">
<emphasis id="BCCE6376A51AFFFCFE1AF812FDEFABDC" bold="true" box="[441,548,1998,2025]" pageId="30" pageNumber="31">
<collectingCountry id="F6ADFFF4A51AFFFCFE1AF812FDEFABDC" box="[441,548,1998,2025]" name="Brazil" pageId="30" pageNumber="31">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="BCCE6376A51AFFFCFD97F813FD67ABDC" bold="true" box="[564,684,1998,2025]" italics="true" pageId="30" pageNumber="31">
<collectingRegion id="4C7E7186A51AFFFCFD97F813FD67ABDC" box="[564,684,1998,2025]" country="Brazil" name="Sao Paulo" pageId="30" pageNumber="31">São Paulo</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6E61251EA51AFFFCFD1FF80CFCE9ABDF" box="[700,802,2000,2026]" pageId="30" pageNumber="31">Iporanga</collectingMunicipality>
,
<location id="8B65E9BFA51AFFFCFC8CF80CFC2EABDF" LSID="urn:lsid:plazi:treatment:06130E72A51AFFC2FF34F92BFD5BAE07:8B65E9BFA51AFFFCFC8CF80CFC2EABDF" box="[815,997,2000,2026]" country="Brazil" latitude="-24.587223" longLatPrecision="21" longitude="-48.594723" municipality="Iporanga" name="Reserva Bertary" pageId="30" pageNumber="31" stateProvince="Sao Paulo">Reserva Bertary</location>
(
<geoCoordinate id="EB8ED9A3A51AFFFCFC55F812FB4AABDC" box="[1014,1153,1998,2025]" degrees="24" direction="south" minutes="35" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="14" value="-24.587223">24°35’14’’S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A51AFFFCFB2DF812FAEFABDC" box="[1166,1316,1998,2025]" degrees="48" direction="west" minutes="35" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="41" value="-48.594723">48°35’41’’W</geoCoordinate>
), 07/2016
</materialsCitation>
,
<materialsCitation id="3ED2B539A51BFFFDFF34FF45FC63ACE2" ID-GBIF-Occurrence="2451575477" collectingDate="2015-04-12" collectionCode="IBSP" collectorName="V. Calvanese &amp; A. Silva" latitude="-23.528055" location="Sao Roque" longLatPrecision="21" longitude="-47.114166" municipality="Parque Ecologico da Mata da Camara" pageId="31" pageNumber="32" specimenCode="IBSP 4068, IBSP 4070" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="holotype">
<collectorName id="234FDAB2A51BFFFDFF34FF45FEE2AC86" box="[151,297,153,179]" pageId="31" pageNumber="32">V. Calvanese</collectorName>
&amp;
<collectorName id="234FDAB2A51BFFFDFEE8FF45FE6CAC86" box="[331,423,153,179]" pageId="31" pageNumber="32">A. Silva</collectorName>
col.,
<specimenCount id="98BC74EDA51BFFFDFE46FF45FDC3AC86" box="[485,520,153,179]" pageId="31" pageNumber="32" type="female">1♀</specimenCount>
[
<typeStatus id="510101C6A51BFFFDFDB6FF44FDB6AC86" box="[533,637,152,179]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
] (
<specimenCode id="DE1C171FA51BFFFDFD31FF45FCDAAC81" box="[658,785,153,180]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="32">IBSP 4068</specimenCode>
),
<specimenCount id="98BC74EDA51BFFFDFC87FF45FC8CAC86" box="[804,839,153,179]" pageId="31" pageNumber="32" type="female">4♀</specimenCount>
and
<specimenCount id="98BC74EDA51BFFFDFCDEFF45FC6BAC86" box="[893,928,153,179]" pageId="31" pageNumber="32" type="male">2♂</specimenCount>
[
<typeStatus id="510101C6A51BFFFDFC0DFF44FBD4AC86" box="[942,1055,152,179]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
] (
<specimenCode id="DE1C171FA51BFFFDFB97FF45FB78AC86" box="[1076,1203,153,179]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="32">IBSP 4070</specimenCode>
);
<location id="8B65E9BFA51BFFFDFB65FF45FA88AC86" LSID="urn:lsid:plazi:treatment:06130E72A51AFFC2FF34F92BFD5BAE07:8B65E9BFA51BFFFDFB65FF45FA88AC86" box="[1222,1347,153,179]" latitude="-23.528055" longLatPrecision="21" longitude="-47.114166" municipality="Parque Ecologico da Mata da Camara" name="Sao Roque" pageId="31" pageNumber="32">São Roque</location>
,
<collectingMunicipality id="6E61251EA51BFFFDFAEDFF45FE21ACE2" pageId="31" pageNumber="32">Parque Ecológico da Mata da Câmara</collectingMunicipality>
(
<geoCoordinate id="EB8ED9A3A51BFFFDFE59FF61FD4AACE2" box="[506,641,189,215]" degrees="23" direction="south" minutes="31" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="41" value="-23.528055">23°31’41”S</geoCoordinate>
,
<geoCoordinate id="EB8ED9A3A51BFFFDFD2EFF60FCDAACE2" box="[653,785,188,215]" degrees="47" direction="west" minutes="6" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="51" value="-47.114166">47°6’51”W</geoCoordinate>
),
<date id="FA0499A4A51BFFFDFC85FF60FC63ACE2" box="[806,936,188,215]" pageId="31" pageNumber="32" value="2015-04-12">
<collectingDate id="EA40604CA51BFFFDFC85FF60FC63ACE2" box="[806,936,188,215]" pageId="31" pageNumber="32" value="2015-04-12">12/04/2015</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3ED2B539A51BFFFDFC10FF61FAF7ACE2" ID-GBIF-Occurrence="2451575474" box="[947,1340,189,215]" collectionCode="IBSP" collectorName="V. Calvanese" pageId="31" pageNumber="32" specimenCode="IBSP 4082" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectorName id="234FDAB2A51BFFFDFC10FF61FB8FACE2" box="[947,1092,189,215]" pageId="31" pageNumber="32">V. Calvanese</collectorName>
col.,
<specimenCount id="98BC74EDA51BFFFDFB20FF61FB6DACE2" box="[1155,1190,189,215]" pageId="31" pageNumber="32" type="male">1♂</specimenCount>
(
<specimenCode id="DE1C171FA51BFFFDFB17FF61FAFFACE2" box="[1204,1332,189,215]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="31" pageNumber="32">IBSP 4082</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51BFFFDFF64FF3DFCEBAD2A" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8E05BF64A51BFFFDFF64FF3DFCEBAD2A" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FF3DFE85ACCE" bold="true" box="[199,334,225,251]" pageId="31" pageNumber="32">Etymology.</emphasis>
The epithet is a noun in the genitive case, in honor of biologist Adele Megan M. S. Silva, who collected the majority of individuals sampled for the species.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51BFFFDFF64FEF4FC77AF0E" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8E05BF64A51BFFFDFF64FEF4FE8DAE9E" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FEF4FE89AD76" bold="true" box="[199,322,296,323]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
A species of
<taxonomicName id="49BAC4E7A51BFFFDFE76FEF5FD81AD76" box="[469,586,297,323]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="BCCE6376A51BFFFDFE76FEF5FD81AD76" box="[469,586,297,323]" italics="true" pageId="31" pageNumber="32">Aphilodon</emphasis>
</taxonomicName>
with: around 37–39 leg-bearing segments; known body length in adults
<quantity id="49421281A51BFFFDFAD1FEF5FF0DAD52" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="31" pageNumber="32" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
; cephalic plate length/width ratio ca. 1.2; antenna ca. 2 times as long as cephalic plate; pectinate lamella of mandible with straight denticles; coxosternite of second maxillae not distinctly shorter in middle part; telopodite of second maxillae with article 3 longer than article 2, with 2 subapical setae and 1 apical sensillum; forcipular pretergite not exposed, covered by cephalic plate; forcipule with posterior lateral margin concave; tarsungulum ca. 0.9 times as long as trochanteroprefemur+femur; forcipule with 3+1 denticles; distal denticle of trochanteroprefemur, denticle of femur and denticle of tibia relatively large and denticle of tarsungulum relatively small; all pleurites conspicuous; each coxopleuron with ca. 7–10 coxal pores in adults, all sparse; ultimate presternite exposed only close to lateral margins of metasternite and apparently divided in middle part; metasternite of ultimate leg-bearing segment length/width ratio ca. 0.7; ultimate legs ca. 2.5 times as long as penultimate legs, with tarsus 1 ca. 1.6 times as long as tibia.
</paragraph>
<paragraph id="8E05BF64A51BFFFDFF64FD69FB49AF22" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<taxonomicName id="49BAC4E7A51BFFFDFF64FD69FE87AEFA" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[199,332,693,719]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="meganae" status="sp. nov.">
<emphasis id="BCCE6376A51BFFFDFF64FD69FE87AEFA" box="[199,332,693,719]" italics="true" pageId="31" pageNumber="32">A. meganae</emphasis>
</taxonomicName>
<emphasis id="BCCE6376A51BFFFDFEF7FD69FE5FAEFA" bold="true" box="[340,404,693,719]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A7FDDE0DA51BFFFDFEF7FD69FE5FAEFA" box="[340,404,693,719]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from all species of
<taxonomicName id="49BAC4E7A51BFFFDFD6BFD69FCF6AEFA" box="[712,829,693,719]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="BCCE6376A51BFFFDFD6BFD69FCF6AEFA" box="[712,829,693,719]" italics="true" pageId="31" pageNumber="32">Aphilodon</emphasis>
</taxonomicName>
by having body length &lt;
<quantity id="49421281A51BFFFDFBC0FD69FB55AEFA" box="[1123,1182,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="cm" value="1.0">1 cm</quantity>
, &lt;41 leg-bearing segments (
<figureCitation id="1681A3E1A51BFFFDFF4AFD05FE99AEC6" box="[233,338,729,755]" captionStart="FIGURE 24" captionStartId="32.[151,250,1737,1762]" captionTargetBox="[181,1412,584,1709]" captionTargetId="figure@32.[174,1414,576,1718]" captionTargetPageId="32" captionText="FIGURE 24. A–E, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, terminal part of body, dorsal; E, same, ventral. Scale bars: 2.2 (A), 0.4 (B–C), 0.2 (D–E) mm." figureDoi="http://doi.org/10.5281/zenodo.3542927" httpUri="https://zenodo.org/record/3542927/files/figure.png" pageId="31" pageNumber="32">Fig. 24A</figureCitation>
), telopodite of second maxillae with article 3 longer than article 2 (
<figureCitation id="1681A3E1A51BFFFDFBE3FD05FB63AEC6" box="[1088,1192,729,755]" captionStart="FIGURE 23" captionStartId="31.[151,250,1886,1911]" captionTargetBox="[158,1428,1389,1859]" captionTargetId="figure@31.[151,1436,1381,1866]" captionTargetPageId="31" captionText="FIGURE 23. A–C, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, right telopodite of second maxillae, ventro-lateral; B, forcipular segment, ventral; C, right forcipule, ventral. Abbreviations: PLM: posterior lateral margin of forcipule; T2: article 2 of telopodite of second maxillae; T3: article 3 of telopodite of second maxillae. Scale bars: 0.02 (A), 0.1 (B–C) mm." figureDoi="http://doi.org/10.5281/zenodo.3542925" httpUri="https://zenodo.org/record/3542925/files/figure.png" pageId="31" pageNumber="32">Fig. 23A</figureCitation>
), forcipular pretergite not exposed (
<figureCitation id="1681A3E1A51BFFFDFE8EFD21FE5AAF22" box="[301,401,765,791]" captionStart="FIGURE 24" captionStartId="32.[151,250,1737,1762]" captionTargetBox="[181,1412,584,1709]" captionTargetId="figure@32.[174,1414,576,1718]" captionTargetPageId="32" captionText="FIGURE 24. A–E, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, habitus, ventral; B, anterior part of body, dorsal; C, same, ventral; D, terminal part of body, dorsal; E, same, ventral. Scale bars: 2.2 (A), 0.4 (B–C), 0.2 (D–E) mm." figureDoi="http://doi.org/10.5281/zenodo.3542927" httpUri="https://zenodo.org/record/3542927/files/figure.png" pageId="31" pageNumber="32">Fig. 24B</figureCitation>
) and forcipule with posterior lateral margin concave (
<figureCitation id="1681A3E1A51BFFFDFC4EFD21FBF5AF22" box="[1005,1086,765,791]" captionStart="FIGURE 23" captionStartId="31.[151,250,1886,1911]" captionTargetBox="[158,1428,1389,1859]" captionTargetId="figure@31.[151,1436,1381,1866]" captionTargetPageId="31" captionText="FIGURE 23. A–C, Aphilodon meganae n. sp. Male paratype (IBSP 4070): A, right telopodite of second maxillae, ventro-lateral; B, forcipular segment, ventral; C, right forcipule, ventral. Abbreviations: PLM: posterior lateral margin of forcipule; T2: article 2 of telopodite of second maxillae; T3: article 3 of telopodite of second maxillae. Scale bars: 0.02 (A), 0.1 (B–C) mm." figureDoi="http://doi.org/10.5281/zenodo.3542925" httpUri="https://zenodo.org/record/3542925/files/figure.png" pageId="31" pageNumber="32">Fig. 23</figureCitation>
B–C).
</paragraph>
<paragraph id="8E05BF64A51BFFFDFF64FCFDFC77AF0E" blockId="31.[151,1437,152,1367]" box="[199,956,800,827]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FCFDFE71AF0E" bold="true" box="[199,442,800,827]" pageId="31" pageNumber="32">Recorded specimens</emphasis>
: 8, including
<specimenCount id="98BC74EDA51BFFFDFDF5FCFCFDB2AF0E" box="[598,633,800,827]" pageId="31" pageNumber="32" type="female">5♀</specimenCount>
and
<specimenCount id="98BC74EDA51BFFFDFD13FCFDFD1EAF0E" box="[688,725,801,827]" pageId="31" pageNumber="32" type="male">3♂</specimenCount>
, from two localities.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51BFFFDFF64FC98FCF7AF6A" box="[199,828,836,863]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8E05BF64A51BFFFDFF64FC98FCF7AF6A" blockId="31.[151,1437,152,1367]" box="[199,828,836,863]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FC98FEAAAF6A" bold="true" box="[199,353,836,863]" pageId="31" pageNumber="32">Distribution.</emphasis>
Brazil—São Paulo: Iporanga; São Roque.
</paragraph>
</subSubSection>
<subSubSection id="C6A0ECEFA51BFFC2FF64FCB4FD5BAE07" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
<paragraph id="8E05BF64A51BFFFDFF64FCB4FD13AFB6" blockId="31.[151,1437,152,1367]" box="[199,728,872,900]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FCB4FE2EAFB6" bold="true" box="[199,485,872,899]" pageId="31" pageNumber="32">
Description of
<typeStatus id="510101C6A51BFFFDFED4FCB5FE2BAFB6" box="[375,480,873,899]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
.
</emphasis>
Female (IBSP 4068).
</paragraph>
<paragraph id="8E05BF64A51BFFFDFF64FC52FADCAFDA" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FC52FE82AF92" box="[199,329,909,935]" italics="true" pageId="31" pageNumber="32">Entire body</emphasis>
: length 8, maximum width 0.28.
<emphasis id="BCCE6376A51BFFFDFD1CFC51FCA8AF92" box="[703,867,909,935]" italics="true" pageId="31" pageNumber="32">Cephalic plate</emphasis>
: sub-rectangular, with straight anterior margin and posterior margin rounded, length 0.25 and width 0.20 (length/width ratio 1.25).
<emphasis id="BCCE6376A51BFFFDFB85FC6DFB48AFFE" box="[1062,1155,945,971]" italics="true" pageId="31" pageNumber="32">Antenna</emphasis>
: length 0.5, 2.0 times as long as cephalic plate; AA II–XIII length/width ratio 0.40–0.86; LAA ca. 2.0 times as long as AA XIII.
</paragraph>
<paragraph id="8E05BF64A51BFFFDFF64FC25FDBEA896" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FC25FEEAA826" box="[199,289,1017,1043]" italics="true" pageId="31" pageNumber="32">Clypeus</emphasis>
: 2+2 LS, 1+1 PS, 6+6 SCS.
<emphasis id="BCCE6376A51BFFFDFDDBFC25FD28A826" box="[632,739,1017,1043]" italics="true" pageId="31" pageNumber="32">Mandible</emphasis>
: PL with 14 denticles.
<emphasis id="BCCE6376A51BFFFDFC57FC26FB5FA826" box="[1012,1172,1017,1043]" italics="true" pageId="31" pageNumber="32">First maxillae</emphasis>
: TFM length 0.02 and width 0.02; DA with 1 apical and 1 subapical SL; MPC with 4+4 SL arranged in a row.
<emphasis id="BCCE6376A51BFFFDFBC6FBC1FAEDA802" box="[1125,1318,1053,1079]" italics="true" pageId="31" pageNumber="32">Second maxillae:</emphasis>
4 little SE distributed in a row along the posterior margin of CSM, between the TSM; left TSM length 0.04 (1.8 times as long as TFM) and maximum width (basal margin) 0.02; in both TSM, T3 with 1 apical and 1 subapical SE and 1 apical SL; left T3 ca. 1.6 times as long as left T2.
</paragraph>
<paragraph id="8E05BF64A51BFFFDFF64FB71FCE1A962" blockId="31.[151,1437,152,1367]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF64FB71FE66A8F2" box="[199,429,1197,1223]" italics="true" pageId="31" pageNumber="32">Forcipular segment:</emphasis>
FTE sub-trapezoidal to rectangular, length 0.07 (0.28 times as long as cephalic plate) and maximum width 0.23, with little scattered SE; FPT not exposed; FCX length 0.18 and width 0.21 (length/width ratio 0.85); FRL length 0.16, reaching the anterior margin of cephalic plate; TE with 2 denticles in TFF and
<quantity id="49421281A51BFFFDFA87FB29FA9BA93A" box="[1316,1360,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="31" pageNumber="32" unit="in" value="1.0">1 in</quantity>
TI, all relatively large; FD1 with 1 apical SE; FD2 with 1 apical and 2 subapical SE; FD3 with 1 apical and 2 subapical SE; TA with relatively short DT and 0.91 times as long as TFF.
</paragraph>
<caption id="DAC5EFECA51BFFFDFF34F882FA5AAB8B" ID-DOI="http://doi.org/10.5281/zenodo.3542925" ID-Zenodo-Dep="3542925" httpUri="https://zenodo.org/record/3542925/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1886,1911]" targetBox="[158,1428,1389,1859]" targetPageId="31">
<paragraph id="8E05BF64A51BFFFDFF34F882FA5AAB8B" blockId="31.[151,1437,1886,1983]" pageId="31" pageNumber="32">
<emphasis id="BCCE6376A51BFFFDFF34F882FE93AB42" bold="true" box="[151,344,1886,1911]" pageId="31" pageNumber="32">FIGURE 23. A–C</emphasis>
,
<taxonomicName id="49BAC4E7A51BFFFDFEC5F882FDF9AB43" authority="Calvanese &amp; Brescovit &amp; Bonato, 2019" authorityName="Calvanese &amp; Brescovit &amp; Bonato" authorityYear="2019" box="[358,562,1886,1910]" class="Chilopoda" family="Aphilodontidae" genus="Aphilodon" kingdom="Animalia" order="Geophilomorpha" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="meganae" status="sp. nov.">
<emphasis id="BCCE6376A51BFFFDFEC5F882FDF9AB43" box="[358,562,1886,1910]" italics="true" pageId="31" pageNumber="32">Aphilodon meganae</emphasis>
</taxonomicName>
<emphasis id="BCCE6376A51BFFFDFD9AF883FDB8AB42" bold="true" box="[569,627,1887,1911]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A7FDDE0DA51BFFFDFD9AF883FDB8AB42" box="[569,627,1887,1911]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male paratype (IBSP 4070):
<emphasis id="BCCE6376A51BFFFDFC03F882FC79AB42" bold="true" box="[928,946,1886,1911]" pageId="31" pageNumber="32">A</emphasis>
, right telopodite of second maxillae, ventro-lateral;
<emphasis id="BCCE6376A51BFFFDFF68F85FFF17ABAE" bold="true" box="[203,220,1923,1947]" pageId="31" pageNumber="32">B</emphasis>
, forcipular segment, ventral;
<emphasis id="BCCE6376A51BFFFDFDABF85EFDD1ABAE" bold="true" box="[520,538,1922,1947]" pageId="31" pageNumber="32">C</emphasis>
, right forcipule, ventral.
<emphasis id="BCCE6376A51BFFFDFCBAF85EFC79ABAE" bold="true" box="[793,946,1922,1947]" pageId="31" pageNumber="32">Abbreviations</emphasis>
: PLM: posterior lateral margin of forcipule; T2: article 2 of telopodite of second maxillae; T3: article 3 of telopodite of second maxillae.
<emphasis id="BCCE6376A51BFFFDFBBBF87AFB4CAB8A" bold="true" box="[1048,1159,1958,1983]" pageId="31" pageNumber="32">Scale bars</emphasis>
: 0.02 (A), 0.1 (B–C) mm.
</paragraph>
</caption>
<paragraph id="8E05BF64A524FFC2FF64FF45FA40AD55" blockId="32.[151,1437,153,562]" pageId="32" pageNumber="33">
<emphasis id="BCCE6376A524FFC2FF64FF45FE9DAC86" box="[199,342,153,179]" italics="true" pageId="32" pageNumber="33">Walking legs</emphasis>
: 39 pairs; FL 0.74 times as long as second leg, with procoxae extended to midline of body; leg 19 length 0.29.
<emphasis id="BCCE6376A524FFC2FE86FF60FC61ACE0" box="[293,938,188,214]" italics="true" pageId="32" pageNumber="33">Tergum from the first to penultimate leg-bearing segment</emphasis>
: TE rectangular, with almost straight edges, relatively elongate when compared to other species; TE 15 length 0.11 and width 0.19 (length/width ratio 0.57), with two rows of 6+6 SE; PT elongated, 0.78 times as long as TE, with a row of 5 SE.
<emphasis id="BCCE6376A524FFC2FB88FEDDFE77AD08" italics="true" pageId="32" pageNumber="33">Sternum from the first to penultimate leg-bearing segment</emphasis>
: STF sub-oval, wider than long; other ST more elongated than in other Aphilodontinae; ST 19 length 0.14 and width 0.11 (length/width ratio 1.3); PST well exposed in middle and terminal body region.
</paragraph>
<paragraph id="8E05BF64A524FFC2FF64FEB5FCF0ADDE" blockId="32.[151,1437,153,562]" pageId="32" pageNumber="33">
<emphasis id="BCCE6376A524FFC2FF64FEB5FDD3ADB6" box="[199,536,361,387]" italics="true" pageId="32" pageNumber="33">Ultimate leg-bearing segment</emphasis>
: MTL slightly trapezoidal, length 0.07 and maximum width 0.10 (length/width ratio 0.7); UPS apparently divided in middle part.
<emphasis id="BCCE6376A524FFC2FD6EFE50FCA8AD93" box="[717,867,396,422]" italics="true" pageId="32" pageNumber="33">Ultimate legs</emphasis>
: thin and elongated, length 0.66, 2.5 times as long as penultimate leg; each CXP with 8–9 COP, weakly grouped, smallest ones closest to MTL; tarsus 1 ca. 1.6 times as long as tibia.
<emphasis id="BCCE6376A524FFC2FEEFFE0DFDE1ADDE" box="[332,554,465,491]" italics="true" pageId="32" pageNumber="33">Postpedal segments</emphasis>
: GF with 1+1 distal SE.
</paragraph>
<paragraph id="8E05BF64A524FFC2FF64FE28FD5BAE07" blockId="32.[151,1437,153,562]" pageId="32" pageNumber="33">
<emphasis id="BCCE6376A524FFC2FF64FE28FE57AE3B" bold="true" box="[199,412,499,526]" pageId="32" pageNumber="33">Known variation.</emphasis>
<emphasis id="BCCE6376A524FFC2FE06FE28FE14AE3B" box="[421,479,500,526]" italics="true" pageId="32" pageNumber="33">Male</emphasis>
: GM biarticulated; GM1 longer than wide, with 3 SE in one row; GM2 triangular, tapering to distal region, with 5 scattered SE.
</paragraph>
</subSubSection>
</treatment>
</document>
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