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<document id="4C5D75C9CB15816CE449CEFB24EA5E79" ID-CLB-Dataset="55792" ID-DOI="10.1080/00222933.2013.807948" ID-GBIF-Dataset="585675a5-0353-44d1-9c9a-866b2a0e8205" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198062" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" checkinTime="1628884751635" checkinUser="felipe" docAuthor="Naderloo, Reza" docDate="2013" docId="03CC87CD9428FFD7395EFBE21078118F" docLanguage="en" docName="JNatHist.47.45-46.2835-2848.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.807948" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Chaenostoma java Naderloo 2013, n. sp." docType="treatment" docVersion="9" lastPageNumber="2842" masterDocId="FFF5FFB5942BFFDF3B70FF8B11511252" masterDocTitle="The sentinel crabs of the genus Chaenostoma (Stimpson, 1858) (Crustacea: Brachyura: Macrophthalmidae), with description of a new species and new records" masterLastPageNumber="2848" masterPageNumber="2835" pageNumber="2837" updateTime="1729188380289" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="12BC38F0F2DAA651B1AC81DC0AA3EC09">The sentinel crabs of the genus Chaenostoma (Stimpson, 1858) (Crustacea: Brachyura: Macrophthalmidae), with description of a new species and new records</mods:title>
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<mods:namePart id="9C1BD565C645272BD1DAF229CD701CE0">Naderloo, Reza</mods:namePart>
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<mods:title id="3962E60337EBE99CAF16FFF4389337E7">Journal of Natural History</mods:title>
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<mods:date id="37F1FBFCCC157EEE654E4B32B5E304C0">2013</mods:date>
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<treatment id="03CC87CD9428FFD7395EFBE21078118F" ID-DOI="http://doi.org/10.5281/zenodo.5198060" ID-GBIF-Taxon="194873727" ID-Zenodo-Dep="5198060" LSID="urn:lsid:plazi:treatment:03CC87CD9428FFD7395EFBE21078118F" httpUri="http://treatment.plazi.org/id/03CC87CD9428FFD7395EFBE21078118F" lastPageId="8" lastPageNumber="2842" pageId="3" pageNumber="2837">
<subSubSection id="C37F65509428FFDC395EFBE2126816D3" box="[558,825,1129,1153]" pageId="3" pageNumber="2837" type="nomenclature">
<paragraph id="8BDA36DB9428FFDC395EFBE2126816D3" blockId="3.[558,825,1129,1185]" box="[558,825,1129,1153]" pageId="3" pageNumber="2837">
<heading id="D09281B79428FFDC395EFBE2126816D3" box="[558,825,1129,1153]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2837" reason="2">
<taxonomicName id="4C654D589428FFDC395EFBE213A616D3" ID-CoL="T9R3" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[558,759,1129,1153]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="2837" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC99428FFDC395EFBE213A616D3" bold="true" box="[558,759,1129,1153]" italics="true" pageId="3" pageNumber="2837">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B29428FFDC398EFBE1126816D3" box="[766,825,1130,1153]" pageId="3" pageNumber="2837" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C37F65509428FFDC3945FB02126316F3" box="[565,818,1161,1185]" pageId="3" pageNumber="2837" type="description">
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(
<figureCitation id="135E2A5E9428FFDC394DFB02138816F2" box="[573,729,1161,1185]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="3" pageNumber="2837">Figures 1AF</figureCitation>
,
<figureCitation id="135E2A5E9428FFDC3994FB02127B16F3" box="[740,810,1161,1185]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="3" pageNumber="2837">2AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F65509428FFDC3BA3FB6E127A17E6" pageId="3" pageNumber="2837" type="materials_examined">
<paragraph id="8BDA36DB9428FFDC3BA3FB6E10F416AF" blockId="3.[211,1172,1253,1349]" box="[211,421,1253,1277]" pageId="3" pageNumber="2837">
<heading id="D09281B79428FFDC3BA3FB6E10F416AF" box="[211,421,1253,1277]" fontSize="10" level="3" pageId="3" pageNumber="2837" reason="5">
<emphasis id="B911EAC99428FFDC3BA3FB6E10F416AF" box="[211,421,1253,1277]" italics="true" pageId="3" pageNumber="2837">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB9428FFDC3BA3FA8610131717" blockId="3.[211,1172,1253,1349]" pageId="3" pageNumber="2837">
<materialsCitation id="3B0D3C869428FFDC3BA3FA8610131717" ID-GBIF-Occurrence="3757902304" collectingDate="1897-07" collectionCode="MHNG" collectorName="Zehntner." location="Rosoeki" pageId="3" pageNumber="2837" specimenCount="1" specimenCount-male="1" stateProvince="Java" typeStatus="holotype">
<emphasis id="B911EAC99428FFDC3BA3FA86106B1777" box="[211,314,1293,1317]" italics="true" pageId="3" pageNumber="2837">
<typeStatus id="54DE88799428FFDC3BA3FA86106B1777" box="[211,314,1293,1317]" pageId="3" pageNumber="2837" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D63FD529428FFDC3A39FA86103A1777" box="[329,363,1293,1317]" pageId="3" pageNumber="2837" type="male">1♂</specimenCount>
(CL
<quantity id="4C9D9B3E9428FFDC3AD8FA8613471777" box="[424,534,1293,1317]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="2837" unit="mm" value="9.6">9. 60 mm</quantity>
, CB
<quantity id="4C9D9B3E9428FFDC393EFA8613EE1777" box="[590,703,1293,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.209" pageId="3" pageNumber="2837" unit="mm" value="12.09">12.09 mm</quantity>
) (
<collectionCode id="ED74AE1E9428FFDC39A6FA8612671777" box="[726,822,1293,1317]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="2837" type="Museum">MHNG</collectionCode>
),
<location id="8EBA60009428FFDC383AFA8612F91777" LSID="urn:lsid:plazi:treatment:03CC87CD9428FFD7395EFBE21078118F:8EBA60009428FFDC383AFA8612F91777" box="[842,936,1293,1317]" name="Rosoeki" pageId="3" pageNumber="2837" stateProvince="Java">Rosoeki</location>
,
<collectingRegion id="49A1F8399428FFDC38C4FA8512B61777" box="[948,999,1294,1317]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="2837">Java</collectingRegion>
,
<date id="FFDB101B9428FFDC3883FA8615311777" box="[1011,1120,1293,1317]" pageId="3" pageNumber="2837" value="1897-07">
<collectingDate id="EF9FE9F39428FFDC3883FA8615311777" box="[1011,1120,1293,1317]" pageId="3" pageNumber="2837" value="1897-07">July 1897</collectingDate>
</date>
, leg.
<collectorName id="2690530D9428FFDC3BA3FAA610131717" box="[211,322,1325,1349]" pageId="3" pageNumber="2837">Zehntner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BDA36DB9428FFDC3BA3FAF5127A17E6" blockId="3.[211,1171,1404,1460]" pageId="3" pageNumber="2837">
<materialsCitation id="3B0D3C869428FFDC3BA3FAF5127617E6" ID-GBIF-Occurrence="3757902301" collectingDate="1897-07" collectionCode="MHNG" collectorName="Zehntner." location="Rosoeki" pageId="3" pageNumber="2837" specimenCode="CL 7.94, CB 10.07" specimenCount="1" specimenCount-female="4" specimenCount-male="1" stateProvince="Jawa Timur" typeStatus="paratype">
<emphasis id="B911EAC99428FFDC3BA3FAF5101117C7" box="[211,320,1406,1429]" italics="true" pageId="3" pageNumber="2837">
<typeStatus id="54DE88799428FFDC3BA3FAF5101117C7" box="[211,320,1406,1429]" pageId="3" pageNumber="2837" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="9D63FD529428FFDC3A26FAF7102917C7" box="[342,376,1404,1429]" pageId="3" pageNumber="2837" type="male">1♂</specimenCount>
(CL
<quantity id="4C9D9B3E9428FFDC3AB3FAF7137E17C6" box="[451,559,1404,1428]" metricMagnitude="-3" metricUnit="m" metricValue="8.97" pageId="3" pageNumber="2837" unit="mm" value="8.97">8.97 mm</quantity>
, CB
<quantity id="4C9D9B3E9428FFDC3904FAF713BE17C6" box="[628,751,1404,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.124" pageId="3" pageNumber="2837" unit="mm" value="11.24">11.24 mm</quantity>
), 4♀ ovig. (CL 7.94
<quantity id="4C9D9B3E9428FFDC3881FAF7150D17C6" box="[1009,1116,1404,1428]" metricMagnitude="-3" metricUnit="m" metricValue="5.07" pageId="3" pageNumber="2837" unit="mm" value="5.07">5.07 mm</quantity>
, CB 10.07
<quantity id="4C9D9B3E9428FFDC3A6DFA1710D117E6" box="[285,384,1436,1460]" metricMagnitude="-3" metricUnit="m" metricValue="6.89" pageId="3" pageNumber="2837" unit="mm" value="6.89">6.89 mm</quantity>
) (
<collectionCode id="ED74AE1E9428FFDC3AE6FA1710A717E6" box="[406,502,1436,1460]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="2837" type="Museum">MHNG</collectionCode>
), locality same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F65509428FFD83BA3FA71156911F4" lastPageId="7" lastPageNumber="2841" pageId="3" pageNumber="2837" type="description">
<paragraph id="8BDA36DB9428FFDC3BA3FA7110031443" blockId="3.[211,1172,1530,1720]" box="[211,338,1530,1553]" pageId="3" pageNumber="2837">
<heading id="D09281B79428FFDC3BA3FA7110031443" box="[211,338,1530,1553]" fontSize="10" level="3" pageId="3" pageNumber="2837" reason="5">
<emphasis id="B911EAC99428FFDC3BA3FA7110031443" box="[211,338,1530,1553]" italics="true" pageId="3" pageNumber="2837">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB9428FFDA3BA3F9AA13E417E9" blockId="3.[211,1172,1530,1720]" lastBlockId="5.[211,1172,1347,1690]" lastPageId="5" lastPageNumber="2839" pageId="3" pageNumber="2837">
Carapace sub-rectangular (
<figureCitation id="135E2A5E9428FFDC397AF9AA13D1146B" box="[522,640,1569,1593]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="3" pageNumber="2837">Figure 1A</figureCitation>
), about 1.21.3 times as broad as long, maximum width on posterior part of posterolateral margin; posterior surface with small granules scarcely on lateral regions, invisible with naked eyes; long setae on lateral regions, continuous to lateral margin. Regions weekly defined; furrows of gastric region distinct on posterior portion; cardiac region slightly elevated, intestinal region depressed. Frontal region (
<figureCitation id="135E2A5E942EFFDA3974FAC813291709" box="[516,632,1347,1371]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="5" pageNumber="2839">Figure 1A</figureCitation>
) downflexed, small granules over it; front wide, about 1/4 as wide as carapace width in anterior part, very slightly constricted medially; anterior edge slightly concave medially, with long setae, lateral margins of front nearly round; frontal furrow faint, shallow.
</paragraph>
<caption id="DF1A6653942FFFDB3BFFFA2610ED1449" ID-DOI="http://doi.org/10.5281/zenodo.5198064" ID-Zenodo-Dep="5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="4" pageNumber="2838" startId="4.[143,210,1453,1475]" targetBox="[144,1096,160,1420]" targetPageId="4">
<paragraph id="8BDA36DB942FFFDB3BFFFA2610ED1449" blockId="4.[143,1104,1453,1563]" pageId="4" pageNumber="2838">
Figure 1.
<taxonomicName id="4C654D58942FFFDB3B87FA2610FD1791" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[247,428,1453,1475]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="2838" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC9942FFFDB3B87FA2610FD1791" box="[247,428,1453,1475]" italics="true" pageId="4" pageNumber="2838">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B2942FFFDB3AC3FA2510B81791" box="[435,489,1454,1475]" pageId="4" pageNumber="2838" rank="species">n. sp.</taxonomicNameLabel>
holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face).
</paragraph>
</caption>
<caption id="DF1A6653942EFFDA3BA3FB1A10CC16AD" ID-DOI="http://doi.org/10.5281/zenodo.5198066" ID-Zenodo-Dep="5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="5" pageNumber="2839" startId="5.[211,278,1169,1191]" targetBox="[212,1172,167,1136]" targetPageId="5">
<paragraph id="8BDA36DB942EFFDA3BA3FB1A10CC16AD" blockId="5.[211,1172,1169,1279]" pageId="5" pageNumber="2839">
Figure 2.
<taxonomicName id="4C654D58942EFFDA3A30FB1A10AB16F5" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[320,506,1169,1191]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="2839" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC9942EFFDA3A30FB1A10AB16F5" box="[320,506,1169,1191]" italics="true" pageId="5" pageNumber="2839">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B2942EFFDA3975FB19131116F5" box="[517,576,1170,1191]" pageId="5" pageNumber="2839" rank="species">n. sp.</taxonomicNameLabel>
holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen.
</paragraph>
</caption>
<paragraph id="8BDA36DB942EFFDA3B8EFA4912171468" blockId="5.[211,1172,1347,1690]" pageId="5" pageNumber="2839">
Eyestalks (
<figureCitation id="135E2A5E942EFFDA3A0BFA4910A11788" box="[379,496,1474,1498]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="5" pageNumber="2839">Figure 1A</figureCitation>
) relatively stout, short, reaching to end of exorbital angle. Upper orbital margin regularly granulate, granules very small, round-tipped, slightly directed outwards, slightly larger laterally. Lower margin tuberculate, tubercles nearly triangular, directed inwards, getting larger in inner part.
</paragraph>
<paragraph id="8BDA36DB942EFFD93B8EF9C910BE134D" blockId="5.[211,1172,1347,1690]" lastBlockId="6.[143,1104,167,1658]" lastPageId="6" lastPageNumber="2840" pageId="5" pageNumber="2839">
Lateral margin (
<figureCitation id="135E2A5E942EFFDA3AC9F9C9137C1408" box="[441,557,1602,1626]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="5" pageNumber="2839">Figure 1A</figureCitation>
) with three teeth (including exorbital angle); first one (exorbital angle) wide triangle to relatively subquadrate, directed forward, making a wide concavity with upper orbital margin; posterior margin moderately arched, granulate; second longer than first one, lower than it, outer margin nearly straight, granulate; third very small, usually distinguishable, indicated by large granules; posterolateral margin relatively straight, diverging posteriorly, with small round granules, wide-based, long setae along it.
</paragraph>
<paragraph id="8BDA36DB942DFFD93BCAFEAC11B6104C" blockId="6.[143,1104,167,1658]" pageId="6" pageNumber="2840">Third maxillipeds with large gap between. Ischium long, about 1.8 times as long as wide, slightly shorter than two times as long as merus, inner margin distally convex, with long setae along it, outer margin nearly straight, with short setae; circular pattern of long setae on proximal portion of outer surface, long setae irregularly along distal margin, short setae scarcely on outer surface. Merus quadrate, inner margin with long setae; outer margin nearly straight, arched proximally, long setae along distal part, outer surface widely depressed near inner margin, narrow depression near along outer margin.</paragraph>
<paragraph id="8BDA36DB942DFFD93BCAFDAD10EE100C" blockId="6.[143,1104,167,1658]" pageId="6" pageNumber="2840">Posteromedian margin of epistome straight, median part of anterior buccal cavity smooth, without any ridge.</paragraph>
<paragraph id="8BDA36DB942DFFD93BCAFDED13C616A9" blockId="6.[143,1104,167,1658]" pageId="6" pageNumber="2840">
Chelipeds (
<figureCitation id="135E2A5E942DFFD93A4BFDED1087102C" box="[315,470,614,638]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="6" pageNumber="2840">Figure 1C, D</figureCitation>
) nearly equal. Ischium denticulate on anterior margin, with long setae. Merus (
<figureCitation id="135E2A5E942DFFD93AEEFD0D135E10CC" box="[414,527,646,670]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="6" pageNumber="2840">Figure 1B</figureCitation>
) upper surface smooth, slightly elevated near inner margin, without chitinous plectrum; outer surface with transverse ridges of very small granules, sparsely covered with short setae; inner surface smooth, densely covered with long setae on distal two third: upper inner margin with small denticles, getting slightly larger distally, densely covered with long setae, setae continuous to anterior surface; inner lower margin with large triangular tubercles, larger distally, covered with scarce setae; outer margin denticulate, long setae scarcely along it. Carpus finely granulate on outer surface; margins with small denticles, some large denticles medially on inner margin, long setae along outer, inner, lower distal margins. Palm (
<figureCitation id="135E2A5E942DFFD938E4FC0E155811CF" box="[916,1033,901,925]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="6" pageNumber="2840">Figure 1C</figureCitation>
) relatively swollen, about 1.2 times as long as high; outer surface (
<figureCitation id="135E2A5E942DFFD9384AFC2E12FA11EF" box="[826,939,933,957]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="6" pageNumber="2840">Figure 1C</figureCitation>
) smooth, with microscopic granules, slightly larger proximally, longitudinal granular ridge on lower portion, running from near tip of immovable finger backward, subparallel to lower margin; lower margin nearly straight, minutely granular, not deflexed near immovable finger; upper margin slightly arched, finely granular; inner surface (
<figureCitation id="135E2A5E942DFFD938E5FBAF155D166E" box="[917,1036,1060,1084]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="6" pageNumber="2840">Figure 1D</figureCitation>
) with small granules on proximal portion, mostly covered with long setae, setal patch continuous along inner surface of fingers. Movable finger curved downward, curved inward distally, cutting edge with subproximal differentiated tooth, large, small denticles distal to large one. Immovable finger short, smooth proximally, without large differentiated tooth, teeth along proximal two-thirds, getting larger distally; large round gap between fingers, tip of fingers spooned, with long setae.
</paragraph>
<paragraph id="8BDA36DB942DFFD93BCAFA88127B1789" blockId="6.[143,1104,167,1658]" pageId="6" pageNumber="2840">
Female chelipeds (
<figureCitation id="135E2A5E942DFFD93AF7FA8810A91749" box="[391,504,1283,1307]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="6" pageNumber="2840">Figure 2A</figureCitation>
) small; palm nearly flat, inferior ridge prominent, parallel to lower margin, gently curving medially, granular, granules small distally; short longitudinal row of granules on middle portion, depressed above this ridge; upper margin granular, with small granules; inner surface of palm smooth, row of long setae near upper margin. Fingers relatively stout, movable finger distinctly longer than palm, long setae along inner surface, longitudinal ridge along outer surface, cutting edges of fingers smooth, chitinous ridge comb-shaped proximally.
</paragraph>
<paragraph id="8BDA36DB942DFFD83BCAFA6813CD130D" blockId="6.[143,1104,167,1658]" lastBlockId="7.[211,1172,167,797]" lastPageId="7" lastPageNumber="2841" pageId="6" pageNumber="2840">Walking legs medium-sized, second, third legs largest, nearly in same length, fourth leg smallest, long setae scarcely along anterior margin of segments, except dactylus. Merus posterior surface smooth; anterior, posterior margins serrate, merus of second and third legs with subdistal tooth on anterior margin, merus of third leg about three times as long as wide. Carpus serrate on anterior and posterior margins; carpus of third leg slightly shorter than propodus. Propodus finely serrate on anterior and posterior margins, slightly longer than dactylus, that of last leg about as long as dactylus. Dactylus with scarce short setae on proximal portion, with six longitudinal smooth ridges; dense setal patch on anterior, posterior margins of daclytus and propodus of first leg; setal patch just on proximal parts of dactylus and propodus of second and third legs; fourth leg without setal patch.</paragraph>
<paragraph id="8BDA36DB942CFFD83B8EFEEC12E9138C" blockId="7.[211,1172,167,797]" pageId="7" pageNumber="2841">
Male abdomen long; segments 3, 4 nearly in same length, segment 5 distinctly longer than segments 3, 4, nearly as long as segment 6; segment 6 (
<figureCitation id="135E2A5E942CFFD838ADFE0C151E13CD" box="[989,1103,391,415]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="7" pageNumber="2841">Figure 2B</figureCitation>
) with lateral margins arched, very slightly swollen proximally, converging distally; telson slightly shorter than segment 6, margins arched, apically rounded.
</paragraph>
<paragraph id="8BDA36DB942CFFD83B8EFE6D12B2100C" blockId="7.[211,1172,167,797]" pageId="7" pageNumber="2841">
G1 gently curved medially (
<figureCitation id="135E2A5E942CFFD83942FE6D13F013AC" box="[562,673,486,510]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="7" pageNumber="2841">Figure 1E</figureCitation>
); apical chitinous process short, directed laterally at nearly 60
<superScript id="7C109B93942CFFD83AE6FD8910CE1041" attach="left" box="[406,415,514,531]" fontSize="7" pageId="7" pageNumber="2841"></superScript>
(
<figureCitation id="135E2A5E942CFFD83ADFFD8D1313104C" box="[431,578,518,542]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="7" pageNumber="2841">Figure 1F, G</figureCitation>
), lateral surface of apical process distinctly depressed, concave on apical surface; long setae around apical process, long feather-shaped setae along ventral margin, short feather-shaped setae along mesial margin.
</paragraph>
<paragraph id="8BDA36DB942CFFD83B8EFDED132610EF" blockId="7.[211,1172,167,797]" pageId="7" pageNumber="2841">
Female abdomen round, segment 6 (
<figureCitation id="135E2A5E942CFFD839E9FDED1258102C" box="[665,777,614,638]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="7" pageNumber="2841">Figure 2E</figureCitation>
) distinctly longer than telson, with arched lateral margins, telson distinctly wider than half as time as segment 6, slightly more than four times as wide as long.
</paragraph>
<paragraph id="8BDA36DB942CFFD83B8EFD4E1386114F" blockId="7.[211,1172,167,797]" pageId="7" pageNumber="2841">
Female gonopore (
<figureCitation id="135E2A5E942CFFD83AACFD4E1303108F" box="[476,594,709,733]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="7" pageNumber="2841">Figure 2C</figureCitation>
) small, with outer margin nearly straight, nearly parallel to median line of sternum; operculum small (
<figureCitation id="135E2A5E942CFFD8384DFD6E12E210AF" box="[829,947,741,765]" captionStart="Figure 2" captionStartId="5.[211,278,1169,1191]" captionTargetBox="[212,1172,167,1136]" captionTargetId="figure-5@5.[212,1172,167,1136]" captionTargetPageId="5" captionText="Figure 2. Chaenostoma java n. sp. holotype (NHMG) (B) and female paratype (A, CE). (A) Female right cheliped (outer face); (B) penultimate segment and telson of male abdomen; (C) female gonopore; (D) female gonopore in closer view; (E) penultimate segment and telson of female abdomen." figureDoi="http://doi.org/10.5281/zenodo.5198066" httpUri="https://zenodo.org/record/5198066/files/figure.png" pageId="7" pageNumber="2841">Figure 2D</figureCitation>
), directed forwards laterally; margins of opening slightly elevated.
</paragraph>
<paragraph id="8BDA36DB942CFFD83BA3FCED107B112C" blockId="7.[211,1080,870,934]" box="[211,298,870,894]" pageId="7" pageNumber="2841">
<heading id="D09281B7942CFFD83BA3FCED107B112C" box="[211,298,870,894]" fontSize="10" level="3" pageId="7" pageNumber="2841" reason="5">
<emphasis id="B911EAC9942CFFD83BA3FCED107B112C" box="[211,298,870,894]" italics="true" pageId="7" pageNumber="2841">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB942CFFD83BA3FC05156911F4" blockId="7.[211,1080,870,934]" box="[211,1080,910,934]" pageId="7" pageNumber="2841">The specimens were collected from the rocky intertidal, under small boulders.</paragraph>
</subSubSection>
<subSubSection id="C37F6550942CFFD83BA3FC641072161D" pageId="7" pageNumber="2841" type="distribution">
<paragraph id="8BDA36DB942CFFD83BA3FC6410091655" blockId="7.[211,1171,1007,1103]" box="[211,344,1007,1031]" pageId="7" pageNumber="2841">
<heading id="D09281B7942CFFD83BA3FC6410091655" box="[211,344,1007,1031]" fontSize="10" level="3" pageId="7" pageNumber="2841" reason="5">
<emphasis id="B911EAC9942CFFD83BA3FC6410091655" box="[211,344,1007,1031]" italics="true" pageId="7" pageNumber="2841">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB942CFFD83BA3FB9C1072161D" blockId="7.[211,1171,1007,1103]" pageId="7" pageNumber="2841">
Presently only known from its
<typeStatus id="54DE8879942CFFD8394DFB93133C1662" box="[573,621,1048,1072]" pageId="7" pageNumber="2841">type</typeStatus>
locality,
<collectingRegion id="49A1F839942CFFD839ABFB93125A167D" box="[731,779,1048,1071]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="2841">Java</collectingRegion>
(
<collectingCountry id="F372764B942CFFD8386CFB9C12DF167D" box="[796,910,1047,1071]" name="Indonesia" pageId="7" pageNumber="2841">Indonesia</collectingCountry>
) in the
<location id="8EBA6000942CFFD8389EFB93104E161D" LSID="urn:lsid:plazi:treatment:03CC87CD9428FFD7395EFBE21078118F:8EBA6000942CFFD8389EFB93104E161D" country="Indonesia" name="eastern Indian Ocean" pageId="7" pageNumber="2841" stateProvince="Jawa Timur">eastern Indian Ocean</location>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6550942CFFD83BA3FB1312DB168A" box="[211,906,1176,1240]" pageId="7" pageNumber="2841" type="etymology">
<paragraph id="8BDA36DB942CFFD83BA3FB13101A16E2" blockId="7.[211,906,1176,1240]" box="[211,331,1176,1200]" pageId="7" pageNumber="2841">
<heading id="D09281B7942CFFD83BA3FB13101A16E2" box="[211,331,1176,1200]" fontSize="10" level="3" pageId="7" pageNumber="2841" reason="5">
<emphasis id="B911EAC9942CFFD83BA3FB13101A16E2" box="[211,331,1176,1200]" italics="true" pageId="7" pageNumber="2841">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB942CFFD83BA3FB4B12DB168A" blockId="7.[211,906,1176,1240]" box="[211,906,1216,1240]" pageId="7" pageNumber="2841">
The species is named after the
<typeStatus id="54DE8879942CFFD8395AFB4B130B168A" box="[554,602,1216,1240]" pageId="7" pageNumber="2841">type</typeStatus>
locality,
<collectingRegion id="49A1F839942CFFD839B0FB4B13A11685" box="[704,752,1216,1239]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="2841">Java</collectingRegion>
in
<collectingCountry id="F372764B942CFFD83863FB4B12D7168A" box="[787,902,1216,1240]" name="Indonesia" pageId="7" pageNumber="2841">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6550942CFFD73BA3FAAA1078118F" lastPageId="8" lastPageNumber="2842" pageId="7" pageNumber="2841" type="discussion">
<paragraph id="8BDA36DB942CFFD83BA3FAAA1064176B" blockId="7.[211,1172,1313,1664]" box="[211,309,1313,1337]" pageId="7" pageNumber="2841">
<heading id="D09281B7942CFFD83BA3FAAA1064176B" box="[211,309,1313,1337]" fontSize="10" level="3" pageId="7" pageNumber="2841" reason="5">
<emphasis id="B911EAC9942CFFD83BA3FAAA1064176B" box="[211,309,1313,1337]" italics="true" pageId="7" pageNumber="2841">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36DB942CFFD73BA3FAC21184106C" blockId="7.[211,1172,1313,1664]" lastBlockId="8.[143,1104,167,989]" lastPageId="8" lastPageNumber="2842" pageId="7" pageNumber="2841">
<taxonomicName id="4C654D58942CFFD83BA3FAC210F21732" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[211,419,1353,1377]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="2841" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC9942CFFD83BA3FAC210F21732" box="[211,419,1353,1377]" italics="true" pageId="7" pageNumber="2841">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B2942CFFD83ADFFAC210A11732" box="[431,496,1353,1376]" pageId="7" pageNumber="2841" rank="species">n. sp.</taxonomicNameLabel>
is mainly characterized by a blunt exorbital angle, nondeflexed male cheliped, a dense setal patch covering the inner surface of the palm and fingers of the male chelipeds, a G1 with a short rounded apical process, and a female gonopore with a small operculum directed forwards. The non-deflexed male chelipeds and the morphology of the male G1 distinguish the new species from
<taxonomicName id="4C654D58942CFFD83BA3FA6210A51452" authority="(Audouin, 1826)" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[211,500,1512,1536]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="2841" phylum="Arthropoda" rank="species" species="boscii">
<emphasis id="B911EAC9942CFFD83BA3FA6211B71452" box="[211,230,1513,1536]" italics="true" pageId="7" pageNumber="2841">C</emphasis>
.
<emphasis id="B911EAC9942CFFD83B85FA6310651452" box="[245,308,1512,1536]" italics="true" pageId="7" pageNumber="2841">boscii</emphasis>
(Audouin, 1826)
</taxonomicName>
and the widely distributed species
<taxonomicName id="4C654D58942CFFD83808FA6210FB1472" authority="(Naderloo and Turkay, 2011)" baseAuthorityName="Naderloo and Turkay" baseAuthorityYear="2011" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="2841" phylum="Arthropoda" rank="species" species="sinuspersici">
<emphasis id="B911EAC9942CFFD83808FA6212DA1452" box="[888,907,1513,1536]" italics="true" pageId="7" pageNumber="2841">C</emphasis>
.
<emphasis id="B911EAC9942CFFD838EBFA6215491452" box="[923,1048,1513,1536]" italics="true" pageId="7" pageNumber="2841">sinuspersici</emphasis>
(Naderloo and Türkay, 2011)
</taxonomicName>
. Male chelipeds of the last two species are more or less deflexed downwards near the base of the immovable finger. The G1 of
<taxonomicName id="4C654D58942CFFD838C6F9A215331412" baseAuthorityName="Naderloo and Turkay" baseAuthorityYear="2011" box="[950,1122,1576,1600]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="2841" phylum="Arthropoda" rank="species" species="sinuspersici">
<emphasis id="B911EAC9942CFFD838C6F9A212811412" box="[950,976,1577,1600]" italics="true" pageId="7" pageNumber="2841">M</emphasis>
.
<emphasis id="B911EAC9942CFFD83895F9A215331412" box="[997,1122,1577,1600]" italics="true" pageId="7" pageNumber="2841">sinuspersici</emphasis>
</taxonomicName>
has the apical process of G1 distinctly mesiolaterally inclined, strongly emarginated, but the apical process of G1 of
<taxonomicName id="4C654D58942CFFD8396BF9E313D914D2" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[539,648,1640,1664]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="2841" phylum="Arthropoda" rank="species" species="boscii">
<emphasis id="B911EAC9942CFFD8396BF9E31364142D" box="[539,565,1640,1663]" italics="true" pageId="7" pageNumber="2841">M</emphasis>
.
<emphasis id="B911EAC9942CFFD83939F9E313D914D2" box="[585,648,1640,1664]" italics="true" pageId="7" pageNumber="2841">boscii</emphasis>
</taxonomicName>
is plate-shaped, directed obliquely in respect to stem (see Naderloo &amp; Türkay 2011, figure 3). The relatively smooth dorsal surface of the carapace of
<taxonomicName id="4C654D589423FFD73AEAFF4310BA128D" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[410,491,200,223]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC99423FFD73AEAFF4310FC128D" box="[410,429,200,223]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD73ACEFF4310BA128D" box="[446,491,200,223]" italics="true" pageId="8" pageNumber="2842">java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B29423FFD73A83FF43137E128D" box="[499,559,200,223]" pageId="8" pageNumber="2842" rank="species">n. sp.</taxonomicNameLabel>
is rather similar to that of
<taxonomicName id="4C654D589423FFD73816FF43101D12AD" authority="(Poupin and Bouchard, 2010)" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD73816FF431228128D" box="[870,889,200,223]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD738F9FF4312EA12B2" box="[905,955,200,224]" italics="true" pageId="8" pageNumber="2842">lisae</emphasis>
(
<bibRefCitation id="EFF44B2A9423FFD738BBFF4C101712AD" author="Poupin J &amp; Bouchard JM" pageId="8" pageNumber="2842" pagination="61 - 67" refId="ref6483" refString="Poupin J, Bouchard JM. 2010. A new dwarf sentinel crab from Mayotte Island, western Indian Ocean (Decapoda: Brachyura: Macrophthalmidae). Zootaxa. 2501: 61 - 67." type="journal article" year="2010">Poupin and Bouchard, 2010</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C654D589423FFD73AFAFF63138012AD" authority="(Miers, 1884)" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[394,721,231,255]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B911EAC99423FFD73AFAFF6310CC12AD" box="[394,413,232,255]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD73ADEFF6C137D12AD" box="[430,556,231,255]" italics="true" pageId="8" pageNumber="2842">punctulatus</emphasis>
(
<bibRefCitation id="EFF44B2A9423FFD7394EFF6C139812AD" author="Miers EJ" box="[574,713,231,255]" pageId="8" pageNumber="2842" pagination="18 - 32" refId="ref6294" refString="Miers EJ. 1884. Crustacea. In: Report on the zoological collections made in the Indo- Pacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part I. The collections from Melanesia. London; p. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
)
</taxonomicName>
. The latter has been described as having a completely smooth dorsal carapace surface (see
<bibRefCitation id="EFF44B2A9423FFD73859FE8C12FB134D" author="Miers EJ" box="[809,938,263,287]" pageId="8" pageNumber="2842" pagination="18 - 32" refId="ref6294" refString="Miers EJ. 1884. Crustacea. In: Report on the zoological collections made in the Indo- Pacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part I. The collections from Melanesia. London; p. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers 1884</bibRefCitation>
;
<bibRefCitation id="EFF44B2A9423FFD738CBFE831519134D" author="Barnes RSK" box="[955,1096,263,287]" pageId="8" pageNumber="2842" pagination="195 - 262" refId="ref5974" refString="Barnes RSK. 1967. The Macrophthalminae of Australasia; with a review of the evolution and morphological diversity of the type genus Macrophthalmus. Trans Zool Soc Lond. 31: 195 - 262." type="journal article" year="1967">Barnes 1967</bibRefCitation>
, p. 229), whereas
<taxonomicName id="4C654D589423FFD73A3DFEA310CC136D" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[333,413,296,319]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC99423FFD73A3DFEA31031136D" box="[333,352,296,319]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD73A00FEA310CC136D" box="[368,413,296,319]" italics="true" pageId="8" pageNumber="2842">java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B29423FFD73AD5FEA310B3136D" box="[421,482,296,319]" pageId="8" pageNumber="2842" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C654D589423FFD7396CFEA3133E136D" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" box="[540,623,295,319]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD7396CFEA3133E136D" box="[540,623,295,319]" italics="true" pageId="8" pageNumber="2842">C. lisae</emphasis>
</taxonomicName>
have few scattered granules on the frontal and the lateral regions of the carapace. It should be noted that
<bibRefCitation id="EFF44B2A9423FFD73826FECC128C130D" author="Miers EJ" box="[854,989,327,351]" pageId="8" pageNumber="2842" pagination="18 - 32" refId="ref6294" refString="Miers EJ. 1884. Crustacea. In: Report on the zoological collections made in the Indo- Pacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part I. The collections from Melanesia. London; p. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers (1884</bibRefCitation>
, pl. XXV, figure A) described
<taxonomicName id="4C654D589423FFD73A01FEEC1340132D" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[369,529,359,383]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B911EAC99423FFD73A01FEEC1340132D" box="[369,529,359,383]" italics="true" pageId="8" pageNumber="2842">C. punctulatus</emphasis>
</taxonomicName>
based on a single female from Port Jackson,
<collectingRegion id="49A1F8399423FFD73F6BFEE3107113CD" country="Australia" name="New South Wales" pageId="8" pageNumber="2842">New South Wales</collectingRegion>
,
<collectingCountry id="F372764B9423FFD73A5DFE0C10C813CD" box="[301,409,391,415]" name="Australia" pageId="8" pageNumber="2842">Australia</collectingCountry>
. Miers mentioned a large differentiated tooth on the cutting edge of the immovable finger and a smaller one on the cutting edge of the movable finger, which are clearly seen in his drawings (
<bibRefCitation id="EFF44B2A9423FFD739E3FE4D1243138C" author="Miers EJ" box="[659,786,454,478]" pageId="8" pageNumber="2842" pagination="18 - 32" refId="ref6294" refString="Miers EJ. 1884. Crustacea. In: Report on the zoological collections made in the Indo- Pacific Ocean during the voyage of H. M. S. &quot; Alert &quot; 1881 - 82. Part I. The collections from Melanesia. London; p. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers 1884</bibRefCitation>
, pl. XXV, figure A).
<bibRefCitation id="EFF44B2A9423FFD73F71FE4C119C13AC" author="Barnes RSK" pageId="8" pageNumber="2842" pagination="195 - 262" refId="ref5974" refString="Barnes RSK. 1967. The Macrophthalminae of Australasia; with a review of the evolution and morphological diversity of the type genus Macrophthalmus. Trans Zool Soc Lond. 31: 195 - 262." type="journal article" year="1967">Barnes (1967</bibRefCitation>
, p. 229, figure 10d) examined a large number of specimens from eastern and southern
<collectingCountry id="F372764B9423FFD73B8AFD8D1033104C" box="[250,354,518,542]" name="Australia" pageId="8" pageNumber="2842">Australia</collectingCountry>
and mentioned that only the movable finger has the differentiated tooth.
</paragraph>
<paragraph id="8BDA36DB9423FFD73BCAFDCC1078118F" blockId="8.[143,1104,167,989]" pageId="8" pageNumber="2842">
Two characters examined here, the male G1 and female gonopore, are proposed to be important diagnostic characters among the species of the group (Naderloo &amp; Türkay 2011).
<taxonomicName id="4C654D589423FFD73A2BFD0D137910CC" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[347,552,646,670]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC99423FFD73A2BFD0D137910CC" box="[347,552,646,670]" italics="true" pageId="8" pageNumber="2842">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B29423FFD73943FD0C132310CC" box="[563,626,647,670]" pageId="8" pageNumber="2842" rank="species">n. sp.</taxonomicNameLabel>
is rather similar to
<taxonomicName id="4C654D589423FFD73810FD0D12E910CC" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" box="[864,952,646,670]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD73810FD0D122210CF" box="[864,883,646,669]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD738F6FD0D12E910CC" box="[902,952,646,670]" italics="true" pageId="8" pageNumber="2842">lisae</emphasis>
</taxonomicName>
in these two characters and these two species are not easily distinguishable from each other.
<taxonomicName id="4C654D589423FFD73BFFFD4D100C108C" authority="Naderloo, 2013" authorityName="Naderloo" authorityYear="2013" box="[143,349,710,734]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="java" status="sp. nov.">
<emphasis id="B911EAC99423FFD73BFFFD4D100C108C" box="[143,349,710,734]" italics="true" pageId="8" pageNumber="2842">Chaenostoma java</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257B29423FFD73A17FD4D10F6108F" box="[359,423,710,733]" pageId="8" pageNumber="2842" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C654D589423FFD73934FD4D13CD108C" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" box="[580,668,710,734]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD73934FD4D1306108F" box="[580,599,710,733]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD7391AFD4D13CD108C" box="[618,668,710,734]" italics="true" pageId="8" pageNumber="2842">lisae</emphasis>
</taxonomicName>
by having a setal patch on the inner surface of the male cheliped palm extending to the fingers (see
<figureCitation id="135E2A5E9423FFD73816FD6E128C10AF" box="[870,989,741,765]" captionStart="Figure 1" captionStartId="4.[143,210,1453,1475]" captionTargetBox="[144,1096,160,1420]" captionTargetId="figure-3@4.[144,1104,160,1420]" captionTargetPageId="4" captionText="Figure 1. Chaenostoma java n. sp. holotype (NHMG). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198064" httpUri="https://zenodo.org/record/5198064/files/figure.png" pageId="8" pageNumber="2842">Figure 1D</figureCitation>
), whereas in
<taxonomicName id="4C654D589423FFD73BDFFC8D1057114C" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" box="[175,262,774,798]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD73BDFFC8D1193114F" box="[175,194,774,797]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD73BA4FC8D1057114C" box="[212,262,774,798]" italics="true" pageId="8" pageNumber="2842">lisae</emphasis>
</taxonomicName>
the setal patch is present only on the inner surface of the palm and the fingers are glabrous. Furthermore, in the new species a proximal large gap is recognized between the male cheliped fingers with a distinctly differentiated tooth on the cutting edge of the movable finger, whereas in
<taxonomicName id="4C654D589423FFD739ABFCED1263112F" baseAuthorityName="Poupin and Bouchard" baseAuthorityYear="2010" box="[731,818,869,893]" class="Malacostraca" family="Macrophthalmidae" genus="Chaenostoma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="2842" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B911EAC99423FFD739ABFCED13BF112F" box="[731,750,870,893]" italics="true" pageId="8" pageNumber="2842">C</emphasis>
.
<emphasis id="B911EAC99423FFD73870FCEE1263112F" box="[768,818,869,893]" italics="true" pageId="8" pageNumber="2842">lisae</emphasis>
</taxonomicName>
there is no gap between the fingers and the tooth of the movable finger is low and not very differentiated as in other species of the group (see
<figureCitation id="135E2A5E9423FFD7392CFC2E125311EF" box="[604,770,933,957]" captionStart="Figure 3" captionStartId="9.[211,278,1254,1276]" captionTargetBox="[212,1162,168,1221]" captionTargetId="figure-5@9.[212,1172,168,1221]" captionTargetPageId="9" captionText="Figure 3. Chaenostoma lisae (Poupin and Bouchard, 2010). Male (MNHN-B10717). (A) Carapace; (B) inner surface of male cheliped (inner face); (C) palm of right cheliped (outer face); (D) palm of right cheliped (inner face); (E) G1, right (ventral face); (F) apical part of the same G1 (lateral face); (G) apical part of the same G1 (ventral face)." figureDoi="http://doi.org/10.5281/zenodo.5198068" httpUri="https://zenodo.org/record/5198068/files/figure.png" pageId="8" pageNumber="2842">Figure 3C, D</figureCitation>
;
<bibRefCitation id="EFF44B2A9423FFD73865FC2E151911EF" author="Poupin J &amp; Bouchard JM" box="[789,1096,933,957]" pageId="8" pageNumber="2842" pagination="61 - 67" refId="ref6483" refString="Poupin J, Bouchard JM. 2010. A new dwarf sentinel crab from Mayotte Island, western Indian Ocean (Decapoda: Brachyura: Macrophthalmidae). Zootaxa. 2501: 61 - 67." type="journal article" year="2010">Poupin &amp; Bouchard 2010</bibRefCitation>
, figure 2F, G).
</paragraph>
</subSubSection>
</treatment>
</document>