264 lines
39 KiB
XML
264 lines
39 KiB
XML
<document id="CD2C18031BC81742BB14AB1D507070C0" ID-DOI="10.5281/zenodo.189408" ID-GBIF-Dataset="8aa2111b-be48-4088-929e-f2cc548447ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="189408" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461101592281" checkinUser="plazi" docAuthor="Minoshima, Yûsuke" docDate="2009" docId="03CC87B84801FF9922D05049FB1CFF5F" docLanguage="en" docName="zt02192p055.pdf" docOrigin="Zootaxa 2192" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oocyclus rupicola Minoshima, sp. n." docType="treatment" docVersion="12" lastPageNumber="51" masterDocId="FFF5FFC04800FF9E22475670FFC9FF87" masterDocTitle="First record of the hygropetric genus Oocyclus Sharp (Coleoptera: Hydrophilidae) from Laos, with description of a new species" masterLastPageNumber="55" masterPageNumber="45" pageNumber="46" updateTime="1698591619799" updateUser="plazi">
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<mods:mods id="2FF4A3D55F50770BF5C45292027853D7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E6A71213B47474A301B8C933E90FE27B">
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<mods:title id="BB1763F296FA779D3A6DC9D3D77F00F9">First record of the hygropetric genus Oocyclus Sharp (Coleoptera: Hydrophilidae) from Laos, with description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name id="373E0397EB4A9673C5D5C55B13407C70" type="personal">
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<mods:role id="14C01063765140C2A166E3A01875003F">
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<mods:roleTerm id="CC9391015535E3B87443B0B5445E100C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9FA4C45118E68FADE073D5E685313CA3">Minoshima, Yûsuke</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="754BE08749B9581AB4959EC14F2F57BD">text</mods:typeOfResource>
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<mods:titleInfo id="41E5B60EB82A91A51A6CE58A2DF95AA3">
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<mods:title id="5C11F9564864EC0CC063AEB7AA533EF4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F112FA6877973F91D38F4A51B94D0EDC">
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<mods:date id="F1829F90810A135307A6CFC917793474">2009</mods:date>
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<mods:detail id="0D53DE8925DFB98D5F56DDC1B01C4E8C" type="volume">
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<mods:number id="603CEB4549AE7335A0FE96E7F310D998">2192</mods:number>
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</mods:detail>
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<mods:extent id="B37130A5F3C3D5C9C3391CB890BE632C" unit="page">
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<mods:start id="8A96D2E405BA3950C25BADC8AA7265C8">45</mods:start>
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<mods:end id="F046A1DC7FD5BA8A8BC2C3330301391A">55</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="1E027F6025DD6B3562C82FDA1222025B">journal article</mods:classification>
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<mods:identifier id="4696652971A029754D174B5A33902A1B" type="DOI">10.5281/zenodo.189408</mods:identifier>
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<mods:identifier id="CA1E149C536CF7FB5706E4892479ECDD" type="GBIF-Dataset">8aa2111b-be48-4088-929e-f2cc548447ab</mods:identifier>
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<mods:identifier id="8DF570BC6F7781142E7E245D82E801ED" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="96DCF2897E399BA52AD9A803FAD1B392" type="Zenodo-Dep">189408</mods:identifier>
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</mods:mods>
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<treatment id="03CC87B84801FF9922D05049FB1CFF5F" ID-DOI="http://doi.org/10.5281/zenodo.6219756" ID-GBIF-Taxon="119622108" ID-Zenodo-Dep="6219756" LSID="urn:lsid:plazi:treatment:03CC87B84801FF9922D05049FB1CFF5F" httpUri="http://treatment.plazi.org/id/03CC87B84801FF9922D05049FB1CFF5F" lastPageId="7" lastPageNumber="51" pageId="1" pageNumber="46">
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<subSubSection id="C37F65254801FF9F22D05049FED1F9F1" pageId="1" pageNumber="46" type="nomenclature">
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<paragraph id="8BDA36AE4801FF9F22D05049FD9EF9D5" blockId="1.[151,599,1592,1654]" box="[151,599,1592,1619]" pageId="1" pageNumber="46">
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<heading id="D09281C24801FF9F22D05049FD9EF9D5" bold="true" box="[151,599,1592,1619]" fontSize="11" level="1" pageId="1" pageNumber="46" reason="1">
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||
<emphasis id="B911EABC4801FF9F22D05049FD9EF9D5" bold="true" box="[151,599,1592,1619]" pageId="1" pageNumber="46">
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<taxonomicName id="4C654D2D4801FF9F22D05049FDC0F9D5" ID-CoL="49RWN" authority="Minoshima" authorityName="Minoshima" box="[151,521,1592,1619]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="rupicola" status="sp. nov.">
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<emphasis id="B911EABC4801FF9F22D05049FEBBF9D4" bold="true" box="[151,370,1593,1619]" italics="true" pageId="1" pageNumber="46">Oocyclus rupicola</emphasis>
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Minoshima
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22257C74801FF9F20525048FD9EF9D5" box="[533,599,1592,1618]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="8BDA36AE4801FF9F22D0502CFED1F9F1" blockId="1.[151,599,1592,1654]" box="[151,280,1628,1654]" pageId="1" pageNumber="46">
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||
(
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<figureCitation id="135E2A2B4801FF9F22D8502CFEC7F9F1" box="[159,270,1628,1654]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,1308,1332]" captionStartId-1="4.[151,256,1940,1964]" captionStartId-2="5.[151,257,1908,1932]" captionTargetBox-0="[188,1398,197,1281]" captionTargetBox-1="[226,1360,197,1914]" captionTargetBox-2="[281,1320,185,1866]" captionTargetId-0="figure@2.[188,1398,197,1283]" captionTargetId-1="figure@4.[226,1360,197,1915]" captionTargetId-2="figure@5.[151,1436,139,1903]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Oocyclus rupicola sp. n., holotype, male: Habitus, dorsal and lateral view." captionText-1="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" captionText-2="FIGURE 3. Oocyclus rupicola sp. n.: A: Aedeagus, dorsal view; B: 9 th sternite, dorsal view; C: 9 th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A – C: male, MiYu-D- 00107; D: male, MiYu-D- 00105; E: female, MiYu-D- 00127]" httpUri-0="https://zenodo.org/record/189409/files/figure.png" httpUri-1="https://zenodo.org/record/189410/files/figure.png" httpUri-2="https://zenodo.org/record/189411/files/figure.png" pageId="1" pageNumber="46">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F65254801FF9F22D050D5FEF7F873" pageId="1" pageNumber="46" type="materials_examined">
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<paragraph id="8BDA36AE4801FF9F22D050D5FE71F961" blockId="1.[151,1438,1701,2036]" pageId="1" pageNumber="46">
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<emphasis id="B911EABC4801FF9F22D050D5FE83F938" bold="true" box="[151,330,1701,1727]" pageId="1" pageNumber="46">
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<typeStatus id="54DE880C4801FF9F22D050D5FF1CF938" box="[151,213,1701,1727]" pageId="1" pageNumber="46">Type</typeStatus>
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Locality.
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</emphasis>
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<collectingCountry id="F372763E4801FF9F231450D5FE58F938" box="[339,401,1701,1727]" name="Laos" pageId="1" pageNumber="46">Laos</collectingCountry>
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: Bolikhamxai Province,
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<quantity id="4C9D9B4B4801FF9F20F150D5FD30F938" box="[694,761,1701,1727]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="46" unit="km" value="11.0">11km</quantity>
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Southwest by West of Ban Pakha, Route 8,
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<geoCoordinate id="EE5150694801FF9F274D50D5FA5EF938" box="[1290,1431,1701,1727]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="9" value="18.197166">N18°11.83´</geoCoordinate>
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,
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<geoCoordinate id="EE5150694801FF9F22D050BCFEE7F961" box="[151,302,1740,1766]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="9" value="104.603165">E104°36.19´</geoCoordinate>
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, alt.
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<specimenCount id="9D63FD274801FF9F232050BCFE71F961" box="[359,440,1740,1766]" pageId="1" pageNumber="46" type="generic">
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||
<quantity id="4C9D9B4B4801FF9F232050BCFE7DF961" box="[359,436,1740,1766]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="46" unit="m" value="580.0">580 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph id="8BDA36AE4801FF9F22825082FEF7F873" blockId="1.[151,1438,1701,2036]" pageId="1" pageNumber="46">
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<emphasis id="B911EABC4801FF9F22825082FDC8F88B" bold="true" box="[197,513,1778,1804]" pageId="1" pageNumber="46">
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||
<typeStatus id="54DE880C4801FF9F22825082FECBF88B" box="[197,258,1778,1804]" pageId="1" pageNumber="46">Type</typeStatus>
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material.
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<typeStatus id="54DE880C4801FF9F23C35082FE32F88B" box="[388,507,1778,1804]" pageId="1" pageNumber="46" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Male. “[
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<collectingCountry id="F372763E4801FF9F20205082FD74F88B" box="[615,701,1778,1804]" name="Laos" pageId="1" pageNumber="46">LAOS</collectingCountry>
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: MiYu-L-08-039] Route 8,” / “
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<quantity id="4C9D9B4B4801FF9F266B5082FBB9F88B" box="[1068,1136,1778,1804]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="46" unit="km" value="11.0">11km</quantity>
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SWbW of Ban Pakha,” / “Bolikhamxai prov.” / “(hygropetric: wet rock);” / “alt.
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<quantity id="4C9D9B4B4801FF9F210F5169FC58F8B4" box="[840,913,1817,1843]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="46" unit="m" value="580.0">580m</quantity>
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||
;
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||
<geoCoordinate id="EE5150694801FF9F21D85169FBF9F8B4" box="[927,1072,1817,1843]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="9" value="18.197166">N18°11.83´</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE5150694801FF9F26795169FB16F8B4" box="[1086,1247,1817,1843]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="9" value="104.603165">E104°36.19´</geoCoordinate>
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;” / “
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<date id="FFDB106E4801FF9F27535169FA5EF8B4" box="[1300,1431,1817,1843]" pageId="1" pageNumber="46" value="2008-05-25">25.V.2008</date>
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; MINOSHIMA Yûsuke leg.” // “MiYu-D-0 0 140” / “MINOSHIMA Yûsuke” / “
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<collectionCode id="ED74AE6B4801FF9F26FB5130FAC6F8DD" box="[1212,1295,1856,1882]" pageId="1" pageNumber="46">SEHU</collectionCode>
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||
,
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||
<collectingCountry id="F372763E4801FF9F27635130FAA6F8DD" box="[1316,1391,1856,1882]" name="Japan" pageId="1" pageNumber="46">Japan</collectingCountry>
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” // “
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<typeStatus id="54DE880C4801FF9F22E55116FE9AF807" box="[162,339,1894,1920]" pageId="1" pageNumber="46" type="holotype">HOLOTYPE</typeStatus>
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||
” / “
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<taxonomicName id="4C654D2D4801FF9F23C05116FD4AF807" box="[391,643,1894,1920]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="rupicola">Oocyclus rupicola</taxonomicName>
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” / “Minoshima” / “det. MINOSHIMA Yûsuke” / “2009” // “0000007945” / “Sys.Ent.” / “Hokkaido Univ.” / “
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<collectingCountry id="F372763E4801FF9F20A951FDFCFDF820" box="[750,820,1933,1959]" name="Japan" pageId="1" pageNumber="46">Japan</collectingCountry>
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||
[
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<collectionCode id="ED74AE6B4801FF9F210351FDFC50F820" box="[836,921,1933,1959]" pageId="1" pageNumber="46">SEHU</collectionCode>
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||
]” [“D-00140”, handwritten on a slide glass card](
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<collectionCode id="ED74AE6B4801FF9F229A51C4FEE6F849" box="[221,303,1972,1998]" pageId="1" pageNumber="46">SEHU</collectionCode>
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||
).
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<emphasis id="B911EABC4801FF9F230451C3FE03F84A" bold="true" box="[323,458,1971,1997]" pageId="1" pageNumber="46">
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<typeStatus id="54DE880C4801FF9F230451C3FE0CF84A" box="[323,453,1971,1997]" pageId="1" pageNumber="46" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D63FD274801FF9F239551C4FDE7F849" box="[466,558,1972,1998]" pageId="1" pageNumber="46" type="male">8 males</specimenCount>
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and
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<specimenCount id="9D63FD274801FF9F202F51C4FD25F849" box="[616,748,1972,1998]" pageId="1" pageNumber="46" type="female">10 females</specimenCount>
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, same locality as
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<typeStatus id="54DE880C4801FF9F21FA51C4FBEDF849" box="[957,1060,1972,1998]" pageId="1" pageNumber="46" type="holotype">holotype</typeStatus>
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||
(
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<collectionCode id="ED74AE6B4801FF9F267251C4FB4FF849" box="[1077,1158,1972,1998]" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="1" pageNumber="46">EUMJ</collectionCode>
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,
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<collectionCode id="ED74AE6B4801FF9F26D551C4FB2EF849" box="[1170,1255,1972,1998]" pageId="1" pageNumber="46">KSEM</collectionCode>
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,
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<collectionCode id="ED74AE6B4801FF9F26B451C4FA89F849" box="[1267,1344,1972,1998]" pageId="1" pageNumber="46">SEHU</collectionCode>
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(7946– 7952),
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<collectionCode id="ED74AE6B4801FF9F22AD51AAFEFBF873" box="[234,306,2010,2036]" pageId="1" pageNumber="46">MYC</collectionCode>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF1A66264802FF9C22D0536CFBF1FAB4" box="[151,1080,1308,1332]" httpUri="https://zenodo.org/record/189409/files/figure.png" pageId="2" pageNumber="47" targetBox="[188,1398,197,1281]" targetPageId="2">
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||
<paragraph id="8BDA36AE4802FF9C22D0536CFBF1FAB4" blockId="2.[151,1080,1308,1332]" box="[151,1080,1308,1332]" pageId="2" pageNumber="47">
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||
<emphasis id="B911EABC4802FF9C22D0536CFED0FAB3" bold="true" box="[151,281,1308,1332]" pageId="2" pageNumber="47">FIGURE 1.</emphasis>
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<taxonomicName id="4C654D2D4802FF9C2367536CFE2DFAB4" box="[288,484,1308,1331]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="rupicola" status="sp. nov.">
|
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<emphasis id="B911EABC4802FF9C2367536CFE2DFAB4" box="[288,484,1308,1331]" italics="true" pageId="2" pageNumber="47">Oocyclus rupicola</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B911EABC4802FF9C23AC536CFDEFFAB3" bold="true" box="[491,550,1308,1332]" pageId="2" pageNumber="47">
|
||
<taxonomicNameLabel id="A22257C74802FF9C23AC536CFDEFFAB3" box="[491,550,1308,1332]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, holotype, male: Habitus, dorsal and lateral view.
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</paragraph>
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||
</caption>
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||
<subSubSection id="C37F65254802FF9D22825314FB40F964" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47" type="description">
|
||
<paragraph id="8BDA36AE4802FF9D22825314FB56FA94" blockId="2.[151,1437,1380,2025]" lastBlockId="3.[151,1437,152,1956]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
|
||
<emphasis id="B911EABC4802FF9C22825314FE79FAF9" bold="true" box="[197,432,1380,1406]" pageId="2" pageNumber="47">Description. Male</emphasis>
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||
. Body broadly oval, moderately convex (
|
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<figureCitation id="135E2A2B4802FF9C21FE5314FBCDFAF9" box="[953,1028,1380,1406]" captionStart="FIGURE 1" captionStartId="2.[151,255,1308,1332]" captionTargetBox="[188,1398,197,1281]" captionTargetId="figure@2.[188,1398,197,1283]" captionTargetPageId="2" captionText="FIGURE 1. Oocyclus rupicola sp. n., holotype, male: Habitus, dorsal and lateral view." httpUri="https://zenodo.org/record/189409/files/figure.png" pageId="2" pageNumber="47">Fig. 1</figureCitation>
|
||
,
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<figureCitation id="135E2A2B4802FF9C26535314FBECFAF9" box="[1044,1061,1380,1406]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="2" pageNumber="47">2</figureCitation>
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||
A).
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<emphasis id="B911EABC4802FF9C26125314FB6CFAF9" bold="true" box="[1109,1189,1380,1406]" pageId="2" pageNumber="47">Color.</emphasis>
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Dorsum black with greenish luster, slightly iridescent; but more vividly colored when wet condition. Ventral surface of head dark brown to black. Antennae, maxillary palpi, labial palpi rather light yellowish-brown; pubescent club of antennae brown; apex of maxillary palpomere 4 darkened. Mentum dark reddish-brown, anterior margin slightly paler; stipes reddish-brown, paler than ventral face of head. Lateral margins of prosternum, epipleura, and pseudepipleura reddish-brown, paler than median part of prosternum, mesoventrite, and metaventrite; epipleura slightly darker than pseudepipleura. Legs reddish brown; coxa and femora slightly darkened; tibia and tarsus paler. Abdominal ventrites dark reddish-brown to dark brown; posterior margins of ventrites slightly paler.
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<emphasis id="B911EABC4802FF9C230850E9FE56F934" bold="true" box="[335,415,1689,1715]" pageId="2" pageNumber="47">Head.</emphasis>
|
||
Labrum about 2.8 times as wide as long; punctation of labrum moderately fine, moderately densely distribution; systematic punctures of labrum composed of a row of coarse punctures bearing fine setae medially, dense mediolaterally, sparse medially. Clypeus with broadly and weakly rounded anterior margin; punctation of clypeus and frons composed of variously sized distinct punctures; interspaces between punctures about 0.5–1.0 times width of a puncture, sometimes more or less distance. Frontoclypeal suture fine, but distinct. Systematic punctures of frons bearing fine, short setae, present roundly along each eye. Antennal scape as long as the combined length of antennomeres 2–4; antennomere 3 short; antennomere 4 very short, about 0.5 times as long as antennomere 3; antennomere 5 moderately long, about the same length as antennomeres 6 and 7 combined; apical segment slightly longer than antennomere 7. Maxillary palpi short; palpomere 3 slightly shorter than palpomere 2; palpomere 4 about 1.3 times as long as palpomere 2. Labial palpi short, slightly shorter than width of mentum; palpomere 2 about as long as palpomere 3; palpomeres 2 and 3 with a few fine setae. Mentum flat, rectangular, anterior margin rounded, about 1.3 times as wide as long; punctation of mentum very fine, sparse; anterolateral margin of mentum with fine, distinct punctures bearing fine, short setae (
|
||
<figureCitation id="135E2A2B4803FF9D23FA5742FDCAFECB" box="[445,515,306,332]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
E).
|
||
<emphasis id="B911EABC4803FF9D206C5742FD46FECB" bold="true" box="[555,655,306,332]" pageId="3" pageNumber="48">Thorax.</emphasis>
|
||
Pronotal punctation moderately coarse, punctures of various sizes, finer and more densely distributed than those of clypeus and frons; interspaces between punctures about 0.5– 1.0 times width of a puncture; a few sparse setiferous punctures bearing fine, short erect seta present on lateral margin. Systematic punctures of pronotum bearing fine short recumbent setae; anterior series distinct, composed of an irregular row, longer than posterior row; posterior series slightly indistinct, more irregular than anterior one. Posterolateral corners of pronotum angulate (
|
||
<figureCitation id="135E2A2B4803FF9D21C45784FC02FD89" box="[899,971,500,526]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
C). Elytra longer than wide, subparallel on anterior half, broadly rounded apically; punctation of elytra moderately coarse, punctures of various sizes, more densely distributed than those of pronotum (
|
||
<figureCitation id="135E2A2B4803FF9D20B35431FCF4FDDC" box="[756,829,577,603]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
D); a few sparse setiferous punctures bearing fine, short erect seta present on outer margin; elytra with five rows of coarse, sparse setiferous punctures bearing recumbent to erect fine setae; punctures of rows slightly undetectable; row 5 with very sparsely distributed punctures. Prosternum with median carina along entire length, slightly projecting anteriorly, with fine setae on anterior margin (
|
||
<figureCitation id="135E2A2B4803FF9D232754ACFE61FD71" box="[352,424,732,758]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
F). Mesoventral process with a few fine setae on median carina; projecting ventrally, from lateral view slightly resembling bird's beak (
|
||
<figureCitation id="135E2A2B4803FF9D209C5572FCE8FC9B" box="[731,801,770,796]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
G). Metaventrite with rather oblong oval glabrous area posteromedially, about 2.8 times as long as wide, length of glabrous area about 0.6 times total length of metaventrite. Procoxae with moderately fine, densely distributed setae, with a few indistinct spines (
|
||
<figureCitation id="135E2A2B4803FF9D27735520FAB2FCED" box="[1332,1403,848,874]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
|
||
F); meso- and metacoxae with coarse setae. Profemur densely pubescent on about basal two-fifths, without distinct punctures; mesofemur with moderately coarse, irregularly and sparsely distributed punctures, interspaces between punctures about 1.0–4.0 times width of a puncture; punctation of metafemur finer, punctures sparser than those of mesofemur. Protarsal segments 1–4 small, subequal in length; apical segment about as long as combined length of segments 1–4.
|
||
<emphasis id="B911EABC4803FF9D21425260FC41FBAD" bold="true" box="[773,904,1040,1066]" pageId="3" pageNumber="48">Abdomen.</emphasis>
|
||
Abdominal ventrites with moderately dense pubescence. Aedeagus (
|
||
<figureCitation id="135E2A2B4803FF9D23F15248FDC1FBD5" box="[438,520,1080,1106]" captionStart="FIGURE 3" captionStartId="5.[151,257,1908,1932]" captionTargetBox="[281,1320,185,1866]" captionTargetId="figure@5.[151,1436,139,1903]" captionTargetPageId="5" captionText="FIGURE 3. Oocyclus rupicola sp. n.: A: Aedeagus, dorsal view; B: 9 th sternite, dorsal view; C: 9 th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A – C: male, MiYu-D- 00107; D: male, MiYu-D- 00105; E: female, MiYu-D- 00127]" httpUri="https://zenodo.org/record/189411/files/figure.png" pageId="3" pageNumber="48">Figs. 3</figureCitation>
|
||
A, D): parameres about as long median lobe, swollen at base, subparallel on basal half of outer margin thence narrowing apically; punctation of parameres fine, moderately densely distributed; median lobe slender, slightly narrowed apically, weakly curved dorsally from lateral view (
|
||
<figureCitation id="135E2A2B4803FF9D272B52F5FF6CFB41" captionStart="FIGURE 3" captionStartId="5.[151,257,1908,1932]" captionTargetBox="[281,1320,185,1866]" captionTargetId="figure@5.[151,1436,139,1903]" captionTargetPageId="5" captionText="FIGURE 3. Oocyclus rupicola sp. n.: A: Aedeagus, dorsal view; B: 9 th sternite, dorsal view; C: 9 th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A – C: male, MiYu-D- 00107; D: male, MiYu-D- 00105; E: female, MiYu-D- 00127]" httpUri="https://zenodo.org/record/189411/files/figure.png" pageId="3" pageNumber="48">Fig. 3</figureCitation>
|
||
D); punctation of median lobe very fine, punctures sparsely distributed; basal aphophysis moderately short; phallobase about 0.6 times as wide as long; punctation of phallobase fine, few sparsely distributed punctures present laterally; manubrium broadly rounded, indistinctly demarcated from phallobase.
|
||
</paragraph>
|
||
<paragraph id="8BDA36AE4803FF9D2282536FFB17FA53" blockId="3.[151,1437,152,1956]" pageId="3" pageNumber="48">
|
||
<emphasis id="B911EABC4803FF9D2282536FFEE2FABE" bold="true" box="[197,299,1311,1337]" pageId="3" pageNumber="48">Female.</emphasis>
|
||
Body size slightly larger than male (see measurements); otherwise almost identical to male.
|
||
<emphasis id="B911EABC4803FF9D22D05336FE5DFAE7" bold="true" box="[151,404,1350,1376]" pageId="3" pageNumber="48">Spermathecal organ</emphasis>
|
||
(
|
||
<figureCitation id="135E2A2B4803FF9D23E35336FE20FAE7" box="[420,489,1350,1376]" captionStart="FIGURE 3" captionStartId="5.[151,257,1908,1932]" captionTargetBox="[281,1320,185,1866]" captionTargetId="figure@5.[151,1436,139,1903]" captionTargetPageId="5" captionText="FIGURE 3. Oocyclus rupicola sp. n.: A: Aedeagus, dorsal view; B: 9 th sternite, dorsal view; C: 9 th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A – C: male, MiYu-D- 00107; D: male, MiYu-D- 00105; E: female, MiYu-D- 00127]" httpUri="https://zenodo.org/record/189411/files/figure.png" pageId="3" pageNumber="48">Fig. 3</figureCitation>
|
||
E). Spermathecal duct long (278–296 (286 ± 7) μm; n = 5), distinctly longer than duct of the spermathecal gland, slender, slightly swollen at base; infundibulum undetectable; spermatheca rather small, slightly oblong oval; cornu undetectable; duct of the spermathecal gland short (58–72 (66 ± 5) μm; n = 5), swollen at medially, slender at base; spermathecal gland slender, slightly swollen.
|
||
</paragraph>
|
||
<paragraph id="8BDA36AE4803FF9D22825390FB40F964" blockId="3.[151,1437,152,1956]" pageId="3" pageNumber="48">
|
||
<emphasis id="B911EABC4803FF9D22825390FE4CFA7D" bold="true" box="[197,389,1504,1530]" pageId="3" pageNumber="48">Measurements.</emphasis>
|
||
Male (n = 8): TL: 4.93–5.55 (5.22 ± 0.21) mm; HW: 1.45–1.60 (1.53 ± 0.06) mm; ED: 0.90–1.00 (0.94 ± 0.04) mm; PL: 1.18–1.28 (1.22 ± 0.04) mm; PW: 3.03–3.35 (3.18 ± 0.12) mm; EL: 3.75– 4.30 (4.01 ± 0.18) mm; EW: 3.40–3.75 (3.56 ± 0.14) mm; ED/HW: 0.59–0.63 (0.61 ± 0.01); PW/PL: 2.56– 2.68 (2.62 ± 0.04); EL/EW: 1.10–1.15 (1.13 ± 0.02). Female (n=10): TL: 5.28–5.85 (5.50 ± 0.19) mm; HW: 1.53–1.70 (1.62 ± 0.05) mm; ED: 0.95–1.08 (1.00 ± 0.04) mm; PL: 1.20–1.35 (1.28 ± 0.05) mm; PW: 3.10– 3.53 (3.31 ± 0.12) mm; EL: 4.05–4.50 (4.23 ± 0.16) mm; EW: 3.58–3.93 (3.75 ± 0.11) mm; ED/HW: 0.61– 0.65 (0.62 ± 0.01); PW/PL: 2.53–2.69 (2.59 ± 0.05); EL/EW: 1.09–1.19 (1.13 ± 0.03).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65254803FF9D2282509FFC99F88D" box="[197,848,1775,1802]" pageId="3" pageNumber="48" type="distribution">
|
||
<paragraph id="8BDA36AE4803FF9D2282509FFC99F88D" blockId="3.[151,1437,152,1956]" box="[197,848,1775,1802]" pageId="3" pageNumber="48">
|
||
<emphasis id="B911EABC4803FF9D2282509FFEACF88E" bold="true" box="[197,357,1775,1801]" pageId="3" pageNumber="48">Distribution.</emphasis>
|
||
<collectingCountry id="F372763E4803FF9D232B5080FE60F88D" box="[364,425,1776,1802]" name="Laos" pageId="3" pageNumber="48">Laos</collectingCountry>
|
||
. Known only from the
|
||
<typeStatus id="54DE880C4803FF9D20FF5080FD23F88D" box="[696,746,1776,1802]" pageId="3" pageNumber="48">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65254803FF9D22825166FCFDF8B7" box="[197,820,1814,1840]" pageId="3" pageNumber="48" type="biology_ecology">
|
||
<paragraph id="8BDA36AE4803FF9D22825166FCFDF8B7" blockId="3.[151,1437,152,1956]" box="[197,820,1814,1840]" pageId="3" pageNumber="48">
|
||
<emphasis id="B911EABC4803FF9D22825166FEE0F8B7" bold="true" box="[197,297,1814,1840]" pageId="3" pageNumber="48">Biology.</emphasis>
|
||
Collected at the hygropetric habitat (
|
||
<figureCitation id="135E2A2B4803FF9D209A5166FCECF8B7" box="[733,805,1814,1840]" captionStart="FIGURE 6. A" captionStartId="8.[151,255,1838,1862]" captionTargetBox="[188,1398,194,1814]" captionTargetId="figure@8.[188,1398,194,1814]" captionTargetPageId="8" captionText="FIGURE 6. A: Type locality and habitat of O. rupicola sp. n., and collecting locality of O. thailensis Short et Swanson, Laos: Route 8, 11 km Southwest by South of Ban Pakha, Bolikhamxai Province, 25 May 2008; B, C: Oocyclus species, same data as Fig. 6 A. Photographs by the author." httpUri="https://zenodo.org/record/189414/files/figure.png" pageId="3" pageNumber="48">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F65254803FF9D2282514DFDA0F823" pageId="3" pageNumber="48" type="etymology">
|
||
<paragraph id="8BDA36AE4803FF9D2282514DFDA0F823" blockId="3.[151,1437,152,1956]" pageId="3" pageNumber="48">
|
||
<emphasis id="B911EABC4803FF9D2282514DFE98F8D0" bold="true" box="[197,337,1853,1879]" pageId="3" pageNumber="48">Etymology.</emphasis>
|
||
The specific epithet is a combination of the Latin word “rupes”, meaning “rock”, and another Latin word “-cola”, meaning "inhabitant". The name was chosen in order to refer to the wet rock surface, which is the habitat of this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A66264804FF9A22D051E4FD2FF876" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="4" pageNumber="49" targetBox="[226,1360,197,1914]" targetPageId="4">
|
||
<paragraph id="8BDA36AE4804FF9A22D051E4FD2FF876" blockId="4.[151,1437,1940,2033]" pageId="4" pageNumber="49">
|
||
<emphasis id="B911EABC4804FF9A22D051E4FED2F82B" bold="true" box="[151,283,1940,1964]" pageId="4" pageNumber="49">FIGURE 2.</emphasis>
|
||
SEM photographs of
|
||
<taxonomicName id="4C654D2D4804FF9A204B51E5FD40F82B" box="[524,649,1941,1964]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="rupicola" status="sp. nov.">
|
||
<emphasis id="B911EABC4804FF9A204B51E5FD40F82B" box="[524,649,1941,1964]" italics="true" pageId="4" pageNumber="49">O. rupicola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EABC4804FF9A20D551E4FD06F82B" bold="true" box="[658,719,1940,1964]" pageId="4" pageNumber="49">
|
||
<taxonomicNameLabel id="A22257C74804FF9A20D551E4FD06F82B" box="[658,719,1940,1964]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D-00140]
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A66264805FF9B22D05104FDC0F874" httpUri="https://zenodo.org/record/189411/files/figure.png" pageId="5" pageNumber="50" targetBox="[281,1320,185,1866]" targetPageId="5">
|
||
<paragraph id="8BDA36AE4805FF9B22D05104FDC0F874" blockId="5.[151,1437,1908,2035]" pageId="5" pageNumber="50">
|
||
<emphasis id="B911EABC4805FF9B22D05104FED4F80B" bold="true" box="[151,285,1908,1932]" pageId="5" pageNumber="50">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C654D2D4805FF9B23635105FE26F80B" box="[292,495,1909,1932]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="rupicola" status="sp. nov.">
|
||
<emphasis id="B911EABC4805FF9B23635105FE26F80B" box="[292,495,1909,1932]" italics="true" pageId="5" pageNumber="50">Oocyclus rupicola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EABC4805FF9B23B05104FDFCF80B" bold="true" box="[503,565,1908,1932]" pageId="5" pageNumber="50">
|
||
<taxonomicNameLabel id="A22257C74805FF9B23B05104FDFCF80B" box="[503,565,1908,1932]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A: Aedeagus, dorsal view; B: 9th sternite, dorsal view; C: 9th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A–C: male, MiYu-D-00107; D: male, MiYu-D- 00105; E: female, MiYu-D-00127]
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A66264806FF9822D05169FBADF8F1" httpUri="https://zenodo.org/record/189412/files/figure.png" pageId="6" pageNumber="51" targetBox="[226,1360,194,1793]" targetPageId="6">
|
||
<paragraph id="8BDA36AE4806FF9822D05169FBADF8F1" blockId="6.[151,1436,1817,1910]" pageId="6" pageNumber="51">
|
||
<emphasis id="B911EABC4806FF9822D05169FED4F8B6" bold="true" box="[151,285,1817,1841]" pageId="6" pageNumber="51">FIGURE 4.</emphasis>
|
||
SEM photographs of
|
||
<taxonomicName id="4C654D2D4806FF982057516AFCA1F8B6" authority="Short et Swanson" authorityName="Short et Swanson" box="[528,872,1818,1841]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="thailensis">
|
||
<emphasis id="B911EABC4806FF982057516AFD55F8B6" box="[528,668,1818,1841]" italics="true" pageId="6" pageNumber="51">O. thailensis</emphasis>
|
||
Short et Swanson
|
||
</taxonomicName>
|
||
: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum and mesoventrite, ventral view. [A, C–D: male, MiYu-D-00136; B, E–F: male, MiYu-D-00138]
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F65254806FF99228251D7FB1CFF5F" lastPageId="7" lastPageNumber="52" pageId="6" pageNumber="51" type="discussion">
|
||
<paragraph id="8BDA36AE4806FF99228251D7FB1CFF5F" blockId="6.[151,1437,1959,2024]" lastBlockId="7.[151,1436,152,216]" lastPageId="7" lastPageNumber="52" pageId="6" pageNumber="51">
|
||
<emphasis id="B911EABC4806FF98228251D7FE89F846" bold="true" box="[197,320,1959,1985]" pageId="6" pageNumber="51">Remarks.</emphasis>
|
||
Judging from descriptions and figures of
|
||
<bibRefCitation id="EFF44B5F4806FF98217B51D7FBA5F846" author="Short" box="[828,1132,1959,1985]" pageId="6" pageNumber="55" refString="Short, A. E. Z. & Swanson, A. P. (2005) A revision of the Oocyclus Sharp of Thailand with description of six new species (Coleoptera: Hydrophilidae). Zootaxa, 1078, 1 - 24." type="journal article" year="2005">Short & Swanson (2005)</bibRefCitation>
|
||
, this new species is most similar to
|
||
<taxonomicName id="4C654D2D4806FF98235251BEFD01F86F" authority="d'Orchymont, 1932" authorityName="d'Orchymont" authorityYear="1932" box="[277,712,1998,2024]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="sumatrensis">
|
||
<emphasis id="B911EABC4806FF98235251BEFE05F860" box="[277,460,1998,2023]" italics="true" pageId="6" pageNumber="51">O. sumatrensis</emphasis>
|
||
d’Orchymont, 1932
|
||
</taxonomicName>
|
||
; it can, however, be distinguished from it by the following characters: 1) spines of procoxae thin, sparse and indistinct, resembling thickened, stiff setae (
|
||
<figureCitation id="135E2A2B4807FF99274856E8FA90FF35" box="[1295,1369,152,178]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="7" pageNumber="52">Fig. 2</figureCitation>
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F); 2) ground punctation of elytra very coarse (
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<figureCitation id="135E2A2B4807FF99203356CEFD72FF5F" box="[628,699,190,216]" captionStart="FIGURE 2" captionStartId="4.[151,256,1940,1964]" captionTargetBox="[226,1360,197,1914]" captionTargetId="figure@4.[226,1360,197,1915]" captionTargetPageId="4" captionText="FIGURE 2. SEM photographs of O. rupicola sp. n.: A: Habitus, dorsal view; B: ditto, ventral view; C: posterolateral corner of pronotum, dorsal view; D: punctation of elytra; E: head, ventral view; F: prosternum, ventral view; G: mesoventrite, ventral view. [Holotype: MiYu-D- 00140]" httpUri="https://zenodo.org/record/189410/files/figure.png" pageId="7" pageNumber="52">Fig. 2</figureCitation>
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D); 3) parameres narrowed apically (
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<figureCitation id="135E2A2B4807FF99262E56CEFB67FF5F" box="[1129,1198,190,216]" captionStart="FIGURE 3" captionStartId="5.[151,257,1908,1932]" captionTargetBox="[281,1320,185,1866]" captionTargetId="figure@5.[151,1436,139,1903]" captionTargetPageId="5" captionText="FIGURE 3. Oocyclus rupicola sp. n.: A: Aedeagus, dorsal view; B: 9 th sternite, dorsal view; C: 9 th tergite, ventral view; D: median lobe, lateral view; E: spermathecal organ. bc: bursa copulatrix; dsg: duct of the spermathecal gland; s: spermatheca; sd: spermathecal duct; sg: spermathecal gland. [Paratypes: A – C: male, MiYu-D- 00107; D: male, MiYu-D- 00105; E: female, MiYu-D- 00127]" httpUri="https://zenodo.org/record/189411/files/figure.png" pageId="7" pageNumber="52">Fig. 3</figureCitation>
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A).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |