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<document id="A97817199C1CDE06EEA624FF4E6AF283" ID-DOI="10.5281/zenodo.5333042" ID-ISSN="2345-7600" ID-Zenodo-Dep="5333042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630173599483" checkinUser="diego" docAuthor="Chen, Hong-Wei, Toda, Masanori J., Lakim, Maklarin B. &amp; Mohamed, Maryati B." docDate="2007" docId="03CC87B29817007CB861688839B4FC6D" docLanguage="en" docName="55rbz23-41.pdf" docOrigin="Raffles Bulletin of Zoology 55 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Phortica (Phortica) membranifera Chen &amp; Toda 2007, new species" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="FFF5FFCA9818006CBB3668493A62FF95" masterDocTitle="The Phortica Sensu Stricto (Insecta: Diptera: Drosophilidae) From Malaysia" masterLastPageNumber="41" masterPageNumber="23" pageNumber="38" updateTime="1699173972972" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B3E17B7737B949FA252225D63778A8C0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EFBFFB3AD840052B33159465AD1A381C">
<mods:title id="4BA15D0BEFE54C295A19E0F09155305D">The Phortica Sensu Stricto (Insecta: Diptera: Drosophilidae) From Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="85312052BFAC95D208D2C3DF11C11CF8" type="personal">
<mods:role id="96EA13A7071D472DD539BE4CA7BD9D00">
<mods:roleTerm id="AE18A9E76CBC3FA319651FD3A3266EF2">Author</mods:roleTerm>
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<mods:namePart id="2BE0D328819B1321E72837E7BC7CDBF7">Chen, Hong-Wei</mods:namePart>
</mods:name>
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<mods:namePart id="E15EE596AC3A4C9E2B05AEE05CF905F6">Toda, Masanori J.</mods:namePart>
</mods:name>
<mods:name id="530BDD0D7BD124A4A478C1BD9A4F05F3" type="personal">
<mods:role id="5F74E2742F41DD6227ABABD7039C60AE">
<mods:roleTerm id="C1E94FFAFD5DF011F3F24E644057E461">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="29B3098FBDCD72A70A4A9E034BC11BDD">Lakim, Maklarin B.</mods:namePart>
</mods:name>
<mods:name id="43249F9F7B855605F09C29B6975813E9" type="personal">
<mods:role id="77A3CB6157F600E0B420CA5C20A9C81F">
<mods:roleTerm id="C141409F70FF119B94EB03BF031823CB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43839D3BF2E957A22E15F083B98DED4F">Mohamed, Maryati B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="186352D59EF8E2FE3F05AA93173B6508">text</mods:typeOfResource>
<mods:relatedItem id="E5BEC34AD65C5F2C73BC01D2F9DEDAC2" type="host">
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<mods:title id="B977F0257B36111B44A80235B20940EB">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="B831EA6FA1C4CAD0EBB7F05F08B41710">
<mods:date id="372222E2A06A3D3C39190BC4C59EB55B">2007</mods:date>
<mods:detail id="ED8143BEE7A83B12391CE978EB434857" type="pubDate">
<mods:number id="323C6CA78AFF6B8496FBAE217BF9822F">2007-02-28</mods:number>
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<mods:detail id="CBC03B84D58D5CFABB2180A1C32BF302" type="volume">
<mods:number id="270986C8A65C6C2CA437229A805961DD">55</mods:number>
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<mods:classification id="55C2F1AAED7D3CF2ED086B2C12CD765F">journal article</mods:classification>
<mods:identifier id="C8226E6094936A9529FD66144B91E4B7" type="DOI">10.5281/zenodo.5333042</mods:identifier>
<mods:identifier id="26744442556DAE7322590646D5D6E7B1" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="1F2FF95268265B0D269E5ED045F731A0" type="Zenodo-Dep">5333042</mods:identifier>
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<treatment id="03CC87B29817007CB861688839B4FC6D" LSID="urn:lsid:plazi:treatment:03CC87B29817007CB861688839B4FC6D" httpUri="http://treatment.plazi.org/id/03CC87B29817007CB861688839B4FC6D" lastPageId="16" lastPageNumber="39" pageId="15" pageNumber="38">
<subSubSection id="C37F652F98170063B86168883ECCFF6D" pageId="15" pageNumber="38" type="nomenclature">
<paragraph id="8BDA36A498170063B86168883ECCFF6D" blockId="15.[855,1408,192,280]" pageId="15" pageNumber="38">
<heading id="D09281C898170063B86168883ECCFF6D" bold="true" centered="true" fontSize="10" level="2" pageId="15" pageNumber="38" reason="2">
<emphasis id="B911EAB698170063B86168883ECCFF6D" bold="true" pageId="15" pageNumber="38">
<taxonomicName id="4C654D2798170063B86168883F1EFF4D" ID-CoL="8DBWC" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" authorityYear="2007" box="[855,1404,192,217]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="membranifera" status="sp. nov." subGenus="Phortica">
<emphasis id="B911EAB698170063B861688839D4FF4C" bold="true" box="[855,950,193,217]" italics="true" pageId="15" pageNumber="38">Phortica</emphasis>
(
<emphasis id="B911EAB698170063B8F168883E44FF4C" bold="true" box="[967,1062,193,217]" italics="true" pageId="15" pageNumber="38">Phortica</emphasis>
)
<emphasis id="B911EAB698170063BF0068883EB1FF4C" bold="true" box="[1078,1235,193,217]" italics="true" pageId="15" pageNumber="38">membranifera</emphasis>
Chen &amp; Toda
</taxonomicName>
,
<taxonomicNameLabel id="A22257CD98170063BF1F68A93ECCFF6D" box="[1065,1198,224,248]" pageId="15" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37F652F98170063BF1569483ED6FE8D" box="[1059,1204,257,280]" pageId="15" pageNumber="38" type="description">
<paragraph id="8BDA36A498170063BF1569483ED6FE8D" blockId="15.[855,1408,192,280]" box="[1059,1204,257,280]" pageId="15" pageNumber="38">
(
<figureCitation id="135E2A2198170063BF1D69483ECFFE8D" box="[1067,1197,257,280]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="15" pageNumber="38">Figs. 5457</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F652F98170063B81A69093F69FD97" pageId="15" pageNumber="38" type="diagnosis">
<paragraph id="8BDA36A498170063B81A69093F69FD97" blockId="15.[812,1452,320,514]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A690939C5FEC0" bold="true" box="[812,935,320,341]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A690939F7FEC0" bold="true" box="[812,917,320,341]" italics="true" pageId="15" pageNumber="38">Diagnosis.</emphasis>
</emphasis>
Vertical process of gonopods with 2 pairs of strongly sclerotized processes connected to each other with membrane: processes of the posterior pair apically fused together (
<figureCitation id="135E2A2198170063BE6269333FFDFE1A" box="[1364,1439,378,399]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="15" pageNumber="38">Fig. 57</figureCitation>
); aedeagal ventral bridge strongly sclerotized, elongated to tip of aedeagal median rod, pointed apically, with large pointed flap submedially (
<figureCitation id="135E2A2198170063B88269993E62FE70" box="[948,1024,464,485]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="15" pageNumber="38">Fig. 57</figureCitation>
); aedeagal median rod slightly sclerotized proximally, with bipartite lobe apically (
<figureCitation id="135E2A2198170063BF8169A43F62FD97" box="[1207,1280,493,514]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="15" pageNumber="38">Fig. 57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652F9817007CB81A6A6E3E72FD8D" lastPageId="16" lastPageNumber="39" pageId="15" pageNumber="38" type="materials_examined">
<paragraph id="8BDA36A498170063B81A6A6E3ED0FD07" blockId="15.[812,1451,550,658]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A6A6E3E76FDAE" bold="true" box="[812,1044,550,572]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A6A6E399FFDA9" bold="true" box="[812,1021,551,572]" italics="true" pageId="15" pageNumber="38">Material examined.</emphasis>
</emphasis>
<materialsCitation id="3B0D3CF998170063BF286A6E3ECAFDE0" collectingDate="1999-10-18" collectionCode="KPSP" collectorName="M. J. Toda" country="Malaysia" location="Sabah" pageId="15" pageNumber="38" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54DE880698170063BF286A6E3E1FFDA9" box="[1054,1149,551,572]" pageId="15" pageNumber="38" type="holotype">Holotype</typeStatus>
male (terminalia dissected),
<collectingCountry id="F372763498170063B81A6A0D39CCFDCC" box="[812,942,580,601]" name="Malaysia" pageId="15" pageNumber="38">MALAYSIA</collectingCountry>
: Ulu Senagang, Crocker Range,
<collectingRegion id="49A1F84698170063BFD66A0D3F7FFDCC" box="[1248,1309,580,601]" country="Malaysia" name="Sabah" pageId="15" pageNumber="38">Sabah</collectingRegion>
, on fallen logs, coll.
<collectorName id="2690537298170063B8696A2939ACFDE0" box="[863,974,608,629]" pageId="15" pageNumber="38">M. J. Toda</collectorName>
,
<date id="FFDB106498170063B8EC6A293E31FDE0" box="[986,1107,608,629]" pageId="15" pageNumber="38" value="1999-10-18">
<collectingDate id="EF9FE98C98170063B8EC6A293E31FDE0" box="[986,1107,608,629]" pageId="15" pageNumber="38" value="1999-10-18">18 Oct.1999</collectingDate>
</date>
(
<collectionCode id="ED74AE6198170063BF546A293EC3FDE0" box="[1122,1185,608,629]" pageId="15" pageNumber="38">KPSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B0D3CF998170063BF856A293ECCFD07" collectingDate="1999-10-18" collectionCode="KPSP, SEHU, ITBC" collectorName="M. J. Toda" country="Malaysia" location="Sabah" pageId="15" pageNumber="38" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54DE880698170063BF856A293F76FDE0" box="[1203,1300,608,629]" pageId="15" pageNumber="38" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D63FD2D98170063BE296A293F0FFDE0" box="[1311,1389,608,629]" pageId="15" pageNumber="38" type="male">3 males</specimenCount>
, same data as holotype (
<collectionCode id="ED74AE6198170063B8EF6A343E74FD07" box="[985,1046,637,658]" pageId="15" pageNumber="38">KPSP</collectionCode>
,
<collectionCode id="ED74AE6198170063BF166A343E03FD07" box="[1056,1121,637,658]" pageId="15" pageNumber="38">SEHU</collectionCode>
,
<collectionCode id="ED74AE6198170063BF5A6A343EC5FD07" box="[1132,1191,637,658]" pageId="15" pageNumber="38">ITBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDA36A498170063B81A6AF33E73FD64" blockId="15.[812,1451,698,753]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A6AF339DEFD47" bold="true" box="[812,956,698,722]" pageId="15" pageNumber="38">
<emphasis id="B911EAB698170063B81A6AF339CAFD47" bold="true" box="[812,936,698,722]" italics="true" pageId="15" pageNumber="38">Decription.</emphasis>
</emphasis>
Male. Head: Frons dark brown on upper half, brown on lower half.
</paragraph>
<paragraph id="8BDA36A498170063B81A6B533E22FCC4" blockId="15.[812,1451,794,849]" pageId="15" pageNumber="38">Thorax: Orange yellow. Scutellum with trifurcate brown patch, pale yellow on tip.</paragraph>
<paragraph id="8BDA36A49817007CB81A6B333961FF6D" blockId="15.[812,1267,890,913]" lastBlockId="16.[136,774,193,248]" lastPageId="16" lastPageNumber="39" pageId="15" pageNumber="38">Wing: Slightly smoky on anterior margin. Legs: All femora dark brown except apical portion. Midleg tibia subapically with 34 longer setae on anterior surface.</paragraph>
<caption id="DF1A662C98170063BB4760463FE5F7D4" ID-DOI="http://doi.org/10.5281/zenodo.5333072" ID-Zenodo-Dep="5333072" httpUri="https://zenodo.org/record/5333072/files/figure.png" pageId="15" pageNumber="38" startId="15.[113,157,2063,2084]" targetBox="[116,1447,1087,2014]" targetPageId="15">
<paragraph id="8BDA36A498170063BB4760463FE5F7D4" blockId="15.[113,1451,2063,2113]" pageId="15" pageNumber="38">
Figs. 5053.
<taxonomicName id="4C654D2798170063BBC7605938D3F7B1" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" authorityYear="2007" box="[241,689,2063,2085]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="38" phylum="Arthropoda" rank="species" species="pasohensis" status="sp. nov." subGenus="Phortica">
<emphasis id="B911EAB698170063BBC760593B27F7B0" box="[241,325,2064,2085]" italics="true" pageId="15" pageNumber="38">Phortica</emphasis>
(
<emphasis id="B911EAB698170063BA6260593BCAF7B0" box="[340,424,2064,2085]" italics="true" pageId="15" pageNumber="38">Phortica</emphasis>
)
<emphasis id="B911EAB698170063BA8E60593843F7B0" box="[440,545,2064,2085]" italics="true" pageId="15" pageNumber="38">pasohensis</emphasis>
Chen &amp; Toda
</taxonomicName>
,
<taxonomicNameLabel id="A22257CD98170063B98D60463951F7B1" box="[699,819,2063,2084]" pageId="15" pageNumber="38" rank="species">new species</taxonomicNameLabel>
, male: 50, epandrium and cercus in lateral view; 51, surstylus in frontal view; 52, tenth sternite and additional plate in frontal view; 53, hypandrium, gonopod, paramere and aedeagus in lateral view.
</paragraph>
</caption>
<paragraph id="8BDA36A49808007CBBBE69683B2AFE4D" blockId="16.[136,775,289,472]" pageId="16" pageNumber="39">Abdomen: First to fifth tergites yellow; second tergite with brown patches submedially; third to fifth tergites each with medially slightly protruded, brown band on posterior margin and 1 pair of yellow patches laterally; sixth tergite nearly entirely dark except yellow medial line, with expanded lobe on lateral margin.</paragraph>
<paragraph id="8BDA36A49808007CBBBE6A4838AFFCCD" blockId="16.[136,775,513,856]" pageId="16" pageNumber="39">
Male terminalia: Epandrium pubescent anteroventrally and dorsomedially, with 1314 setae on dorsal to posterolateral portion per side (
<figureCitation id="135E2A219808007CBA6F6A083BD7FDCD" box="[345,437,577,600]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="16" pageNumber="39">Fig. 54</figureCitation>
). Surstylus with pubescence subbasally and several wedge-shaped prensisetae on apical margin to inner surface (
<figureCitation id="135E2A219808007CBAA56AC83B86FD0D" box="[403,484,641,664]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="16" pageNumber="39">Fig. 55</figureCitation>
). Additional plate between cerci and 10th sternite pubescent, medially connected to 10th sternite (
<figureCitation id="135E2A219808007CBBC26A883B32FD4D" box="[244,336,705,728]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="16" pageNumber="39">Fig. 56</figureCitation>
). Posterolateral lobe of hypandrium undeveloped. Paramere medially pubescent; apical process slender, with 1 tooth; median process with 1 sensillum and 3 teeth; proximal process round apically (
<figureCitation id="135E2A219808007CB9016B6838EBFCAD" box="[567,649,801,824]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="16" pageNumber="39">Fig. 57</figureCitation>
). Aedeagus with 2 pairs of bridges; basal bridge looped (
<figureCitation id="135E2A219808007CB9466B0838A3FCCD" box="[624,705,833,856]" captionStart="Figs" captionStartId="16.[136,180,2065,2086]" captionTargetBox="[141,1476,1099,1998]" captionTargetId="figure-8@16.[112,1507,1063,2027]" captionTargetPageId="16" captionText="Figs. 5457. Phortica (Phortica) membranifera Chen &amp; Toda, new species, male: 54, epandrium and cercus in lateral view; 55, surstylus in frontal view; 56, tenth sternite and additional plate in frontal view; 57, hypandrium, gonopod, paramere and aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5333076" httpUri="https://zenodo.org/record/5333076/files/figure.png" pageId="16" pageNumber="39">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A49808007CBBBE6BC83B34FC0D" blockId="16.[136,342,897,920]" box="[136,342,897,920]" pageId="16" pageNumber="39">Female: Unknown.</paragraph>
<paragraph id="8BDA36A49808007CBBBE6B883F09FF4D" blockId="16.[136,775,961,1016]" lastBlockId="16.[835,1474,193,536]" pageId="16" pageNumber="39">
Measurements: BL =
<quantity id="4C9D9B419808007CBA476B883BB2FC4D" box="[369,464,961,984]" metricMagnitude="-3" metricUnit="m" metricValue="4.02" pageId="16" pageNumber="39" unit="mm" value="4.02">4.02 mm</quantity>
in
<typeStatus id="54DE88069808007CBAC56B883832FC4D" box="[499,592,961,984]" pageId="16" pageNumber="39" type="holotype">holotype</typeStatus>
(range in
<specimenCount id="9D63FD2D9808007CB9886B883965FC4D" box="[702,775,961,984]" pageId="16" pageNumber="39" type="male">3 male</specimenCount>
<typeStatus id="54DE88069808007CBBBE6BA83A90FC6D" box="[136,242,993,1016]" pageId="16" pageNumber="39" type="paratype">paratypes</typeStatus>
: 4.20-4.60); ThL = 2.00 mm (2.00-2.16); WL =
<quantity id="4C9D9B419808007CB875688839C1FF4D" box="[835,931,193,216]" metricMagnitude="-3" metricUnit="m" metricValue="3.12" pageId="16" pageNumber="39" unit="mm" value="3.12">3.12 mm</quantity>
(3.04-3.16); WW =
<quantity id="4C9D9B419808007CBFB268883E86FF4D" box="[1156,1252,193,216]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="16" pageNumber="39" unit="mm" value="1.32">1.32 mm</quantity>
(1.32-1.44).
</paragraph>
<paragraph id="8BDA36A49808007CB87568A83E72FD8D" blockId="16.[835,1474,193,536]" pageId="16" pageNumber="39">Indices: arb = 4-5/2-3 (5-6/2-3), avd = 0.50 (0.50-0.55), adf = 1.85 (1.80-1.90), flw = 1.50 (1.40-1.55), FW/HW = 0.40 (0.35-0.40), ch/o = 0.06 (0.06-0.07), prorb = 1.00 (1.05-1.10), rcorb = 0.50 (0.45-0.50), vb = 0.30 (0.35-0.40), dcl = 0.60 (0.60), presctl = 0.80 (0.70-0.75), sctl = 1.10 (1.05-1.10), sterno = 0.90 (0.90-0.95), orbito = 1.60 (1.55-1.65), dcp = 0.23 (0.22-0.23), sctlp = 1.10 (1.00-1.10), C = 2.19 (2.25- 2.37), 4c = 1.62 (1.43-1.46), 4v = 2.92 (2.67-2.86), 5x = 0.89 (0.78-0.80), ac = 4.20 (3.33-3.80), M = 0.62 (0.50-0.57), C3F = 0.71 (0.74-0.75).</paragraph>
</subSubSection>
<subSubSection id="C37F652F9808007CB8756A083FFCFDED" pageId="16" pageNumber="39" type="etymology">
<paragraph id="8BDA36A49808007CB8756A083FFCFDED" blockId="16.[835,1473,576,632]" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756A0839B3FDCD" bold="true" box="[835,977,576,601]" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756A0839DDFDCC" bold="true" box="[835,959,577,601]" italics="true" pageId="16" pageNumber="39">Etymology.</emphasis>
</emphasis>
From the Latin word “membranifer”, referring to the vertical process of gonopods partly membranous.
</paragraph>
</subSubSection>
<subSubSection id="C37F652F9808007CB8756AE83ED2FD2D" box="[835,1200,672,697]" pageId="16" pageNumber="39" type="distribution">
<paragraph id="8BDA36A49808007CB8756AE83ED2FD2D" blockId="16.[835,1200,672,697]" box="[835,1200,672,697]" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756AE839ADFD2D" box="[835,975,673,697]" italics="true" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756AE839AAFD2C" bold="true" box="[835,968,673,697]" italics="true" pageId="16" pageNumber="39">Distribution</emphasis>
.
</emphasis>
<emphasis id="B911EAB69808007CB8E16AE93986FD2D" bold="true" box="[983,996,672,696]" pageId="16" pageNumber="39"></emphasis>
<collectingCountry id="F37276349808007CB8DA6AE83E2CFD2D" box="[1004,1102,673,696]" name="Malaysia" pageId="16" pageNumber="39">Malaysia</collectingCountry>
(
<collectingRegion id="49A1F8469808007CBF686AE83EC7FD2D" box="[1118,1189,673,696]" country="Malaysia" name="Sabah" pageId="16" pageNumber="39">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652F9808007CB8756AA839B4FC6D" pageId="16" pageNumber="39" type="discussion">
<paragraph id="8BDA36A49808007CB8756AA839B4FC6D" blockId="16.[835,1474,736,1016]" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756AA839A4FD6D" bold="true" box="[835,966,736,761]" pageId="16" pageNumber="39">
<emphasis id="B911EAB69808007CB8756AA839CCFD6C" bold="true" box="[835,942,737,761]" italics="true" pageId="16" pageNumber="39">Remarks.</emphasis>
</emphasis>
This species is somewhat similar to
<emphasis id="B911EAB69808007CBE406AA83FE4FD6D" box="[1398,1414,737,760]" italics="true" pageId="16" pageNumber="39">P</emphasis>
.
<taxonomicName id="4C654D279808007CBEAE6AA83EF6FC8D" authority="(Toda &amp; Peng, 1990)" baseAuthorityName="Toda &amp; Peng" baseAuthorityYear="1990" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="lambda" subGenus="Phortica">
(
<emphasis id="B911EAB69808007CBE976AA83FD3FD6D" box="[1441,1457,737,760]" italics="true" pageId="16" pageNumber="39">P</emphasis>
.)
<emphasis id="B911EAB69808007CB8756B4839F7FC8D" box="[835,917,769,792]" italics="true" pageId="16" pageNumber="39">lambda</emphasis>
(Toda &amp; Peng, 1990)
</taxonomicName>
from
<collectingCountry id="F37276349808007CBFE96B483F42FC8D" box="[1247,1312,769,792]" name="China" pageId="16" pageNumber="39">China</collectingCountry>
in having the vertical process of gonopods partly sclerotized, but can be distinguished from it by the aedeagal median rod (in
<taxonomicName id="4C654D279808007CBE9C6B0839F1FCED" baseAuthorityName="Toda &amp; Peng" baseAuthorityYear="1990" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="lambda">
<emphasis id="B911EAB69808007CBE9C6B083FD8FCCD" box="[1450,1466,833,856]" italics="true" pageId="16" pageNumber="39">P</emphasis>
.
<emphasis id="B911EAB69808007CB8756B2839F1FCED" box="[835,915,865,888]" italics="true" pageId="16" pageNumber="39">lambda</emphasis>
</taxonomicName>
: ventral bridge between aedeagus and gonopods not reaching to tip of aedeagal median rod, without submedial branch; aedeagal median rod entirely sclerotized, with 1 process subapically and 1 pair of long lateral processes submedially).
</paragraph>
</subSubSection>
</treatment>
</document>