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<document id="F4DC579F7A713AA7A419BE9950B3FF72" ID-CLB-Dataset="39228" ID-DOI="10.11646/zootaxa.4059.1.10" ID-GBIF-Dataset="5d527dc2-b981-4d4f-8dc5-9ead824602db" ID-ISSN="1175-5326" ID-Zenodo-Dep="245490" ID-ZooBank="C785A607-FCCE-43E0-9F75-53E49ACFCB07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461361220935" checkinUser="plazi" docAuthor="Driauach, Ouafaa, Krzemińska, Ewa &amp; Belqat, Boutaïna" docDate="2015" docId="03CC87AAFF8EFFB7FF790011BE38F978" docLanguage="en" docName="zt04059p190.pdf" docOrigin="Zootaxa 4059 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichocera (Saltrichocera) pappi Krzeminska 2003" docType="treatment" docVersion="13" lastPageNumber="184" masterDocId="FFF5FFD2FF8DFFB4FFEE0350BC2AFFCB" masterDocTitle="Genus Trichocera in Morocco: first records from Africa and a new species (Diptera: Trichoceridae)" masterLastPageNumber="190" masterPageNumber="181" pageNumber="184" updateTime="1698661808762" updateUser="ExternalLinkService">
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<mods:title id="EECA0A267B9C8BB7478892B37C795FBD">Genus Trichocera in Morocco: first records from Africa and a new species (Diptera: Trichoceridae)</mods:title>
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<mods:namePart id="C8584A9FE9A712B1DE6F409C4030A64A">Driauach, Ouafaa</mods:namePart>
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<mods:namePart id="41D39E6B24DE58CFBEF7ECDD3AC47614">Krzemińska, Ewa</mods:namePart>
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<mods:namePart id="DFBD6B8F1766523118B21ACD23DD1770">Belqat, Boutaïna</mods:namePart>
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<subSubSection id="C37F6537FF8EFFB7FF790011BF27FC90" box="[151,781,833,859]" pageId="3" pageNumber="184" type="nomenclature">
<paragraph id="8BDA36BCFF8EFFB7FF790011BF27FC90" blockId="3.[151,781,833,859]" box="[151,781,833,859]" pageId="3" pageNumber="184">
<heading id="D09281D0FF8EFFB7FF790011BF27FC90" bold="true" box="[151,781,833,859]" fontSize="11" level="1" pageId="3" pageNumber="184" reason="1">
<taxonomicName id="4C654D3FFF8EFFB7FF790011BF27FC90" ID-CoL="8FH48" authority="Krzeminska, 2003" authorityName="Krzeminska" authorityYear="2003" box="[151,781,833,859]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi" subGenus="Saltrichocera">
<emphasis id="B911EAAEFF8EFFB7FF790011BF27FC90" bold="true" box="[151,781,833,859]" pageId="3" pageNumber="184">
<emphasis id="B911EAAEFF8EFFB7FF790011BE09FC90" bold="true" box="[151,547,833,859]" italics="true" pageId="3" pageNumber="184">Trichocera (Saltrichocera) pappi</emphasis>
<bibRefCitation id="EFF44B4DFF8EFFB7FDC40011BF27FC90" author="Krzeminska" box="[554,781,833,859]" pageId="3" pageNumber="184" refString="Krzeminska, E. (2003) A new species of Trichocera Meigen from Hungary and further remarks on the Hungarian Tichoceridae (Diptera: Trichoceridae). Folia entomologica hungarica, 64, 227 - 284." type="journal article" year="2003">Krzemińska, 2003</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BDA36BCFF8EFFB7FF7900D8BF11FC55" blockId="3.[151,827,904,926]" box="[151,827,904,926]" pageId="3" pageNumber="184">
<treatmentCitationGroup id="AB751192FF8EFFB7FF7900D8BF11FC55" box="[151,827,904,926]" pageId="3" pageNumber="184">
<taxonomicName id="4C654D3FFF8EFFB7FF7900D8BEFCFC55" ID-CoL="8FH48" authority="Krzeminska 2003: 278" authorityName="Krzeminska" authorityPageNumber="278" authorityYear="2003" box="[151,726,904,926]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi" subGenus="Saltrichocera">
<emphasis id="B911EAAEFF8EFFB7FF7900D8BDCDFC55" box="[151,487,904,926]" italics="true" pageId="3" pageNumber="184">Trichocera (Saltrichocera) pappi</emphasis>
<treatmentCitation id="0AC410ADFF8EFFB7FE0300D8BEFCFC55" author="Krzeminska" box="[493,726,904,926]" page="278" pageId="3" pageNumber="184" year="2003">
<bibRefCitation id="EFF44B4DFF8EFFB7FE0300D8BE8EFC55" author="Krzeminska" box="[493,676,904,926]" pageId="3" pageNumber="184" refString="Krzeminska, E. (2003) A new species of Trichocera Meigen from Hungary and further remarks on the Hungarian Tichoceridae (Diptera: Trichoceridae). Folia entomologica hungarica, 64, 227 - 284." type="journal article" year="2003">Krzemińska 2003</bibRefCitation>
: 278
</treatmentCitation>
</taxonomicName>
, fig. 111
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</paragraph>
</subSubSection>
<subSubSection id="C37F6537FF8EFFB7FF79009ABEDBFAED" pageId="3" pageNumber="184" type="materials_examined">
<paragraph id="8BDA36BCFF8EFFB7FF79009ABDE9FB84" blockId="3.[151,1437,970,1715]" pageId="3" pageNumber="184">
<emphasis id="B911EAAEFF8EFFB7FF79009ABD53FC28" bold="true" box="[151,377,970,995]" pageId="3" pageNumber="184">Material examined</emphasis>
.
<collectingCountry id="F372762CFF8EFFB7FE69009BBDDBFC29" box="[391,497,971,994]" name="Morocco" pageId="3" pageNumber="184">Morocco</collectingCountry>
: Seguia El Hajeb, 9.
<date id="FFDB107CFF8EFFB7FD3F009BBF79FC28" box="[721,851,971,995]" pageId="3" pageNumber="184" value="2011-12-01">
XII.2011,
<specimenCount id="9D63FD35FF8EFFB7FCA9009BBF79FC28" box="[839,851,971,995]" pageId="3" pageNumber="184" type="generic">1</specimenCount>
</date>
♂; Daya Fifi, 30.
<date id="FFDB107CFF8EFFB7FBFA009BB8A0FC28" box="[1044,1162,971,995]" pageId="3" pageNumber="184" value="2012-03-01">
III.2012,
<specimenCount id="9D63FD35FF8EFFB7FB90009BB8A0FC28" box="[1150,1162,971,995]" pageId="3" pageNumber="184" type="generic">1</specimenCount>
</date>
♂; Affluent Oued Akrir, 23.
<date id="FFDB107CFF8EFFB7FF2F00BFBD68FBCC" box="[193,322,1007,1031]" pageId="3" pageNumber="184" value="2012-11-01">XI. 2012, 1</date>
♀,
<specimenCount id="9D63FD35FF8EFFB7FE8B00BFBDADFBCC" box="[357,391,1007,1031]" pageId="3" pageNumber="184" type="generic">3♂</specimenCount>
; Cascade Grotte des Pigeons, 25.
<date id="FFDB107CFF8EFFB7FCEE00BEBFA8FBCC" box="[768,898,1006,1031]" pageId="3" pageNumber="184" value="2014-11-01">XI. 2014, 1</date>
♀,
<specimenCount id="9D63FD35FF8EFFB7FC4B00BFBFE2FBCD" box="[933,968,1007,1031]" pageId="3" pageNumber="184" type="generic">1♂</specimenCount>
; Oued Amsemlil,
<date id="FFDB107CFF8EFFB7FB7600BFB93DFBCC" box="[1176,1303,1006,1031]" pageId="3" pageNumber="184" value="2015-02-28">28.II. 2015</date>
, Site 2, 2♀,
<specimenCount id="9D63FD35FF8EFFB7FF790743BC93FBE0" box="[151,185,1043,1067]" pageId="3" pageNumber="184" type="generic">2♂</specimenCount>
; Site 3, 1♀,
<specimenCount id="9D63FD35FF8EFFB7FEBE0743BD5CFBE1" box="[336,374,1043,1067]" pageId="3" pageNumber="184" type="generic">4♂</specimenCount>
; Affluent Oued Amsemlil, 28.
<date id="FFDB107CFF8EFFB7FD3D0743BF61FBE0" box="[723,843,1042,1067]" pageId="3" pageNumber="184" value="2015-02-02">II. 2015, 2</date>
f,
<specimenCount id="9D63FD35FF8EFFB7FC8A0743BFA0FBE0" box="[868,906,1043,1067]" pageId="3" pageNumber="184" type="generic">4♂</specimenCount>
; Ruisseau Agoummir, 28.
<date id="FFDB107CFF8EFFB7FB5F0742B908FBE0" box="[1201,1314,1042,1067]" pageId="3" pageNumber="184" value="2015-02-03">
II.2015,
<specimenCount id="9D63FD35FF8EFFB7FAFA0743B908FBE0" box="[1300,1314,1043,1067]" pageId="3" pageNumber="184" type="generic">3</specimenCount>
</date>
♂; Maison forestière, 21.
<date id="FFDB107CFF8EFFB7FEC30766BD8EFB84" box="[301,420,1078,1103]" pageId="3" pageNumber="184" value="2015-04-01">
IV.2015,
<specimenCount id="9D63FD35FF8EFFB7FE760767BD8EFB84" box="[408,420,1079,1103]" pageId="3" pageNumber="184" type="generic">1</specimenCount>
</date>
♂.
</paragraph>
<paragraph id="8BDA36BCFF8EFFB7FF29070AB813FB5C" blockId="3.[151,1437,970,1715]" pageId="3" pageNumber="184">
<emphasis id="B911EAAEFF8EFFB7FF29070ABDFCFBB8" bold="true" box="[199,470,1114,1139]" pageId="3" pageNumber="184">Additional description</emphasis>
. Body colour dark brown to black. Wing membrane dark pigmented; venation very variable, regarding the shape of discal cell and proportions of veins R2+3+4/R2+3.
</paragraph>
<paragraph id="8BDA36BCFF8EFFB7FF2907F2BEDBFAED" blockId="3.[151,1437,970,1715]" pageId="3" pageNumber="184">Genitalia, female: The genital segment and the ovipositor are dark yellow, contrasting with dark colour of the remainder of abdomen. The apical portion of genital fork is characteristic, wider than long, bowl-shaped, with lateral portions slightly inclined to inside and more sclerotized (darker) than remainder. Some females bear four bristles on supragenital plate instead of the usual two.</paragraph>
</subSubSection>
<subSubSection id="C37F6537FF8EFFB7FF290662BE38F978" pageId="3" pageNumber="184" type="discussion">
<paragraph id="8BDA36BCFF8EFFB7FF290662B87DF9E9" blockId="3.[151,1437,970,1715]" pageId="3" pageNumber="184">
<emphasis id="B911EAAEFF8EFFB7FF290662BF5DFA80" bold="true" box="[199,887,1330,1355]" pageId="3" pageNumber="184">
Remarks on discerning
<taxonomicName id="4C654D3FFF8EFFB7FE000663BEB0FA80" box="[494,666,1330,1355]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saltator" subGenus="Saltrichocera">
<emphasis id="B911EAAEFF8EFFB7FE000663BDD7FA81" bold="true" box="[494,509,1331,1354]" italics="true" pageId="3" pageNumber="184">T</emphasis>
. (
<emphasis id="B911EAAEFF8EFFB7FDF60662BE0DFA80" bold="true" box="[536,551,1330,1355]" italics="true" pageId="3" pageNumber="184">S</emphasis>
.)
<emphasis id="B911EAAEFF8EFFB7FDAD0662BEB0FA80" bold="true" box="[579,666,1330,1355]" italics="true" pageId="3" pageNumber="184">saltator</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D3FFF8EFFB7FD320663BF5BFA80" box="[732,881,1330,1355]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi" subGenus="Saltrichocera">
<emphasis id="B911EAAEFF8EFFB7FD320663BEC1FA81" bold="true" box="[732,747,1331,1354]" italics="true" pageId="3" pageNumber="184">T</emphasis>
. (
<emphasis id="B911EAAEFF8EFFB7FCE80662BF3FFA80" bold="true" box="[774,789,1330,1355]" italics="true" pageId="3" pageNumber="184">S</emphasis>
.)
<emphasis id="B911EAAEFF8EFFB7FCDF0662BF5BFA80" bold="true" box="[817,881,1330,1355]" italics="true" pageId="3" pageNumber="184">pappi</emphasis>
</taxonomicName>
.
</emphasis>
Males of the two species are more difficult to discern in the Moroccan localities than in Europe.
<taxonomicName id="4C654D3FFF8EFFB7FD360607BF1FFAA5" box="[728,821,1367,1390]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi">
<emphasis id="B911EAAEFF8EFFB7FD360607BF1FFAA5" box="[728,821,1367,1390]" italics="true" pageId="3" pageNumber="184">T. pappi</emphasis>
</taxonomicName>
is most often as black as
<taxonomicName id="4C654D3FFF8EFFB7FBB00607B8FBFAA4" box="[1118,1233,1366,1391]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAAEFF8EFFB7FBB00607B8FBFAA4" box="[1118,1233,1366,1391]" italics="true" pageId="3" pageNumber="184">T. saltator</emphasis>
</taxonomicName>
, and the shape of gonostyles and bridge are in many specimens very similar. When in doubt, the males were discerned by ventral setation of vein Sc, which in
<taxonomicName id="4C654D3FFF8EFFB7FE0206CFBE74FA7C" box="[492,606,1438,1463]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAAEFF8EFFB7FE0206CFBE74FA7C" box="[492,606,1438,1463]" italics="true" pageId="3" pageNumber="184">T. saltator</emphasis>
</taxonomicName>
begins in the distal half of Sc, and is scarce (up to c. 20 setae grouped mainly in distal section). In
<taxonomicName id="4C654D3FFF8EFFB7FE210694BE00FA10" box="[463,554,1476,1499]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi">
<emphasis id="B911EAAEFF8EFFB7FE210694BE00FA10" box="[463,554,1476,1499]" italics="true" pageId="3" pageNumber="184">T. pappi</emphasis>
</taxonomicName>
the setae begin in c. of Sc, and are more dense (3060), especially in mid vein. In both species the vein Rs has setae, contrary to the results of
<bibRefCitation id="EFF44B4DFF8EFFB7FC8906B7B802FA34" author="Lawrence" box="[871,1064,1510,1535]" pageId="3" pageNumber="184" refString="Lawrence, B. R. (1957) The British species of Trichocera (Diptera: Trichoceridae). Proceedings of the Royal Entomological Society of London, (A), 32, 132 - 138." type="journal article" year="1957">Lawrence (1957)</bibRefCitation>
recorded for
<taxonomicName id="4C654D3FFF8EFFB7FB2D06B7B91DFA34" box="[1219,1335,1510,1535]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAAEFF8EFFB7FB2D06B7B91DFA34" box="[1219,1335,1510,1535]" italics="true" pageId="3" pageNumber="184">T. saltator</emphasis>
</taxonomicName>
in
<collectingCountry id="F372762CFF8EFFB7FAB006B7BCC2F9E8" name="United Kingdom" pageId="3" pageNumber="184">Great Britain</collectingCountry>
. Additionally, the hind tarsal claw in male is longer and stronger in
<taxonomicName id="4C654D3FFF8EFFB7FC31055CB87AF9E8" box="[991,1104,1546,1571]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAAEFF8EFFB7FC31055CBFC6F9E8" box="[991,1004,1548,1571]" italics="true" pageId="3" pageNumber="184">T</emphasis>
.
<emphasis id="B911EAAEFF8EFFB7FC15055AB87AF9E8" box="[1019,1104,1546,1571]" italics="true" pageId="3" pageNumber="184">saltator</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA36BCFF8EFFB7FF29057EBE38F978" blockId="3.[151,1437,970,1715]" pageId="3" pageNumber="184">
Females of the two species are easily distinguishable by the shape of the ovipositor and genital plate, although remarkably some females of
<taxonomicName id="4C654D3FFF8EFFB7FE370504BE1EF9A0" box="[473,564,1620,1643]" class="Insecta" family="Trichoceridae" genus="Trichocera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="pappi">
<emphasis id="B911EAAEFF8EFFB7FE370504BE1EF9A0" box="[473,564,1620,1643]" italics="true" pageId="3" pageNumber="184">T. pappi</emphasis>
</taxonomicName>
have 46 bristles on supragenital plate, while other have usual two. This latter finding, and the unusual similarity of some males may indicate casual interbreeding between the two species living close to their distributional limits.
</paragraph>
</subSubSection>
</treatment>
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