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<document id="AF3B07EFBA63AA9A3BD2C914530EAB55" ID-DOI="10.5281/zenodo.185202" ID-GBIF-Dataset="4bcdebee-203f-4c45-8904-026876fc4a08" ID-ISSN="1175-5326" ID-Zenodo-Dep="185202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459969944348" checkinUser="plazi" docAuthor="Ng, Heok Hee" docDate="2009" docId="03CC879A820AFFF4FF1BA1D2B14BFB2E" docLanguage="en" docName="zt01977p048.pdf" docOrigin="Zootaxa 1977" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Parakysis hystriculus Ng, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFF5FFE2820BFFF0FF8CA145B32CFF95" masterDocTitle="Parakysis hystriculus, a new species of catfish (Siluriformes: Akysidae) from Sumatra" masterLastPageNumber="48" masterPageNumber="39" pageNumber="40" updateTime="1698231362669" updateUser="plazi">
<mods:mods id="E1F2FABD89E1CAD0A584611485EA2BCD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="984EB1F42C5C43DEED3A532E4A0AB7BB">
<mods:title id="3225A439EB2890789F89F4C6C2A8E2BA">Parakysis hystriculus, a new species of catfish (Siluriformes: Akysidae) from Sumatra</mods:title>
</mods:titleInfo>
<mods:name id="90063D67BDC8F3F5CC929650BCE8A296" type="personal">
<mods:role id="C9B603B8741F99AC0607045CF953C1AE">
<mods:roleTerm id="5AA42CE4AE1357C279B5973CA04D6A86">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD679EDC81BE2019B16227B1396FF446">Ng, Heok Hee</mods:namePart>
</mods:name>
<mods:typeOfResource id="8243E4015081088DC787DB9302A60961">text</mods:typeOfResource>
<mods:relatedItem id="4DA902EC2EB115D395A169A15B8567FF" type="host">
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<mods:title id="E26F979F04216F34F88374C9B6EEB73C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7CF0A7378512AEA9F6408D6DC1D848A5">
<mods:date id="5CCFF526A667EBAA8698B3CC1ED11D68">2009</mods:date>
<mods:detail id="71D7B9E302C25DF9E3E416C6ED786035" type="volume">
<mods:number id="7B7E168062BFDE5B347D4E205B570546">1977</mods:number>
</mods:detail>
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<mods:classification id="7B57BA5B8BBB50EEAE365EF26087469F">journal article</mods:classification>
<mods:identifier id="D426E42BD4D13E2E59F8D723F913D536" type="DOI">10.5281/zenodo.185202</mods:identifier>
<mods:identifier id="41D6A55FCBAC3A402377BABA7420E4FA" type="GBIF-Dataset">4bcdebee-203f-4c45-8904-026876fc4a08</mods:identifier>
<mods:identifier id="D5363C98390874B000DB8A1B310AF5DB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="74109FBADE528D3EB25F4D951A0C725B" type="Zenodo-Dep">185202</mods:identifier>
</mods:mods>
<treatment id="03CC879A820AFFF4FF1BA1D2B14BFB2E" ID-DOI="http://doi.org/10.5281/zenodo.5696978" ID-GBIF-Taxon="119377777" ID-Zenodo-Dep="5696978" LSID="urn:lsid:plazi:treatment:03CC879A820AFFF4FF1BA1D2B14BFB2E" httpUri="http://treatment.plazi.org/id/03CC879A820AFFF4FF1BA1D2B14BFB2E" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40">
<subSubSection id="C37F6507820AFFF1FF1BA1D2B28FFF41" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="8BDA368C820AFFF1FF1BA1D2B128FF24" blockId="1.[151,516,151,212]" box="[151,516,151,177]" pageId="1" pageNumber="40">
<heading id="D09281E0820AFFF1FF1BA1D2B128FF24" bold="true" box="[151,516,151,177]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
<emphasis id="B911EA9E820AFFF1FF1BA1D2B128FF24" bold="true" box="[151,516,151,177]" pageId="1" pageNumber="40">
<taxonomicName id="4C654D0F820AFFF1FF1BA1D2B2B4FF24" ID-CoL="4CXNG" box="[151,408,151,177]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="hystriculus" status="sp. nov.">
<emphasis id="B911EA9E820AFFF1FF1BA1D2B2B4FF24" bold="true" box="[151,408,151,177]" italics="true" pageId="1" pageNumber="40">Parakysis hystriculus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22257E5820AFFF1FE2BA1D2B128FF24" box="[423,516,151,177]" pageId="1" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA368C820AFFF1FF1BA1FFB28FFF41" blockId="1.[151,516,151,212]" box="[151,419,186,212]" pageId="1" pageNumber="40">
(
<figureCitation id="135E2A09820AFFF1FF13A1FFB3DDFF41" box="[159,241,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,255,2005,2029]" captionTargetBox="[157,1436,464,1995]" captionTargetId="figure@2.[157,1442,464,1996]" captionTargetPageId="2" captionText="FIGURE 1. Parakysis hystriculus, MZB 10985, holotype, 27.6 mm SL; Sumatra: swamp forest at Sentang." httpUri="https://zenodo.org/record/185203/files/figure.png" pageId="1" pageNumber="40">Figs. 1</figureCitation>
,
<figureCitation id="135E2A09820AFFF1FF73A1FFB223FF41" box="[255,271,186,212]" captionStart="FIGURE 2" captionStartId="4.[151,255,963,987]" captionTargetBox="[151,1435,194,939]" captionTargetId="figure@4.[151,1436,194,939]" captionTargetPageId="4" captionText="FIGURE 2. Posterior portion of bodies in: a. Parakysis hystriculus, ZRC 51418, male paratype, 26.0 mm SL (above) and ZRC 51418, female paratype, 25.7 mm SL (below); b. P. grandis, ZRC 38632, male paratype, 24.2 mm SL (above) and ZRC 39178, female paratype, 29.3 mm SL (below), showing difference in anal-fin shape. Images not to scale." httpUri="https://zenodo.org/record/185204/files/figure.png" pageId="1" pageNumber="40">2</figureCitation>
a, 4a, 5a, 6a)
</paragraph>
</subSubSection>
<subSubSection id="C37F6507820AFFF1FF1BA04FB1FAFE79" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="8BDA368C820AFFF1FF1BA04FB0EDFEE1" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">
<emphasis id="B911EA9E820AFFF1FF1BA04FB261FEB1" bold="true" box="[151,333,266,292]" pageId="1" pageNumber="40">
<typeStatus id="54DE882E820AFFF1FF1BA04FB3FEFEB1" box="[151,210,266,292]" pageId="1" pageNumber="40">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DE882E820AFFF1FED5A04FB2E6FEB1" box="[345,458,266,292]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED74AE49820AFFF1FE54A04FB134FEB1" box="[472,536,266,292]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="40">MZB</collectionCode>
10985,
<quantity id="4C9D9B69820AFFF1FDF0A04FB1CBFEB1" box="[636,743,266,292]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="1" pageNumber="40" unit="mm" value="27.6">27.6 mm</quantity>
<collectionCode id="ED74AE49820AFFF1FD7CA04FB03AFEB1" box="[752,790,266,292]" pageId="1" pageNumber="40">SL</collectionCode>
; Sumatra: Sumatera Selatan, Lalang drainage, swamp forest at Sentang, ca.
<quantity id="4C9D9B69820AFFF1FE24A077B2C4FED9" box="[424,488,306,332]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="40" unit="km" value="5.0">5 km</quantity>
after turnoff ca.
<quantity id="4C9D9B69820AFFF1FD4EA077B03EFED9" box="[706,786,306,332]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="40" unit="km" value="12.0">12 km</quantity>
after Bayung Lencir on Bayung Lencir–Jambi road,
<geoCoordinate id="EE51504B820AFFF1FF1BA01FB23BFEE1" box="[151,279,346,372]" direction="south" orientation="latitude" pageId="1" pageNumber="40" precision="1" value="-1.9365333">1°56.192'S</geoCoordinate>
<geoCoordinate id="EE51504B820AFFF1FE93A01FB293FEE1" box="[287,447,346,372]" direction="east" orientation="longitude" pageId="1" pageNumber="40" precision="1" value="103.70887">103º42.532'E</geoCoordinate>
; M. Kottelat &amp; H. H. Ng,
<date id="FFDB104C820AFFF1FD7EA01FB091FEE1" box="[754,957,346,372]" pageId="1" pageNumber="40" value="2008-05-17" valueMax="2008-05-20" valueMin="2008-05-17">17–20 May 2008</date>
.
</paragraph>
<paragraph id="8BDA368C820AFFF1FF49A0C7B1FAFE79" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">
<emphasis id="B911EA9E820AFFF1FF49A0C7B260FE09" bold="true" box="[197,332,386,412]" pageId="1" pageNumber="40">
<typeStatus id="54DE882E820AFFF1FF49A0C7B26AFE09" box="[197,326,386,412]" pageId="1" pageNumber="40" type="paratype">Paratypes</typeStatus>
:
</emphasis>
CMK 20676 (10),
<quantity id="4C9D9B69820AFFF1FDBEA0C7B1F1FE09" box="[562,733,386,412]" metricMagnitude="-2" metricUnit="m" metricValue="2.575" metricValueMax="2.76" metricValueMin="2.3899999999999997" pageId="1" pageNumber="40" unit="mm" value="25.75" valueMax="27.6" valueMin="23.9">23.9–27.6 mm</quantity>
<collectionCode id="ED74AE49820AFFF1FD69A0C7B027FE09" box="[741,779,386,412]" pageId="1" pageNumber="40">SL</collectionCode>
;
<collectionCode id="ED74AE49820AFFF1FC94A0C7B07DFE09" box="[792,849,386,412]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
51418 (9), 23.0–28.2; data as for
<typeStatus id="54DE882E820AFFF1FB6AA0C7B67CFE09" box="[1254,1360,386,412]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED74AE49820AFFF1FAD0A0C7B6B0FE09" box="[1372,1436,386,412]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="40">MZB</collectionCode>
10986 (10),
<quantity id="4C9D9B69820AFFF1FEABA0EFB2FDFE51" box="[295,465,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="2.58" metricValueMin="2.07" pageId="1" pageNumber="40" unit="mm" value="23.25" valueMax="25.8" valueMin="20.7">20.7–25.8 mm</quantity>
<collectionCode id="ED74AE49820AFFF1FE54A0EFB2D2FE51" box="[472,510,426,452]" pageId="1" pageNumber="40">SL</collectionCode>
;
<collectionCode id="ED74AE49820AFFF1FD86A0EFB149FE51" box="[522,613,426,452]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="1" pageNumber="40">UMMZ</collectionCode>
248763 (12),
<quantity id="4C9D9B69820AFFF1FC80A0EFB09AFE51" box="[780,950,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="2.485" metricValueMax="2.71" metricValueMin="2.26" pageId="1" pageNumber="40" unit="mm" value="24.85" valueMax="27.1" valueMin="22.6">22.6–27.1 mm</quantity>
<collectionCode id="ED74AE49820AFFF1FC31A0EFB0CFFE51" box="[957,995,426,452]" pageId="1" pageNumber="40">SL</collectionCode>
;
<collectionCode id="ED74AE49820AFFF1FC63A0EFB704FE51" box="[1007,1064,426,452]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="40">ZRC</collectionCode>
43088 (68),
<quantity id="4C9D9B69820AFFF1FB4CA0EFB646FE51" box="[1216,1386,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="2.3299999999999996" metricValueMax="2.92" metricValueMin="1.7399999999999998" pageId="1" pageNumber="40" unit="mm" value="23.299999999999997" valueMax="29.2" valueMin="17.4">17.4–29.2 mm</quantity>
<collectionCode id="ED74AE49820AFFF1FAFDA0EFB6BBFE51" box="[1393,1431,426,452]" pageId="1" pageNumber="40">SL</collectionCode>
; locality as for
<typeStatus id="54DE882E820AFFF1FEB3A097B285FE79" box="[319,425,466,492]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
; H. H. Tan,
<date id="FFDB104C820AFFF1FDBBA097B1FEFE79" box="[567,722,466,492]" pageId="1" pageNumber="40" value="1997-07-27">27 July 1997</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6507820AFFF1FF49A0BFB295FC31" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="8BDA368C820AFFF1FF49A0BFB295FC31" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">
<emphasis id="B911EA9E820AFFF1FF49A0BFB269FD81" bold="true" box="[197,325,506,532]" pageId="1" pageNumber="40">Diagnosis.</emphasis>
<taxonomicName id="4C654D0F820AFFF1FEC1A0BFB166FD86" box="[333,586,506,531]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="hystriculus">
<emphasis id="B911EA9E820AFFF1FEC1A0BFB166FD86" box="[333,586,506,531]" italics="true" pageId="1" pageNumber="40">Parakysis hystriculus</emphasis>
</taxonomicName>
is distinguished from other members of the genus in having a combination of: branched pelvic-fin rays with anterior branch always shorter than posterior branch (vs. almost same length in congeners), lateral edges of head evenly sloping in dorsal view (vs. gently curving in congeners), anteriormost pair of accessory inner mandibular barbels longer than (vs. equal to or shorter than in congeners) distance separating bases of left and right barbel, outer mandibular barbel with one accessory barbel (vs. absent in
<taxonomicName id="4C654D0F820AFFF1FE8BA387B2F8FD4E" box="[263,468,706,731]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="anomalopteryx">
<emphasis id="B911EA9E820AFFF1FE8BA387B2F8FD4E" box="[263,468,706,731]" italics="true" pageId="1" pageNumber="40">P. anomalopteryx</emphasis>
</taxonomicName>
), mandibular laterosensory pore between bases of inner and outer mandibular barbels present (vs. absent in
<taxonomicName id="4C654D0F820AFFF1FE44A3AFB1B8FC96" box="[456,660,746,771]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="anomalopteryx">
<emphasis id="B911EA9E820AFFF1FE44A3AFB1B8FC96" box="[456,660,746,771]" italics="true" pageId="1" pageNumber="40">P. anomalopteryx</emphasis>
</taxonomicName>
), margin of lower lip with medial concavity (vs. entire in
<taxonomicName id="4C654D0F820AFFF1FAB9A3AFB22AFCBE" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="anomalopteryx">
<emphasis id="B911EA9E820AFFF1FAB9A3AFB22AFCBE" italics="true" pageId="1" pageNumber="40">P. anomalopteryx</emphasis>
</taxonomicName>
), anal fin rounded (vs. polygonal or rectangular in
<taxonomicName id="4C654D0F820AFFF1FCEFA257B0F6FCBE" box="[867,986,786,811]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B911EA9E820AFFF1FCEFA257B0F6FCBE" box="[867,986,786,811]" italics="true" pageId="1" pageNumber="40">P. grandis</emphasis>
</taxonomicName>
), caudal fin with narrow lobes having evenly-tapering posteromedial margins (vs. broad lobes with curved posteromedial margins in
<taxonomicName id="4C654D0F820AFFF1FB7FA27FB646FCC6" box="[1267,1386,826,851]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B911EA9E820AFFF1FB7FA27FB646FCC6" box="[1267,1386,826,851]" italics="true" pageId="1" pageNumber="40">P. grandis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D0F820AFFF1FF1BA227B217FCEE" box="[151,315,866,891]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="longirostris">
<emphasis id="B911EA9E820AFFF1FF1BA227B217FCEE" box="[151,315,866,891]" italics="true" pageId="1" pageNumber="40">P. longirostris</emphasis>
</taxonomicName>
) and 10 (vs.
<quantity id="4C9D9B69820AFFF1FE5FA227B121FCE9" box="[467,525,866,892]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="40" unit="in" value="11.0">11 in</quantity>
<taxonomicName id="4C654D0F820AFFF1FD98A227B19CFCEE" box="[532,688,866,891]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="verrucosus">
<emphasis id="B911EA9E820AFFF1FD98A227B19CFCEE" box="[532,688,866,891]" italics="true" pageId="1" pageNumber="40">P. verrucosus</emphasis>
</taxonomicName>
) branched rays without secondary branches (vs. with secondary branching in
<taxonomicName id="4C654D0F820AFFF1FEBDA2CFB28BFC36" box="[305,423,906,931]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="40" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B911EA9E820AFFF1FEBDA2CFB28BFC36" box="[305,423,906,931]" italics="true" pageId="1" pageNumber="40">P. grandis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6507820AFFF3FF49A2F7B7F9F83A" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40" type="description">
<paragraph id="8BDA368C820AFFF1FF49A2F7B71FFA99" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">
<emphasis id="B911EA9E820AFFF1FF49A2F7B271FC59" bold="true" box="[197,349,946,972]" pageId="1" pageNumber="40">Description.</emphasis>
Morphometric data are in
<tableCitation id="C6E70337820AFFF1FD25A2F7B029FC59" box="[681,773,946,972]" captionStart="TABLE 1" captionStartId="3.[151,239,471,495]" captionTargetBox="[159,1299,528,1660]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data for Parakysis hystriculus (n = 16)." httpUri="http://table.plazi.org/id/DF1A66048208FFF3FF1BA092B069FE7A" pageId="1" pageNumber="40" tableUuid="DF1A66048208FFF3FF1BA092B069FE7A">Table 1</tableCitation>
. Body slightly compressed, becoming more so toward caudal peduncle. Dorsal profile rising evenly but not steeply from tip of snout to origin of dorsal fin and sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile horizontal to posterior end of anal fin base, then sloping very gently dorsally from there to end of caudal peduncle. Skin rugose, covered with evenly distributed, rounded tubercles of uniform size. Lateral line absent, with postcranial laterosensory system on body consisting only of single pore on lateral surface of body immediately posterior to vertical through base of last pectoral-fin ray. Mandibular laterosensory canals with two pairs of pores: one medial to base of inner mandibular barbel and another between bases of inner and outer mandibular barbels. Vertebrae 18+12=30 (1), 19+12=31 (5), 19+13=32 (3), 20+12=32* (6), or 19+14=33 (1).
</paragraph>
<paragraph id="8BDA368C820AFFF1FF49A45FB718FA39" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">Head slightly depressed, dorsal surface covered with thick, rugose skin; dorsal profile convex; lateral margins of head straight in dorsal view, making head look broadly triangular. Eye very small and ovoid, subcutaneous, horizontal axis longest; located entirely in dorsal half of head. Gill openings restricted, extending from just above to just below cleithral process. Gill membranes united with isthmus.</paragraph>
<paragraph id="8BDA368C820AFFF1FF49A4FFB2FBF9D9" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">Mouth subterminal, fleshy upper lip with anterior medial lobe projecting anteriorly; lower lip with medial concavity in outline through which lobe on upper lip is visible. Oral teeth small, villiform, and in irregular rows on all tooth-bearing surfaces. Premaxillary and dentary tooth bands narrow and of equal width throughout. Vomerine teeth absent.</paragraph>
<paragraph id="8BDA368C820AFFF1FF49A71FB187F821" blockId="1.[151,1437,266,1972]" pageId="1" pageNumber="40">Barbels in four pairs. Nasal barbel slender, extending to two thirds of distance between its base and anterior base of dorsal spine. Maxillary barbel long and slender, with broad base originating at rictus of jaws and continuous with both upper and lower lips; extending to vertical through anterior base of dorsal spine. Inner mandibular barbel origin close to midline; barbel thicker and longer than nasal barbel and extending to horizontal line through anterior bases of pectoral spine. Base of inner mandibular barbel with two accessory barbels (branches) not confluent at base; anterior accessory barbel longer than distance between bases of left and right barbel. Outer mandibular barbel origin close to midline of ventral surface of head, thicker and longer than nasal barbel, and extending to vertical line through bases of posteriormost pectoral fin ray. Base of outer mandibular barbel with one accessory barbel.</paragraph>
<paragraph id="8BDA368C8209FFF3FF49A1DDB3D8FE31" blockId="2.[151,1436,152,418]" lastBlockId="3.[151,1437,152,420]" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
Dorsal fin located above middle of body; origin nearer to tip of snout than to caudal flexure. Dorsal fin margin convex, with I,3,i (30) rays; spine short and robust, lacking serrations on posterior edge. Pectoral fin with I,5 (2), I,5,i* (24), I,6 (3) or I,6,i (1) rays; branched fin rays with anterior branches shorter than posterior branches. Pectoral-fin spine short and stout; anterior and posterior spine margins smooth and without serrations. Pectoral fin margin straight anteriorly, convex posteriorly. Pelvic fin origin at vertical through posterior end of dorsal fin base; with i,4,i (30) rays. Posterior branch of branched pelvic-fin rays markedly longer than anterior branch; tip of adpressed fin not reaching anal fin origin. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Long dorsomedian adipose ridge originating just posterior to dorsal fin and confluent with upper procurrent caudal fin rays. Anal fin base ventral to posterior half of adipose fin; fin with iii,3,i (3), iii,4,i *(8), iii,5 (1), iv,3,I (1), iv, 4 (4), iv,4,i (8), iii,5,i (3), iii,6 (1) or iv,5 (1) rays. Anal-fin shape sexually dimorphic: fin in males more ventroposteriorly produced than in females (
<figureCitation id="135E2A098208FFF3FC3CA047B0DDFE8C" box="[944,1009,258,281]" captionStart="FIGURE 2" captionStartId="4.[151,255,963,987]" captionTargetBox="[151,1435,194,939]" captionTargetId="figure@4.[151,1436,194,939]" captionTargetPageId="4" captionText="FIGURE 2. Posterior portion of bodies in: a. Parakysis hystriculus, ZRC 51418, male paratype, 26.0 mm SL (above) and ZRC 51418, female paratype, 25.7 mm SL (below); b. P. grandis, ZRC 38632, male paratype, 24.2 mm SL (above) and ZRC 39178, female paratype, 29.3 mm SL (below), showing difference in anal-fin shape. Images not to scale." httpUri="https://zenodo.org/record/185204/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
a). Caudal peduncle moderately slender. Caudal fin deeply forked, with i,6,4,i (30) principal rays; upper and lower lobes pointed. Procurrent rays symmetrical, extending only slightly anterior to fin base. Mature male with long, conical genital papilla located immediately posterior to anus. Immature males with shorter papilla. Female lacking genital papilla, but with short, tubular extension of genital opening.
</paragraph>
<caption id="DF1A66048209FFF2FF1BA690B63FF878" box="[151,1299,2005,2029]" httpUri="https://zenodo.org/record/185203/files/figure.png" pageId="2" pageNumber="41" targetBox="[157,1436,464,1995]" targetPageId="2">
<paragraph id="8BDA368C8209FFF2FF1BA690B63FF878" blockId="2.[151,1299,2005,2029]" box="[151,1299,2005,2029]" pageId="2" pageNumber="41">
<emphasis id="B911EA9E8209FFF2FF1BA690B235F878" bold="true" box="[151,281,2005,2029]" pageId="2" pageNumber="41">FIGURE 1.</emphasis>
<taxonomicName id="4C654D0F8209FFF2FEACA693B129F878" box="[288,517,2006,2029]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="hystriculus">
<emphasis id="B911EA9E8209FFF2FEACA693B129F878" box="[288,517,2006,2029]" italics="true" pageId="2" pageNumber="41">Parakysis hystriculus</emphasis>
</taxonomicName>
, MZB 10985, holotype, 27.6 mm SL; Sumatra: swamp forest at Sentang.
</paragraph>
</caption>
<caption id="DF1A66048208FFF3FF1BA092B069FE7A" ID-Table-UUID="DF1A66048208FFF3FF1BA092B069FE7A" box="[151,837,471,495]" httpUri="http://table.plazi.org/id/DF1A66048208FFF3FF1BA092B069FE7A" pageId="3" pageNumber="42" targetBox="[159,1299,528,1660]" targetIsTable="true" targetPageId="3">
<paragraph id="8BDA368C8208FFF3FF1BA092B069FE7A" blockId="3.[151,837,471,495]" box="[151,837,471,495]" pageId="3" pageNumber="42">
<emphasis id="B911EA9E8208FFF3FF1BA092B224FE7A" bold="true" box="[151,264,471,495]" pageId="3" pageNumber="42">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="4C654D0F8208FFF3FD86A09DB1C3FE7A" box="[522,751,472,495]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="hystriculus">
<emphasis id="B911EA9E8208FFF3FD86A09DB1C3FE7A" box="[522,751,472,495]" italics="true" pageId="3" pageNumber="42">Parakysis hystriculus</emphasis>
</taxonomicName>
(n=16).
</paragraph>
</caption>
<paragraph id="8BDA368C8208FFF3FD12A355B63FF9E9" pageId="3" pageNumber="42">
<table id="F965C42C8208000FFF13A355B63FF9E9" box="[159,1299,528,1660]" gridcols="4" gridrows="27" pageId="3" pageNumber="42">
<tr id="355534CE8208000FFF13A355B63FFDB2" box="[159,1299,528,551]" gridrow="0" pageId="3" pageNumber="42" rowspan-0="1">
<th id="76845DB28208000FFD12A355B02DFDB2" box="[670,769,528,551]" gridcol="1" gridrow="0" pageId="3" pageNumber="42">Holotype</th>
<th id="76845DB28208000FFCF0A355B72CFDB2" box="[892,1024,528,551]" gridcol="2" gridrow="0" pageId="3" pageNumber="42">Range</th>
<th id="76845DB28208000FFB00A355B63FFDB2" box="[1164,1299,528,551]" gridcol="3" gridrow="0" pageId="3" pageNumber="42">Mean±SD</th>
</tr>
<tr id="355534CE8208000FFF13A37FB63FFDC4" box="[159,1299,570,593]" gridrow="1" pageId="3" pageNumber="42" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76845DB28208000FFF13A37FB2C2FDC4" box="[159,494,570,593]" gridcol="0" gridrow="1" pageId="3" pageNumber="42">%SL</th>
</tr>
<tr id="355534CE8208000FFF13A320B63FFDE9" box="[159,1299,613,636]" gridrow="2" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A320B2C2FDE9" box="[159,494,613,636]" gridcol="0" gridrow="2" pageId="3" pageNumber="42">Predorsal length</th>
<td id="76845DB28208000FFD12A320B02DFDE9" box="[670,769,613,636]" gridcol="1" gridrow="2" pageId="3" pageNumber="42">34.8</td>
<td id="76845DB28208000FFCF0A320B72CFDE9" box="[892,1024,613,636]" gridcol="2" gridrow="2" pageId="3" pageNumber="42">33.5–37.6</td>
<td id="76845DB28208000FFB00A320B63FFDE9" box="[1164,1299,613,636]" gridcol="3" gridrow="2" pageId="3" pageNumber="42">35.6±1.52</td>
</tr>
<tr id="355534CE8208000FFF13A3D5B63FFD32" box="[159,1299,656,679]" gridrow="3" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A3D5B2C2FD32" box="[159,494,656,679]" gridcol="0" gridrow="3" pageId="3" pageNumber="42">Preanal length</th>
<td id="76845DB28208000FFD12A3D5B02DFD32" box="[670,769,656,679]" gridcol="1" gridrow="3" pageId="3" pageNumber="42">71.0</td>
<td id="76845DB28208000FFCF0A3D5B72CFD32" box="[892,1024,656,679]" gridcol="2" gridrow="3" pageId="3" pageNumber="42">69.4–75.0</td>
<td id="76845DB28208000FFB00A3D5B63FFD32" box="[1164,1299,656,679]" gridcol="3" gridrow="3" pageId="3" pageNumber="42">72.6±1.95</td>
</tr>
<tr id="355534CE8208000FFF13A3FFB63FFD44" box="[159,1299,698,721]" gridrow="4" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A3FFB2C2FD44" box="[159,494,698,721]" gridcol="0" gridrow="4" pageId="3" pageNumber="42">Prepelvic length</th>
<td id="76845DB28208000FFD12A3FFB02DFD44" box="[670,769,698,721]" gridcol="1" gridrow="4" pageId="3" pageNumber="42">49.3</td>
<td id="76845DB28208000FFCF0A3FFB72CFD44" box="[892,1024,698,721]" gridcol="2" gridrow="4" pageId="3" pageNumber="42">47.7–53.6</td>
<td id="76845DB28208000FFB00A3FFB63FFD44" box="[1164,1299,698,721]" gridcol="3" gridrow="4" pageId="3" pageNumber="42">50.9±2.11</td>
</tr>
<tr id="355534CE8208000FFF13A3A0B63FFD69" box="[159,1299,741,764]" gridrow="5" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A3A0B2C2FD69" box="[159,494,741,764]" gridcol="0" gridrow="5" pageId="3" pageNumber="42">Prepectoral length</th>
<td id="76845DB28208000FFD12A3A0B02DFD69" box="[670,769,741,764]" gridcol="1" gridrow="5" pageId="3" pageNumber="42">17.0</td>
<td id="76845DB28208000FFCF0A3A0B72CFD69" box="[892,1024,741,764]" gridcol="2" gridrow="5" pageId="3" pageNumber="42">17.0–20.0</td>
<td id="76845DB28208000FFB00A3A0B63FFD69" box="[1164,1299,741,764]" gridcol="3" gridrow="5" pageId="3" pageNumber="42">18.6±0.95</td>
</tr>
<tr id="355534CE8208000FFF13A255B63FFCB2" box="[159,1299,784,807]" gridrow="6" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A255B2C2FCB2" box="[159,494,784,807]" gridcol="0" gridrow="6" pageId="3" pageNumber="42">Length of dorsal-fin base</th>
<td id="76845DB28208000FFD12A255B02DFCB2" box="[670,769,784,807]" gridcol="1" gridrow="6" pageId="3" pageNumber="42">10.5</td>
<td id="76845DB28208000FFCF0A255B72CFCB2" box="[892,1024,784,807]" gridcol="2" gridrow="6" pageId="3" pageNumber="42">9.4–14.1</td>
<td id="76845DB28208000FFB00A255B63FFCB2" box="[1164,1299,784,807]" gridcol="3" gridrow="6" pageId="3" pageNumber="42">11.3±1.42</td>
</tr>
<tr id="355534CE8208000FFF13A27FB63FFCC4" box="[159,1299,826,849]" gridrow="7" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A27FB2C2FCC4" box="[159,494,826,849]" gridcol="0" gridrow="7" pageId="3" pageNumber="42">Dorsal-spine length</th>
<td id="76845DB28208000FFD12A27FB02DFCC4" box="[670,769,826,849]" gridcol="1" gridrow="7" pageId="3" pageNumber="42">12.0</td>
<td id="76845DB28208000FFCF0A27FB72CFCC4" box="[892,1024,826,849]" gridcol="2" gridrow="7" pageId="3" pageNumber="42">7.9–14.5</td>
<td id="76845DB28208000FFB00A27FB63FFCC4" box="[1164,1299,826,849]" gridcol="3" gridrow="7" pageId="3" pageNumber="42">10.7±1.98</td>
</tr>
<tr id="355534CE8208000FFF13A220B63FFCE9" box="[159,1299,869,892]" gridrow="8" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A220B2C2FCE9" box="[159,494,869,892]" gridcol="0" gridrow="8" pageId="3" pageNumber="42">Length of anal-fin base</th>
<td id="76845DB28208000FFD12A220B02DFCE9" box="[670,769,869,892]" gridcol="1" gridrow="8" pageId="3" pageNumber="42">13.8</td>
<td id="76845DB28208000FFCF0A220B72CFCE9" box="[892,1024,869,892]" gridcol="2" gridrow="8" pageId="3" pageNumber="42">11.4–13.8</td>
<td id="76845DB28208000FFB00A220B63FFCE9" box="[1164,1299,869,892]" gridcol="3" gridrow="8" pageId="3" pageNumber="42">13.0±0.93</td>
</tr>
<tr id="355534CE8208000FFF13A2D5B63FFC32" box="[159,1299,912,935]" gridrow="9" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A2D5B2C2FC32" box="[159,494,912,935]" gridcol="0" gridrow="9" pageId="3" pageNumber="42">Pelvic-fin length</th>
<td id="76845DB28208000FFD12A2D5B02DFC32" box="[670,769,912,935]" gridcol="1" gridrow="9" pageId="3" pageNumber="42">17.4</td>
<td id="76845DB28208000FFCF0A2D5B72CFC32" box="[892,1024,912,935]" gridcol="2" gridrow="9" pageId="3" pageNumber="42">15.7–21.9</td>
<td id="76845DB28208000FFB00A2D5B63FFC32" box="[1164,1299,912,935]" gridcol="3" gridrow="9" pageId="3" pageNumber="42">17.7±2.18</td>
</tr>
<tr id="355534CE8208000FFF13A2FFB63FFC44" box="[159,1299,954,977]" gridrow="10" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A2FFB2C2FC44" box="[159,494,954,977]" gridcol="0" gridrow="10" pageId="3" pageNumber="42">Pectoral-fin length</th>
<td id="76845DB28208000FFD12A2FFB02DFC44" box="[670,769,954,977]" gridcol="1" gridrow="10" pageId="3" pageNumber="42">18.8</td>
<td id="76845DB28208000FFCF0A2FFB72CFC44" box="[892,1024,954,977]" gridcol="2" gridrow="10" pageId="3" pageNumber="42">18.8–22.7</td>
<td id="76845DB28208000FFB00A2FFB63FFC44" box="[1164,1299,954,977]" gridcol="3" gridrow="10" pageId="3" pageNumber="42">20.5±1.55</td>
</tr>
<tr id="355534CE8208000FFF13A2A0B63FFC69" box="[159,1299,997,1020]" gridrow="11" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A2A0B2C2FC69" box="[159,494,997,1020]" gridcol="0" gridrow="11" pageId="3" pageNumber="42">Pectoral-spine length</th>
<td id="76845DB28208000FFD12A2A0B02DFC69" box="[670,769,997,1020]" gridcol="1" gridrow="11" pageId="3" pageNumber="42">14.9</td>
<td id="76845DB28208000FFCF0A2A0B72CFC69" box="[892,1024,997,1020]" gridcol="2" gridrow="11" pageId="3" pageNumber="42">12.5–15.2</td>
<td id="76845DB28208000FFB00A2A0B63FFC69" box="[1164,1299,997,1020]" gridcol="3" gridrow="11" pageId="3" pageNumber="42">13.6±1.01</td>
</tr>
<tr id="355534CE8208000FFF13A555B63FFBB2" box="[159,1299,1040,1063]" gridrow="12" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A555B2C2FBB2" box="[159,494,1040,1063]" gridcol="0" gridrow="12" pageId="3" pageNumber="42">Caudal-fin length</th>
<td id="76845DB28208000FFD12A555B02DFBB2" box="[670,769,1040,1063]" gridcol="1" gridrow="12" pageId="3" pageNumber="42">27.2</td>
<td id="76845DB28208000FFCF0A555B72CFBB2" box="[892,1024,1040,1063]" gridcol="2" gridrow="12" pageId="3" pageNumber="42">25.9–35.2</td>
<td id="76845DB28208000FFB00A555B63FFBB2" box="[1164,1299,1040,1063]" gridcol="3" gridrow="12" pageId="3" pageNumber="42">29.1±3.36</td>
</tr>
<tr id="355534CE8208000FFF13A57FB63FFBC4" box="[159,1299,1082,1105]" gridrow="13" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A57FB2C2FBC4" box="[159,494,1082,1105]" gridcol="0" gridrow="13" pageId="3" pageNumber="42">Caudal peduncle length</th>
<td id="76845DB28208000FFD12A57FB02DFBC4" box="[670,769,1082,1105]" gridcol="1" gridrow="13" pageId="3" pageNumber="42">14.9</td>
<td id="76845DB28208000FFCF0A57FB72CFBC4" box="[892,1024,1082,1105]" gridcol="2" gridrow="13" pageId="3" pageNumber="42">14.0–17.0</td>
<td id="76845DB28208000FFB00A57FB63FFBC4" box="[1164,1299,1082,1105]" gridcol="3" gridrow="13" pageId="3" pageNumber="42">15.4±1.00</td>
</tr>
<tr id="355534CE8208000FFF13A520B63FFBE9" box="[159,1299,1125,1148]" gridrow="14" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A520B2C2FBE9" box="[159,494,1125,1148]" gridcol="0" gridrow="14" pageId="3" pageNumber="42">Caudal peduncle depth</th>
<td id="76845DB28208000FFD12A520B02DFBE9" box="[670,769,1125,1148]" gridcol="1" gridrow="14" pageId="3" pageNumber="42">10.9</td>
<td id="76845DB28208000FFCF0A520B72CFBE9" box="[892,1024,1125,1148]" gridcol="2" gridrow="14" pageId="3" pageNumber="42">9.3–13.2</td>
<td id="76845DB28208000FFB00A520B63FFBE9" box="[1164,1299,1125,1148]" gridcol="3" gridrow="14" pageId="3" pageNumber="42">10.9±1.08</td>
</tr>
<tr id="355534CE8208000FFF13A5D5B63FFB32" box="[159,1299,1168,1191]" gridrow="15" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A5D5B2C2FB32" box="[159,494,1168,1191]" gridcol="0" gridrow="15" pageId="3" pageNumber="42">Body depth at anus</th>
<td id="76845DB28208000FFD12A5D5B02DFB32" box="[670,769,1168,1191]" gridcol="1" gridrow="15" pageId="3" pageNumber="42">14.5</td>
<td id="76845DB28208000FFCF0A5D5B72CFB32" box="[892,1024,1168,1191]" gridcol="2" gridrow="15" pageId="3" pageNumber="42">13.3–15.6</td>
<td id="76845DB28208000FFB00A5D5B63FFB32" box="[1164,1299,1168,1191]" gridcol="3" gridrow="15" pageId="3" pageNumber="42">14.2±0.73</td>
</tr>
<tr id="355534CE8208000FFF13A5FFB63FFB44" box="[159,1299,1210,1233]" gridrow="16" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A5FFB2C2FB44" box="[159,494,1210,1233]" gridcol="0" gridrow="16" pageId="3" pageNumber="42">Head length</th>
<td id="76845DB28208000FFD12A5FFB02DFB44" box="[670,769,1210,1233]" gridcol="1" gridrow="16" pageId="3" pageNumber="42">23.9</td>
<td id="76845DB28208000FFCF0A5FFB72CFB44" box="[892,1024,1210,1233]" gridcol="2" gridrow="16" pageId="3" pageNumber="42">21.6–25.2</td>
<td id="76845DB28208000FFB00A5FFB63FFB44" box="[1164,1299,1210,1233]" gridcol="3" gridrow="16" pageId="3" pageNumber="42">23.1±1.14</td>
</tr>
<tr id="355534CE8208000FFF13A5A0B63FFB69" box="[159,1299,1253,1276]" gridrow="17" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A5A0B2C2FB69" box="[159,494,1253,1276]" gridcol="0" gridrow="17" pageId="3" pageNumber="42">Head width</th>
<td id="76845DB28208000FFD12A5A0B02DFB69" box="[670,769,1253,1276]" gridcol="1" gridrow="17" pageId="3" pageNumber="42">21.4</td>
<td id="76845DB28208000FFCF0A5A0B72CFB69" box="[892,1024,1253,1276]" gridcol="2" gridrow="17" pageId="3" pageNumber="42">21.4–24.4</td>
<td id="76845DB28208000FFB00A5A0B63FFB69" box="[1164,1299,1253,1276]" gridcol="3" gridrow="17" pageId="3" pageNumber="42">23.2±1.05</td>
</tr>
<tr id="355534CE8208000FFF13A455B63FFAB2" box="[159,1299,1296,1319]" gridrow="18" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A455B2C2FAB2" box="[159,494,1296,1319]" gridcol="0" gridrow="18" pageId="3" pageNumber="42">Head depth</th>
<td id="76845DB28208000FFD12A455B02DFAB2" box="[670,769,1296,1319]" gridcol="1" gridrow="18" pageId="3" pageNumber="42">14.5</td>
<td id="76845DB28208000FFCF0A455B72CFAB2" box="[892,1024,1296,1319]" gridcol="2" gridrow="18" pageId="3" pageNumber="42">11.9–15.4</td>
<td id="76845DB28208000FFB00A455B63FFAB2" box="[1164,1299,1296,1319]" gridcol="3" gridrow="18" pageId="3" pageNumber="42">14.1±1.03</td>
</tr>
<tr id="355534CE8208000FFF13A47FB63FFAC4" box="[159,1299,1338,1361]" gridrow="19" pageId="3" pageNumber="42" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76845DB28208000FFF13A47FB2C2FAC4" box="[159,494,1338,1361]" gridcol="0" gridrow="19" pageId="3" pageNumber="42">%HL</th>
</tr>
<tr id="355534CE8208000FFF13A420B63FFAE9" box="[159,1299,1381,1404]" gridrow="20" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A420B2C2FAE9" box="[159,494,1381,1404]" gridcol="0" gridrow="20" pageId="3" pageNumber="42">Snout length</th>
<td id="76845DB28208000FFD12A420B02DFAE9" box="[670,769,1381,1404]" gridcol="1" gridrow="20" pageId="3" pageNumber="42">34.8</td>
<td id="76845DB28208000FFCF0A420B72CFAE9" box="[892,1024,1381,1404]" gridcol="2" gridrow="20" pageId="3" pageNumber="42">29.6–36.8</td>
<td id="76845DB28208000FFB00A420B63FFAE9" box="[1164,1299,1381,1404]" gridcol="3" gridrow="20" pageId="3" pageNumber="42">33.9±2.80</td>
</tr>
<tr id="355534CE8208000FFF13A4D5B63FFA32" box="[159,1299,1424,1447]" gridrow="21" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A4D5B2C2FA32" box="[159,494,1424,1447]" gridcol="0" gridrow="21" pageId="3" pageNumber="42">Interorbital distance</th>
<td id="76845DB28208000FFD12A4D5B02DFA32" box="[670,769,1424,1447]" gridcol="1" gridrow="21" pageId="3" pageNumber="42">40.9</td>
<td id="76845DB28208000FFCF0A4D5B72CFA32" box="[892,1024,1424,1447]" gridcol="2" gridrow="21" pageId="3" pageNumber="42">36.6–41.7</td>
<td id="76845DB28208000FFB00A4D5B63FFA32" box="[1164,1299,1424,1447]" gridcol="3" gridrow="21" pageId="3" pageNumber="42">40.0±1.77</td>
</tr>
<tr id="355534CE8208000FFF13A4FFB63FFA44" box="[159,1299,1466,1489]" gridrow="22" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A4FFB2C2FA44" box="[159,494,1466,1489]" gridcol="0" gridrow="22" pageId="3" pageNumber="42">Eye diameter</th>
<td id="76845DB28208000FFD12A4FFB02DFA44" box="[670,769,1466,1489]" gridcol="1" gridrow="22" pageId="3" pageNumber="42">9.1</td>
<td id="76845DB28208000FFCF0A4FFB72CFA44" box="[892,1024,1466,1489]" gridcol="2" gridrow="22" pageId="3" pageNumber="42">8.5–10.5</td>
<td id="76845DB28208000FFB00A4FFB63FFA44" box="[1164,1299,1466,1489]" gridcol="3" gridrow="22" pageId="3" pageNumber="42">9.5±0.61</td>
</tr>
<tr id="355534CE8208000FFF13A4A0B63FFA69" box="[159,1299,1509,1532]" gridrow="23" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A4A0B2C2FA69" box="[159,494,1509,1532]" gridcol="0" gridrow="23" pageId="3" pageNumber="42">Nasal barbel length</th>
<td id="76845DB28208000FFD12A4A0B02DFA69" box="[670,769,1509,1532]" gridcol="1" gridrow="23" pageId="3" pageNumber="42">65.2</td>
<td id="76845DB28208000FFCF0A4A0B72CFA69" box="[892,1024,1509,1532]" gridcol="2" gridrow="23" pageId="3" pageNumber="42">65.2–112.7</td>
<td id="76845DB28208000FFB00A4A0B63FFA69" box="[1164,1299,1509,1532]" gridcol="3" gridrow="23" pageId="3" pageNumber="42">92.8±14.82</td>
</tr>
<tr id="355534CE8208000FFF13A755B63FF9B2" box="[159,1299,1552,1575]" gridrow="24" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A755B2C2F9B2" box="[159,494,1552,1575]" gridcol="0" gridrow="24" pageId="3" pageNumber="42">Maxillary barbel length</th>
<td id="76845DB28208000FFD12A755B02DF9B2" box="[670,769,1552,1575]" gridcol="1" gridrow="24" pageId="3" pageNumber="42">116.7</td>
<td id="76845DB28208000FFCF0A755B72CF9B2" box="[892,1024,1552,1575]" gridcol="2" gridrow="24" pageId="3" pageNumber="42">115.8–144.4</td>
<td id="76845DB28208000FFB00A755B63FF9B2" box="[1164,1299,1552,1575]" gridcol="3" gridrow="24" pageId="3" pageNumber="42">130.2±11.54</td>
</tr>
<tr id="355534CE8208000FFF13A77FB63FF9C4" box="[159,1299,1594,1617]" gridrow="25" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A77FB2C2F9C4" box="[159,494,1594,1617]" gridcol="0" gridrow="25" pageId="3" pageNumber="42">Inner mandibular barbel length</th>
<td id="76845DB28208000FFD12A77FB02DF9C4" box="[670,769,1594,1617]" gridcol="1" gridrow="25" pageId="3" pageNumber="42">28.8</td>
<td id="76845DB28208000FFCF0A77FB72CF9C4" box="[892,1024,1594,1617]" gridcol="2" gridrow="25" pageId="3" pageNumber="42">28.8–66.7</td>
<td id="76845DB28208000FFB00A77FB63FF9C4" box="[1164,1299,1594,1617]" gridcol="3" gridrow="25" pageId="3" pageNumber="42">52.4±12.29</td>
</tr>
<tr id="355534CE8208000FFF13A720B63FF9E9" box="[159,1299,1637,1660]" gridrow="26" pageId="3" pageNumber="42">
<th id="76845DB28208000FFF13A720B2C2F9E9" box="[159,494,1637,1660]" gridcol="0" gridrow="26" pageId="3" pageNumber="42">Outer mandibular barbel length</th>
<td id="76845DB28208000FFD12A720B02DF9E9" box="[670,769,1637,1660]" gridcol="1" gridrow="26" pageId="3" pageNumber="42">86.4</td>
<td id="76845DB28208000FFCF0A720B72CF9E9" box="[892,1024,1637,1660]" gridcol="2" gridrow="26" pageId="3" pageNumber="42">86.4–124.2</td>
<td id="76845DB28208000FFB00A720B63FF9E9" box="[1164,1299,1637,1660]" gridcol="3" gridrow="26" pageId="3" pageNumber="42">102.5±12.69</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDA368C8208FFF3FF49A788B7F9F83A" blockId="3.[151,1437,1741,1967]" pageId="3" pageNumber="42">
<emphasis id="B911EA9E8208FFF3FF49A788B23EF972" bold="true" box="[197,274,1741,1767]" pageId="3" pageNumber="42">Color.</emphasis>
In 70% ethanol: dorsal and lateral surfaces of head and body dark brown. Belly, chest, and ventral surface of head cream. A series of about five to six light brown spots on dorsolateral surface of body, arranged longitudinally from below dorsal fin base to caudal peduncle, frequently partially coalescent with elongate light brown patch on dorsal surface of body running along entire postdorsal length. Proximal three-quarters of dorsal fin dark brown, with distal one-quarter hyaline. Pectoral, pelvic, anal, and caudal fins hyaline with scattered brown spots, denser on rays than on membranes. Barbels cream with dark brown rings.
</paragraph>
</subSubSection>
<caption id="DF1A6604820FFFF4FF1BA286B675FBB5" httpUri="https://zenodo.org/record/185204/files/figure.png" pageId="4" pageNumber="43" targetBox="[151,1435,194,939]" targetPageId="4">
<paragraph id="8BDA368C820FFFF4FF1BA286B675FBB5" blockId="4.[151,1435,963,1056]" pageId="4" pageNumber="43">
<emphasis id="B911EA9E820FFFF4FF1BA286B237FC4E" bold="true" box="[151,283,963,987]" pageId="4" pageNumber="43">FIGURE 2.</emphasis>
Posterior portion of bodies in: a.
<taxonomicName id="4C654D0F820FFFF4FD02A286B059FC4F" box="[654,885,963,986]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="hystriculus">
<emphasis id="B911EA9E820FFFF4FD02A286B059FC4F" box="[654,885,963,986]" italics="true" pageId="4" pageNumber="43">Parakysis hystriculus</emphasis>
</taxonomicName>
, ZRC 51418, male paratype, 26.0 mm SL (above) and ZRC 51418, female paratype, 25.7 mm SL (below); b.
<taxonomicName id="4C654D0F820FFFF4FC9FA2A3B0ACFC68" box="[787,896,998,1021]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B911EA9E820FFFF4FC9FA2A3B0ACFC68" box="[787,896,998,1021]" italics="true" pageId="4" pageNumber="43">P. grandis</emphasis>
</taxonomicName>
, ZRC 38632, male paratype, 24.2 mm SL (above) and ZRC 39178, female paratype, 29.3 mm SL (below), showing difference in anal-fin shape. Images not to scale.
</paragraph>
</caption>
<subSubSection id="C37F6507820FFFF4FF49A516B14BFB2E" pageId="4" pageNumber="43" type="distribution">
<paragraph id="8BDA368C820FFFF4FF49A516B14BFB2E" blockId="4.[151,1437,1107,1211]" pageId="4" pageNumber="43">
<emphasis id="B911EA9E820FFFF4FF49A516B24AFBF8" bold="true" box="[197,358,1107,1133]" pageId="4" pageNumber="43">Distribution.</emphasis>
Known from the Lalang River drainage in southern Sumatra (
<figureCitation id="135E2A09820FFFF4FBB3A516B7AAFBF8" box="[1087,1158,1107,1133]" captionStart="FIGURE 3" captionStartId="5.[151,255,1813,1837]" captionTargetBox="[151,1433,200,1776]" captionTargetId="figure@5.[151,1436,194,1790]" captionTargetPageId="5" captionText="FIGURE 3. Map showing distribution of Parakysis species in Sumatra." httpUri="https://zenodo.org/record/185205/files/figure.png" pageId="4" pageNumber="43">Fig. 3</figureCitation>
).
<emphasis id="B911EA9E820FFFF4FF1BA53EB208FB00" bold="true" box="[151,292,1147,1173]" pageId="4" pageNumber="43">Etymology.</emphasis>
From the Latin adjective
<taxonomicName id="4C654D0F820FFFF4FDD1A53EB1CCFB00" box="[605,736,1147,1173]" class="Actinopterygii" family="Akysidae" genus="Parakysis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="hystriculus">hystriculus</taxonomicName>
, meaning hairy or bearded; in reference to the long accessory mandibular barbels of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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