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<document id="6065ADD0DE2525F0AB9BDECDC9D9608B" ID-DOI="10.5281/zenodo.12521148" ID-ISSN="1407-8619" ID-Zenodo-Dep="12521148" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719252154146" checkinUser="felipe" docAuthor="Rukmane, Anita" docDate="2019" docId="03CC878DFFD7DB4BDE77503EFD7E14BD" docLanguage="en" docName="BalticJColeopterol.19.2.141-150.pdf" docOrigin="Baltic Journal of Coleopterology 19 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.3:BalticJColeop.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeop.2012-2021.journal_article" docStyleVersion="3" docTitle="Pachyrhynchus phaleratus subsp. dannylayroni Rukmane, 2019, subsp. nov." docType="treatment" docVersion="2" lastPageNumber="149" masterDocId="FFF5FFF5FFD1DB43DD245341FFCD152A" masterDocTitle="Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines" masterLastPageNumber="150" masterPageNumber="141" pageNumber="147" updateTime="1719254198936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="0F58DF2A240127A46D67C0592AEA5B4E">Four new species and two subspecies of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae: Pachyrhynchini) from Luzon Island, Philippines</mods:title>
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<mods:namePart id="B55026D020193C1BE6C09CE77D24F137">Rukmane, Anita</mods:namePart>
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<mods:title id="01135C4BC35FBC1D4C9047E43A6D0A62">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="1C63AF1F3A615E54B38DBF648AE8C92F">
<mods:date id="146F4A2DBB361F674FAA424EB0AF9C05">2019</mods:date>
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<mods:number id="E7EF3D878C190C82EF254C41D8D74BF6">2019-12-31</mods:number>
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<mods:number id="7F81157C4C794583351A76C51BC98113">19</mods:number>
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<mods:identifier id="0ED8F09792A4B80C62C7F2F47A88BBCD" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="B4A5BFB1AD7005EF5A9DEEEAA96C69A2" type="Zenodo-Dep">12521148</mods:identifier>
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<treatment id="03CC878DFFD7DB4BDE77503EFD7E14BD" LSID="urn:lsid:plazi:treatment:03CC878DFFD7DB4BDE77503EFD7E14BD" httpUri="http://treatment.plazi.org/id/03CC878DFFD7DB4BDE77503EFD7E14BD" lastPageId="8" lastPageNumber="149" pageId="6" pageNumber="147">
<subSubSection id="C37F6510FFD7DB45DE77503EFC0B169D" pageId="6" pageNumber="147" type="nomenclature">
<paragraph id="8BDA369BFFD7DB45DE77503EFC0B169D" blockId="6.[851,1361,895,983]" pageId="6" pageNumber="147">
<emphasis id="B911EA89FFD7DB45DE77503EFC0B169D" bold="true" pageId="6" pageNumber="147">
<heading id="D09281F7FFD7DB45DE77503EFA9C16BD" bold="true" box="[851,1361,895,919]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="147" reason="4">
<taxonomicName id="4C654D18FFD7DB45DE77503EFA9C16BD" authorityName="Rukmane" authorityYear="2019" box="[851,1361,895,919]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="subSpecies" species="phaleratus" status="subsp. nov." subSpecies="dannylayroni">
<emphasis id="B911EA89FFD7DB45DE77503EFB4D16BD" bold="true" box="[851,1152,895,919]" italics="true" pageId="6" pageNumber="147">Pachyrhynchus phaleratus</emphasis>
ssp.
<emphasis id="B911EA89FFD7DB45D998503EFA9C16BD" bold="true" box="[1212,1361,895,919]" italics="true" pageId="6" pageNumber="147">dannylayroni</emphasis>
</taxonomicName>
</heading>
<taxonomicNameLabel id="A22257F2FFD7DB45DE7750DEFC0B169D" box="[851,966,927,951]" pageId="6" pageNumber="147" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD7DB45DE7750FEFC1216FD" box="[851,991,959,983]" pageId="6" pageNumber="147" type="description">
<paragraph id="8BDA369BFFD7DB45DE7750FEFC1216FD" blockId="6.[851,1361,895,983]" box="[851,991,959,983]" pageId="6" pageNumber="147">
<figureCitation id="135E2A1EFFD7DB45DE7750FEFC4116FD" box="[851,908,959,983]" captionStart="Fig" captionStartId="5.[774,816,1682,1706]" captionTargetBox="[848,1217,962,1617]" captionTargetId="figure-212@5.[842,1225,957,1621]" captionTargetPageId="5" captionText="Fig. 3. Dorsal habitus of P.phaleratus ssp. dannylayroni subsp. nov. female" figureDoi="http://doi.org/10.5281/zenodo.12521154" httpUri="https://zenodo.org/record/12521154/files/figure.png" pageId="6" pageNumber="147">
Fig.
<geoCoordinate id="EE51505CFFD7DB45DE5A50FEFC4116FD" box="[894,908,959,983]" degrees="3, 4" direction="east" orientation="longitude" pageId="6" pageNumber="147" precision="5555" value="3.4">3</geoCoordinate>
</figureCitation>
,
<figureCitation id="135E2A1EFFD7DB45DEB150FEFC1116FD" box="[917,988,959,983]" captionStart="Fig" captionStartId="7.[227,267,1283,1307]" captionTargetBox="[239,1246,229,1199]" captionTargetId="figure-4@7.[234,1313,209,1209]" captionTargetPageId="7" captionText="Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B – P.disargus sp. nov.; C, D – P.congestus ssp. aedamlayroni subsp. nov.; H, I – P.kirklayroni sp. nov.; J, K – P.sagittatus sp. nov.; L, M – P.tetramaculatus sp.nov.; E, F, G – female genitalia of P.phaleratus ssp. dannylayroni subsp. nov.; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="6" pageNumber="147">4E,F,G</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD7DB45DE7750BEFACC119D" pageId="6" pageNumber="147" type="materials_examined">
<paragraph id="8BDA369BFFD7DB45DE7750BEFACC119D" blockId="6.[851,1361,1023,1208]" pageId="6" pageNumber="147">
<emphasis id="B911EA89FFD7DB45DE7750BEFC20113D" bold="true" box="[851,1005,1023,1047]" pageId="6" pageNumber="147">Type material.</emphasis>
<materialsCitation id="3B0D3CC6FFD7DB45DED750BEFB30119D" collectingDate="2019-01" collectorName="Rukmane" country="Philippines" determinerName="Rukmane A." location="Dingalan" municipality="Luzon" pageId="6" pageNumber="147" specimenCount="2" stateProvince="Aurora" typeStatus="holotype">
Female.
<typeStatus id="54DE8839FFD7DB45D96350BEFB69113D" box="[1095,1188,1023,1047]" pageId="6" pageNumber="147" type="holotype">Holotype</typeStatus>
. ‘’
<collectingCountry id="F372760BFFD7DB45D99050BEFA9C113D" box="[1204,1361,1023,1047]" name="Philippines" pageId="6" pageNumber="147">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6BBEACE1FFD7DB45DE7B5761FC68111D" box="[863,933,1056,1079]" pageId="6" pageNumber="147">Luzon</collectingMunicipality>
,
<location id="8EBA6040FFD7DB45DE8A575EFBDE111D" LSID="urn:lsid:plazi:treatment:03CC878DFFD7DB4BDE77503EFD7E14BD:8EBA6040FFD7DB45DE8A575EFBDE111D" box="[942,1043,1055,1079]" country="Philippines" municipality="Luzon" name="Dingalan" pageId="6" pageNumber="147" stateProvince="Aurora">Dingalan</location>
,
<collectingRegion id="49A1F879FFD7DB45D9385761FBAA1112" box="[1052,1127,1056,1080]" country="Philippines" name="Aurora" pageId="6" pageNumber="147">Aurora</collectingRegion>
/
<date id="FFDB105BFFD7DB45D95C5761FB0C111D" box="[1144,1217,1055,1079]" pageId="6" pageNumber="147" value="2019-01">
<collectingDate id="EF9FE9B3FFD7DB45D95C5761FB0C111D" box="[1144,1217,1055,1079]" pageId="6" pageNumber="147" value="2019-01">I. 2019</collectingDate>
</date>
/ local collector leg.’’ (white rectangular label, printed); ‘’
<typeStatus id="54DE8839FFD7DB45DE7F571EFC3F115D" box="[859,1010,1119,1143]" pageId="6" pageNumber="147" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C654D18FFD7DB45D920571EFC2811BD" authorityName="Rukmane" authorityYear="2019" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="subSpecies" species="phaleratus" subSpecies="dannylayroni">
<emphasis id="B911EA89FFD7DB45D920571EFAEF115D" box="[1028,1314,1119,1143]" italics="true" pageId="6" pageNumber="147">Pachyrhynchus phaleratus</emphasis>
ssp.
<emphasis id="B911EA89FFD7DB45DE77573EFC2811BD" box="[851,997,1151,1175]" italics="true" pageId="6" pageNumber="147">dannylayroni</emphasis>
</taxonomicName>
/
<collectorName id="2690534DFFD7DB45DED9573EFBA511BD" box="[1021,1128,1151,1175]" pageId="6" pageNumber="147">Rukmane</collectorName>
, 2019 / det.
<determinerName id="7E06DA5FFFD7DB45D9CD573EFCA01192" pageId="6" pageNumber="147">Rukmane A.</determinerName>
2019’’ (red rectangular label, printed)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD7DB45DE77579EFAFA11DD" box="[851,1335,1247,1272]" pageId="6" pageNumber="147" type="distribution">
<paragraph id="8BDA369BFFD7DB45DE77579EFAFA11DD" blockId="6.[851,1335,1247,1272]" box="[851,1335,1247,1272]" pageId="6" pageNumber="147">
<emphasis id="B911EA89FFD7DB45DE77579EFC2B11DD" bold="true" box="[851,998,1247,1271]" pageId="6" pageNumber="147">Distribution:</emphasis>
Luzon Island,
<collectingRegion id="49A1F879FFD7DB45D9A557A1FAFE11DD" box="[1153,1331,1247,1272]" country="Philippines" name="Aurora" pageId="6" pageNumber="147">Aurora Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD7DB4BDE77565EFE60143D" lastPageId="8" lastPageNumber="149" pageId="6" pageNumber="147" type="description">
<paragraph id="8BDA369BFFD7DB45DE77565EFA9D139D" blockId="6.[851,1362,1311,1719]" pageId="6" pageNumber="147">
<emphasis id="B911EA89FFD7DB45DE77565EFC11101D" bold="true" box="[851,988,1311,1335]" pageId="6" pageNumber="147">Description.</emphasis>
Female. Measurements (
<typeStatus id="54DE8839FFD7DB45D9C4565EFA8B101D" box="[1248,1350,1311,1335]" pageId="6" pageNumber="147" type="holotype">Holotype</typeStatus>
): LB: 13.1; LR: 1.9; WR: 1.8; LP: 4.0; WP: 3.7; LE: 8.9; WE: 6.2. Rostrum wider than long, LR/WR 1,05; pronotum slightly longer than wide, LP/WP 1.08; elytra nearly 1.5 times longer than wide, LE/ WE 1.44; elytra more than two times as long as prothorax, LE/LP 2.23; elytra wider than prothorax, WE/WP 1.66. Body surface black, shiny, with pale green markings of round to recumbent scales. Dorsal habitus as shown in
<figureCitation id="135E2A1EFFD7DB45D837557EFA80137D" box="[1299,1357,1599,1623]" captionStart="Fig" captionStartId="5.[774,816,1682,1706]" captionTargetBox="[848,1217,962,1617]" captionTargetId="figure-212@5.[842,1225,957,1621]" captionTargetPageId="5" captionText="Fig. 3. Dorsal habitus of P.phaleratus ssp. dannylayroni subsp. nov. female" figureDoi="http://doi.org/10.5281/zenodo.12521154" httpUri="https://zenodo.org/record/12521154/files/figure.png" pageId="6" pageNumber="147">Fig.3</figureCitation>
. Prothorax with same markings as in
<taxonomicName id="4C654D18FFD7DB45D9E9551EFC3813BD" authorityName="G.R. Waterhouse" authorityYear="1841" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="147" phylum="Arthropoda" rank="subSpecies" species="phaleratus" subSpecies="phaleratus">
<emphasis id="B911EA89FFD7DB45D9E9551EFA9C135D" box="[1229,1361,1631,1655]" italics="true" pageId="6" pageNumber="147">P.phaleratus</emphasis>
ssp.
<emphasis id="B911EA89FFD7DB45DEA6553EFC3813BD" box="[898,1013,1663,1687]" italics="true" pageId="6" pageNumber="147">phaleratus</emphasis>
</taxonomicName>
. Each elytron with the following markings: 1) three longitudinal stripes at basal
</paragraph>
<paragraph id="8BDA369BFFD6DB44DF9E53D0FCE3158E" blockId="7.[698,814,145,164]" box="[698,814,145,164]" pageId="7" pageNumber="148">
<emphasis id="B911EA89FFD6DB44DF9E53D0FCE3158E" box="[698,814,145,164]" italics="true" pageId="7" pageNumber="148">Rukmane A.</emphasis>
</paragraph>
<caption id="DF1A6613FFD6DB44DDC75642FDFF10B1" ID-DOI="http://doi.org/10.5281/zenodo.12521156" ID-Zenodo-Dep="12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="7" pageNumber="148" startId="7.[227,267,1283,1307]" targetBox="[239,1246,229,1199]" targetPageId="7" targetType="figure">
<paragraph id="8BDA369BFFD6DB44DDC75642FDFF10B1" blockId="7.[227,1288,1283,1435]" pageId="7" pageNumber="148">
Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B –
<taxonomicName id="4C654D18FFD6DB44DDC75662FE9E1011" authorityName="Rukmane" authorityYear="2019" box="[227,339,1315,1339]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="disargus" status="sp. nov.">
<emphasis id="B911EA89FFD6DB44DDC75662FE9E1011" box="[227,339,1315,1339]" italics="true" pageId="7" pageNumber="148">P.disargus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44DC7C5665FE621011" box="[344,431,1316,1339]" pageId="7" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
; C, D –
<taxonomicName id="4C654D18FFD6DB44DF205664FC831011" authorityName="Rukmane" authorityYear="2019" box="[516,846,1315,1340]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="subSpecies" species="congestus" status="subsp. nov." subSpecies="aedamlayroni">
<emphasis id="B911EA89FFD6DB44DF205664FD491016" box="[516,644,1317,1340]" italics="true" pageId="7" pageNumber="148">P.congestus</emphasis>
ssp.
<emphasis id="B911EA89FFD6DB44DF9C5662FC831011" box="[696,846,1315,1339]" italics="true" pageId="7" pageNumber="148">aedamlayroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44DE705662FC021011" box="[852,975,1315,1339]" pageId="7" pageNumber="148" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
; H, I –
<taxonomicName id="4C654D18FFD6DB44D93F5662FB651011" authorityName="Rukmane" authorityYear="2019" box="[1051,1192,1315,1339]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="kirklayroni" status="sp. nov.">
<emphasis id="B911EA89FFD6DB44D93F5662FB651011" box="[1051,1192,1315,1339]" italics="true" pageId="7" pageNumber="148">P.kirklayroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44D9895665FACE1011" box="[1197,1283,1316,1339]" pageId="7" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
; J, K –
<taxonomicName id="4C654D18FFD6DB44DC075604FE531076" authorityName="Rukmane" authorityYear="2019" box="[291,414,1349,1372]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="sagittatus" status="sp. nov.">
<emphasis id="B911EA89FFD6DB44DC075604FE531076" box="[291,414,1349,1372]" italics="true" pageId="7" pageNumber="148">P.sagittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44DC875605FE3A1071" box="[419,503,1348,1371]" pageId="7" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
; L, M –
<taxonomicName id="4C654D18FFD6DB44DF6F5602FD331071" authorityName="Rukmane" authorityYear="2019" box="[587,766,1347,1371]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp.nov.">
<emphasis id="B911EA89FFD6DB44DF6F5602FD331071" box="[587,766,1347,1371]" italics="true" pageId="7" pageNumber="148">P.tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44DE205605FC9F1071" box="[772,850,1348,1371]" pageId="7" pageNumber="148" rank="species">sp.nov.</taxonomicNameLabel>
; E, F, G – female genitalia of
<taxonomicName id="4C654D18FFD6DB44D9A65602FE691051" authorityName="Rukmane" authorityYear="2019" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="subSpecies" species="phaleratus" status="subsp. nov." subSpecies="dannylayroni">
<emphasis id="B911EA89FFD6DB44D9A65602FAC51071" box="[1154,1288,1347,1371]" italics="true" pageId="7" pageNumber="148">P.phaleratus</emphasis>
ssp.
<emphasis id="B911EA89FFD6DB44DC375622FE691051" box="[275,420,1379,1403]" italics="true" pageId="7" pageNumber="148">dannylayroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F2FFD6DB44DC8E5622FDEB1051" box="[426,550,1379,1403]" pageId="7" pageNumber="148" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm
</paragraph>
</caption>
<paragraph id="8BDA369BFFD6DB44DDC75688FC3A1363" blockId="7.[227,737,1480,1728]" lastBlockId="7.[774,1285,1489,1641]" pageId="7" pageNumber="148">
part, from base to submedial part; 2) longitudinal line along lateral margin in all length; 3) two elongated sutural patches at apical part; 4) two roundish patches medially, two roundish patches at apical ½; 5) longitudinal stripe from the middle to apical 1/2 along interval IV; 6) triangular patch near apex laterally; elytra widest just in the middle, apex firmly rounded. Metasternum covered with round pale green scales; two patches on ventrite I, each redirected laterally; round patch of scales on each side of ventrite II. Genitalia as illustrated in
<figureCitation id="135E2A1EFFD6DB44DEA95570FC0B1363" box="[909,966,1585,1609]" captionStart="Fig" captionStartId="7.[227,267,1283,1307]" captionTargetBox="[239,1246,229,1199]" captionTargetId="figure-4@7.[234,1313,209,1209]" captionTargetPageId="7" captionText="Fig. 4. A, C, H, J, L – aedeagus in lateral view; B, D, I, K, M – aedeagus in ventral view; A, B – P.disargus sp. nov.; C, D – P.congestus ssp. aedamlayroni subsp. nov.; H, I – P.kirklayroni sp. nov.; J, K – P.sagittatus sp. nov.; L, M – P.tetramaculatus sp.nov.; E, F, G – female genitalia of P.phaleratus ssp. dannylayroni subsp. nov.; E – sternite VIII in ventral view; F – ovipositor in dorsal view; G – spermatheca. Scale bar 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.12521156" httpUri="https://zenodo.org/record/12521156/files/figure.png" pageId="7" pageNumber="148">Fig. 4</figureCitation>
E-G.
</paragraph>
<paragraph id="8BDA369BFFD6DB44DE225510FC621343" blockId="7.[774,1285,1489,1641]" box="[774,943,1617,1641]" pageId="7" pageNumber="148">Male unknown.</paragraph>
<paragraph id="8BDA369BFFD6DB4BDE2255D3FE60143D" blockId="7.[774,1285,1681,1737]" lastBlockId="8.[303,813,191,279]" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148">
<emphasis id="B911EA89FFD6DB44DE2255D3FC321380" bold="true" box="[774,1023,1682,1706]" pageId="7" pageNumber="148">Differential analyses.</emphasis>
This new subspecies is delimitated from
<taxonomicName id="4C654D18FFD6DB44DEEE55F0FAC813E3" authorityName="G.R. Waterhouse" authorityYear="1841" box="[970,1285,1713,1737]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="148" phylum="Arthropoda" rank="subSpecies" species="phaleratus" subSpecies="phaleratus">
<emphasis id="B911EA89FFD6DB44DEEE55F0FB9E13E3" box="[970,1107,1713,1737]" italics="true" pageId="7" pageNumber="148">P.phaleratus</emphasis>
ssp.
<emphasis id="B911EA89FFD6DB44D9B455F0FAC813E3" box="[1168,1285,1713,1737]" italics="true" pageId="7" pageNumber="148">phaleratus</emphasis>
</taxonomicName>
mainly by its unique markings on elytra, which are much more thick than in
<taxonomicName id="4C654D18FFD9DB4BDF56539EFE64143D" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="149" phylum="Arthropoda" rank="subSpecies" species="phaleartus" subSpecies="phaleratus">
<emphasis id="B911EA89FFD9DB4BDF56539EFD3615DD" box="[626,763,223,247]" italics="true" pageId="8" pageNumber="149">P.phaleartus</emphasis>
ssp.
<emphasis id="B911EA89FFD9DB4BDC0B53BEFE64143D" box="[303,425,255,279]" italics="true" pageId="8" pageNumber="149">phaleratus</emphasis>
</taxonomicName>
<emphasis id="B911EA89FFD9DB4BDC8D53BEFE60143D" box="[425,429,255,279]" italics="true" pageId="8" pageNumber="149">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37F6510FFD9DB4BDC0B527EFD7E14BD" pageId="8" pageNumber="149" type="etymology">
<paragraph id="8BDA369BFFD9DB4BDC0B527EFD7E14BD" blockId="8.[303,813,319,407]" pageId="8" pageNumber="149">
<emphasis id="B911EA89FFD9DB4BDC0B527EFE7D147D" bold="true" box="[303,432,319,343]" pageId="8" pageNumber="149">Etymology.</emphasis>
This subspecies was named after Danny Layron in appreciation of cooperation and possibility to study current material.
</paragraph>
</subSubSection>
</treatment>
</document>
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