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<document id="659393D7D9B37C45CC99476E1D38C279" ID-CLB-Dataset="50963" ID-DOI="10.5281/zenodo.169839" ID-GBIF-Dataset="6e63434b-16a3-4599-a191-7f66b91d5d21" ID-ISSN="1175­5326" ID-Zenodo-Dep="169839" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459746906191" checkinUser="plazi" docAuthor="Ziegler, Thomas &amp; Quyet, Le Khac" docDate="2005" docId="03DF87A6C37B977AFE907667FD65FE16" docLanguage="en" docName="zt01042p038.pdf" docOrigin="Zootaxa 1042" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Calamaria thanhi Ziegler &amp; Quyet, 2005, sp. n." docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FFE6FFDEC3799773FF98777BFFEBFF90" masterDocTitle="A new species of reed snake, Calamaria (Squamata: Colubridae), from the Central Truong Son (Annamite mountain range), Vietnam" masterLastPageNumber="38" masterPageNumber="27" pageNumber="29" updateTime="1732488054323" updateUser="ExternalLinkService">
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<mods:title id="D3AE488A318B91C8A9DA925919AD23E6">A new species of reed snake, Calamaria (Squamata: Colubridae), from the Central Truong Son (Annamite mountain range), Vietnam</mods:title>
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<mods:namePart id="C3141DDE75259DEB88D6F8F7CA6B7C6D">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="185A1D623682CF381E212C346E7DA675">Quyet, Le Khac</mods:namePart>
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<mods:typeOfResource id="DA160F7E2A51F10FCD51BCCC1F21E725">text</mods:typeOfResource>
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<mods:date id="34973469ECFA0E283CA2CB36C6F49E59">2005</mods:date>
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<treatment id="03DF87A6C37B977AFE907667FD65FE16" ID-DOI="http://doi.org/10.5281/zenodo.6267236" ID-GBIF-Taxon="119336600" ID-Zenodo-Dep="6267236" LSID="urn:lsid:plazi:treatment:03DF87A6C37B977AFE907667FD65FE16" httpUri="http://treatment.plazi.org/id/03DF87A6C37B977AFE907667FD65FE16" lastPageId="9" lastPageNumber="35" pageId="2" pageNumber="29" scope_class="Squamata" scope_family="Colubridae">
<subSubSection id="C36C653BC37B9771FE907667FE73FEC9" pageId="2" pageNumber="29" type="nomenclature">
<paragraph id="8BC936B0C37B9771FE907667FDC2FEA5" blockId="2.[264,553,283,345]" box="[264,553,283,310]" pageId="2" pageNumber="29">
<heading id="D08181DCC37B9771FE907667FDC2FEA5" bold="true" box="[264,553,283,310]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
<emphasis id="B902EAA2C37B9771FE907667FDC2FEA5" bold="true" box="[264,553,283,310]" pageId="2" pageNumber="29">
<taxonomicName id="4C764D33C37B9771FE907667FE0BFEA6" ID-CoL="PD8Y" box="[264,480,284,310]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37B9771FE907667FE0BFEA6" bold="true" box="[264,480,284,310]" italics="true" pageId="2" pageNumber="29">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37B9771FE7F7660FDC2FEA5" box="[487,553,283,309]" pageId="2" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936B0C37B9771FE907644FE73FEC9" blockId="2.[264,553,283,345]" box="[264,408,319,345]" pageId="2" pageNumber="29">
<figureCitation id="134D2A35C37B9771FE907644FE7FFEC9" box="[264,404,319,345]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[264,367,1010,1034]" captionStartId-1="4.[264,368,1819,1843]" captionStartId-2="5.[264,368,1044,1068]" captionStartId-3="5.[264,367,1765,1789]" captionStartId-4="6.[264,368,1039,1063]" captionStartId-5="6.[264,368,1811,1835]" captionTargetBox-0="[264,1322,323,1002]" captionTargetBox-1="[264,1322,1140,1801]" captionTargetBox-2="[264,1323,326,1020]" captionTargetBox-3="[264,1322,1140,1758]" captionTargetBox-4="[264,1323,326,1014]" captionTargetBox-5="[264,1322,1135,1787]" captionTargetId-0="figure@4.[264,1323,323,1003]" captionTargetId-1="figure@4.[264,1323,1139,1802]" captionTargetId-2="figure@5.[264,1323,326,1020]" captionTargetId-3="figure@5.[264,1323,1136,1758]" captionTargetId-4="figure@6.[264,1323,326,1015]" captionTargetId-5="figure@6.[264,1323,1135,1787]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionText-0="FIGURE 1. Dorsal view of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-1="FIGURE 2. Ventral view of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-2="FIGURE 3. Oblique view of the head and neck region of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-3="FIGURE 4. Lateral view of the head of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-4="FIGURE 5. Ventral view of the head of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-5="FIGURE 6. Dorsal view of the tail of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920). Photos by Thomas Ziegler." httpUri-0="https://zenodo.org/record/169840/files/figure.png" httpUri-1="https://zenodo.org/record/169841/files/figure.png" httpUri-2="https://zenodo.org/record/169842/files/figure.png" httpUri-3="https://zenodo.org/record/169843/files/figure.png" httpUri-4="https://zenodo.org/record/169844/files/figure.png" httpUri-5="https://zenodo.org/record/169845/files/figure.png" pageId="2" pageNumber="29">Figures 1–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37B9771FE9076F5FDA9FDB1" pageId="2" pageNumber="29" type="materials_examined">
<paragraph id="8BC936B0C37B9771FE9076F5FDA9FDB1" blockId="2.[264,1323,398,1865]" pageId="2" pageNumber="29">
<emphasis id="B902EAA2C37B9771FE9076F5FE75FE38" bold="true" box="[264,414,398,424]" pageId="2" pageNumber="29">
<typeStatus id="54CD8812C37B9771FE9076F5FE90FE38" box="[264,379,398,424]" pageId="2" pageNumber="29" type="holotype">Holotype</typeStatus>
.—
</emphasis>
An adult female (
<collectionCode id="ED67AE75C37B9771FDF676F4FD29FE39" LSID="urn:lsid:biocol.org:col:34613" box="[622,706,399,425]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="29">ZFMK</collectionCode>
82920), adjacent to Phong Nha
<emphasis id="B902EAA2C37B9771FBDB76F5FBA6FE38" bold="true" box="[1091,1101,398,424]" pageId="2" pageNumber="29">­</emphasis>
Ke Bang National Park, Dan Hoa Commune, Minh Hoa District, Quang Binh Province,
<collectingCountry id="F3617620C37B9771FBD576CCFB58FE41" box="[1101,1203,439,465]" name="Vietnam" pageId="2" pageNumber="29">Vietnam</collectingCountry>
, collected on
<date id="FFC81070C37B9771FEB476A4FE22FE69" box="[300,457,479,505]" pageId="2" pageNumber="29" value="2003-06-16">16 June 2003</date>
by local people in a limestone cave of primary forest; for injuries prior to collection see
<figureCitation id="134D2A35C37B9771FE37757CFDD5FDB1" box="[431,574,519,545]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[264,367,1010,1034]" captionStartId-1="4.[264,368,1819,1843]" captionStartId-2="5.[264,368,1044,1068]" captionStartId-3="5.[264,367,1765,1789]" captionStartId-4="6.[264,368,1039,1063]" captionStartId-5="6.[264,368,1811,1835]" captionTargetBox-0="[264,1322,323,1002]" captionTargetBox-1="[264,1322,1140,1801]" captionTargetBox-2="[264,1323,326,1020]" captionTargetBox-3="[264,1322,1140,1758]" captionTargetBox-4="[264,1323,326,1014]" captionTargetBox-5="[264,1322,1135,1787]" captionTargetId-0="figure@4.[264,1323,323,1003]" captionTargetId-1="figure@4.[264,1323,1139,1802]" captionTargetId-2="figure@5.[264,1323,326,1020]" captionTargetId-3="figure@5.[264,1323,1136,1758]" captionTargetId-4="figure@6.[264,1323,326,1015]" captionTargetId-5="figure@6.[264,1323,1135,1787]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionText-0="FIGURE 1. Dorsal view of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-1="FIGURE 2. Ventral view of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-2="FIGURE 3. Oblique view of the head and neck region of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-3="FIGURE 4. Lateral view of the head of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-4="FIGURE 5. Ventral view of the head of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920)." captionText-5="FIGURE 6. Dorsal view of the tail of the ethanol­preserved holotype of Calamaria thanhi sp. n. (ZFMK 82920). Photos by Thomas Ziegler." httpUri-0="https://zenodo.org/record/169840/files/figure.png" httpUri-1="https://zenodo.org/record/169841/files/figure.png" httpUri-2="https://zenodo.org/record/169842/files/figure.png" httpUri-3="https://zenodo.org/record/169843/files/figure.png" httpUri-4="https://zenodo.org/record/169844/files/figure.png" httpUri-5="https://zenodo.org/record/169845/files/figure.png" pageId="2" pageNumber="29">Figures 1–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37B9771FEA07555FB83FB61" pageId="2" pageNumber="29" type="diagnosis">
<paragraph id="8BC936B0C37B9771FEA07555FB83FB61" blockId="2.[264,1323,398,1865]" pageId="2" pageNumber="29">
<emphasis id="B902EAA2C37B9771FEA07555FE3EFDD8" bold="true" box="[312,469,558,584]" pageId="2" pageNumber="29">Diagnosis.—</emphasis>
<taxonomicName id="4C764D33C37B9771FE4D7554FD43FDD8" box="[469,680,559,584]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="29" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37B9771FE4D7554FD43FDD8" box="[469,680,559,584]" italics="true" pageId="2" pageNumber="29">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37B9771FD297554FD1AFDD9" box="[689,753,559,585]" pageId="2" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from all congeners on the basis of the following combination of characters: 1) rostral wider than high, portion visible from above longer than prefrontal suture; prefrontal shorter than frontal, touching first two supralabials; 2) single nasal; 3) frontal hexagonal, more than 2.5 times maximum width of supraocular, and about two­thirds length of parietal; 4) paraparietal surrounded by six shields and scales; 5) eye diameter larger than eye­mouth distance; 6) preocular absent; 7) absent loreals and anterior temporals; 8) four supralabials, second and third entering orbit; fourth supralabial longest and in broad contact with parietal, third ¾ of second, first slightly shorter than third; 9) modified maxillary teeth; 10) five infralabials, three touching anterior chin shield; 11) mental not touching anterior chin shields; only first pair of chin shields meeting in midline; 12) three gular scales in midline between posterior chin shields and first ventral; 13) 198 ventrals; 21 divided subcaudal scales; 14) single anal plate; 15) smooth dorsals; dorsal scale row reduction on tail to five opposite last subcaudal anterior to terminal scute; 16) large size (total length up to
<quantity id="4C8E9B55C37B9771FCCC734CFC50FBC1" box="[852,955,1079,1105]" metricMagnitude="-1" metricUnit="m" metricValue="4.55" pageId="2" pageNumber="29" unit="mm" value="455.0">455 mm</quantity>
); tail moderately long, tapering gradually to a point; 17) dark bluish greyish dorsum, iridescent; anterior body with four pale zigzag bands; yellowish markings on dorsum of tail base and tail tip; 18) ventral side of body immaculate beige yellowish; underside of head greyish brownish; beige yellowish tail base with small dark flecks and dark outermost corners of subcaudals.
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37B9770FEA07385FE96FB41" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="description">
<paragraph id="8BC936B0C37B9771FEA07385FAC0F8D9" blockId="2.[264,1323,398,1865]" pageId="2" pageNumber="29">
<emphasis id="B902EAA2C37B9771FEA07385FD65FA88" bold="true" box="[312,654,1278,1304]" pageId="2" pageNumber="29">
Description of
<typeStatus id="54CD8812C37B9771FE667385FD80FA88" box="[510,619,1278,1304]" pageId="2" pageNumber="29" type="holotype">holotype</typeStatus>
.—
</emphasis>
Rostral wider (
<quantity id="4C8E9B55C37B9771FCD37384FC44FA89" box="[843,943,1279,1305]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="29" unit="mm" value="2.6">2.6 mm</quantity>
) than high (
<quantity id="4C8E9B55C37B9771FBCC7384FB51FA89" box="[1108,1210,1279,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="29" unit="mm" value="1.8">1.8 mm</quantity>
), portion visible from above longer (
<quantity id="4C8E9B55C37B9771FDCD725CFD5FFAD1" box="[597,692,1319,1345]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="29" unit="mm" value="1.7">1.7 mm</quantity>
) than prefrontal suture (
<quantity id="4C8E9B55C37B9771FC47725CFBA9FAD1" box="[991,1090,1319,1345]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="29" unit="mm" value="1.3">1.3 mm</quantity>
). Prefrontal shorter (
<quantity id="4C8E9B55C37B9771FE887234FE85FAF9" box="[272,366,1359,1385]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="29" unit="mm" value="2.7">2.7 mm</quantity>
) than frontal (
<quantity id="4C8E9B55C37B9771FD817234FD91FAF9" box="[537,634,1359,1385]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="29" unit="mm" value="3.2">3.2 mm</quantity>
), touching first two supralabials. Frontal hexagonal, more than 2.5 times (
<quantity id="4C8E9B55C37B9771FE24720CFDF2FA01" box="[444,537,1399,1425]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="29" unit="mm" value="2.6">2.6 mm</quantity>
) maximum width of supraocular (1.0 mm), length about two­thirds that of the parietal (
<quantity id="4C8E9B55C37B9771FE6372E4FDB4FA29" box="[507,607,1439,1465]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="29" unit="mm" value="4.7">4.7 mm</quantity>
). Parietal about 1.7 times length of prefrontal. Paraparietal surrounded by six shields and scales. Nasal smaller than postocular. No preocular. Postocular higher than wide, not as high (
<quantity id="4C8E9B55C37B9771FD737294FCA3F999" box="[747,840,1519,1545]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="29" unit="mm" value="0.9">0.9 mm</quantity>
) as eye diameter (
<quantity id="4C8E9B55C37B9771FB837294FB91F999" box="[1051,1146,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="29" unit="mm" value="1.4">1.4 mm</quantity>
). Eye diameter larger than eye­mouth distance (1.0 mm). Distance from anterior corner of eye to centre of nostril
<quantity id="4C8E9B55C37B9771FEC37144FE52F9C9" box="[347,441,1599,1625]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="29" unit="mm" value="2.4">2.4 mm</quantity>
, and from anterior corner of eye to tip of snout
<quantity id="4C8E9B55C37B9771FC767144FBA0F9C9" box="[1006,1099,1599,1625]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="29" unit="mm" value="3.8">3.8 mm</quantity>
. Four supralabials, second and third entering orbit, fourth longest, third ¾ of second, first slightly shorter than third and about half as long as second. Mental semicircular to triangular, not touching anterior chin shields. Five infralabials, three touching anterior chin shield. First pair of chin shields meeting in midline, second pair diverging and only touching anterior. Three gulars in midline between posterior chin shields and first ventral. 198 ventral scales, and 21 divided subcaudals. A single anal scale. Thirteen smooth scales across the midbody.
</paragraph>
<paragraph id="8BC936B0C37A9770FE907664FEBDFE19" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">The number of dorsal scale rows is reduced to six rows on tail opposite 8th subcaudal posterior to cloaca (reduction to five rows only opposite last subcaudal anterior to terminal scute).</paragraph>
<paragraph id="8BC936B0C37A9770FEA076ECFC93FDB9" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
Snout­vent length
<quantity id="4C8E9B55C37A9770FD8C76ECFD91FE21" box="[532,634,407,433]" metricMagnitude="-1" metricUnit="m" metricValue="4.24" pageId="3" pageNumber="30" unit="mm" value="424.0">424 mm</quantity>
, tail length
<quantity id="4C8E9B55C37A9770FC9076ECFC8BFE21" box="[776,864,407,433]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="30" unit="mm" value="31.0">31 mm</quantity>
. Body thickness about
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, tail root
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thick. Tail moderately long, relatively thin, tapering gradually to a point. Ratio of tail to total length (
<quantity id="4C8E9B55C37A9770FDC5769CFD28FD91" box="[605,707,487,513]" metricMagnitude="-1" metricUnit="m" metricValue="4.55" pageId="3" pageNumber="30" unit="mm" value="455.0">455 mm</quantity>
) is 0.068. Vermiform habitus, head indistinct from neck. Modified (most probably nine) maxillary teeth.
</paragraph>
<paragraph id="8BC936B0C37A9770FEA0754CFE96FB41" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
Colour preserved in ethanol dark bluish to greyish above, iridescent. Parietals and subsequent scales with indistinct light beige to yellowish markings. Lower parts of supralabials yellowish, most distinct below and somewhat behind the eyes. Dorsal scales posterior in part with more or less light markings, especially on the sides of the anterior body half. Outermost dorsal scale rows light yellowish beige below. Four beige to yellowish dorsal bands are well discernible in the anterior body half: each band is half a scale to two scales broad, somewhat zigzag shaped and turns into the yellowish beige coloration of the venter. The first yellowish dorsal band is located about
<quantity id="4C8E9B55C37A9770FBF27434FB4DFCF9" box="[1130,1190,847,873]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="30" unit="cm" value="1.0">1 cm</quantity>
behind the head, the others follow each with distances of about
<quantity id="4C8E9B55C37A9770FCF4740CFC54FC01" box="[876,959,887,913]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="30" unit="cm" value="3.5">3.5 cm</quantity>
. Small yellowish markings can be found on the dorsum of the tail base as well as on the dorsal tail tip: whereas the anterior light dorsal area, situated about half a centimetre behind the anal scale, is recognizable as rudiment of a thin zigzag shaped band, comprising five scale rows, the yellow fleck on the tail’s end covers only the hind part of a single scale bordering the tail tip. The ventral side of the body is immaculate yellowish to beige. Similar to the body venter the tail base is yellowish beige except for some small dark flecks and the dark outermost corners of the subcaudals. The underside of the head appears greyish to brownish.
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37A9770FEA073A5FBA1F981" pageId="3" pageNumber="30" type="etymology">
<paragraph id="8BC936B0C37A9770FEA073A5FC33FAE1" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
<emphasis id="B902EAA2C37A9770FEA073A5FE09FB68" bold="true" box="[312,482,1246,1272]" pageId="3" pageNumber="30">Etymology.—</emphasis>
We dedicate this new species to our friend and colleague Vu Ngoc Thanh (
<collectingCountry id="F3617620C37A9770FEFD727CFE21FAB1" box="[357,458,1287,1313]" name="Vietnam" pageId="3" pageNumber="30">Vietnam</collectingCountry>
National University, Hanoi, University of Science, Faculty of Biology, Department of Vertebrate Zoology, Zoological Museum) in recognition of his valuable support during numerous biodiversity excursions in
<collectingCountry id="F3617620C37A9770FCF5722CFC3FFAE1" box="[877,980,1367,1393]" name="Vietnam" pageId="3" pageNumber="30">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BC936B0C37A9770FEA07204FB0BFA51" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
<collectingCountry id="F3617620C37A9770FEA07204FEB2FA09" box="[312,345,1407,1433]" name="American Samoa" pageId="3" pageNumber="30">As</collectingCountry>
common names we suggest Thanh’s reed snake (English), Ran mai gam Thanh (Vietnamese), Calamaire de Thanh (French), and Thanhs Zwergschlange (
<collectingCountry id="F3617620C37A9770FBF672DCFB3FFA51" box="[1134,1236,1447,1473]" name="Germany" pageId="3" pageNumber="30">German</collectingCountry>
).
</paragraph>
<paragraph id="8BC936B0C37A9770FEA072B5FBA1F981" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
<emphasis id="B902EAA2C37A9770FEA072B5FDE9FA78" bold="true" box="[312,514,1486,1512]" pageId="3" pageNumber="30">Comparisons.—</emphasis>
<taxonomicName id="4C764D33C37A9770FD9A72B4FD3CFA78" box="[514,727,1487,1512]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37A9770FD9A72B4FD3CFA78" box="[514,727,1487,1512]" italics="true" pageId="3" pageNumber="30">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37A9770FD7A72B4FCCEFA79" box="[738,805,1487,1513]" pageId="3" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
differs according to
<bibRefCitation id="EFE74B41C37A9770FBB072B4FAC7FA79" author="Inger" box="[1064,1324,1487,1513]" pageId="3" pageNumber="38" refString="Inger, R. F. &amp; Marx, H. (1965) The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria. Fieldiana Zoology, 49, 1 - 304." type="journal article" year="1965">Inger &amp; Marx (1965)</bibRefCitation>
and
<bibRefCitation id="EFE74B41C37A9770FEA2728CFD82F981" author="Darevsky" box="[314,617,1527,1553]" lastPageId="3" lastPageNumber="38" pageId="3" pageNumber="37" refString="Darevsky, I. S. &amp; Orlov, N. L. (1992) A new subspecies of the dwarf snake Calamaria lowi ingermarxi ssp. nov. (Serpentes, Colubridae) from southern Vietnam. Asiatic Herpetological Research, 4, 13 - 17." type="journal article" year="1992">Darevsky &amp; Orlov (1992)</bibRefCitation>
from the Vietnamese species as follows:
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37A9774FEA07164FE41F916" lastPageId="7" lastPageNumber="34" pageId="3" pageNumber="30" type="description">
<paragraph id="8BC936B0C37A9770FEA07164FBF9F8B9" blockId="3.[264,1325,284,1833]" pageId="3" pageNumber="30">
<taxonomicName id="4C764D33C37A9770FEA07164FE75F9A8" box="[312,414,1567,1592]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="buchi">
<emphasis id="B902EAA2C37A9770FEA07164FE75F9A8" box="[312,414,1567,1592]" italics="true" pageId="3" pageNumber="30">C. buchi</emphasis>
</taxonomicName>
is blackish above, each dorsal scale with small light spots; in addition,
<taxonomicName id="4C764D33C37A9770FA887164FEA2F9F0" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="buchi">
<emphasis id="B902EAA2C37A9770FA887164FEA2F9F0" italics="true" pageId="3" pageNumber="30">C. buchi</emphasis>
</taxonomicName>
has a higher count of ventral scales (221–236 vs.
<quantity id="4C8E9B55C37A9770FC2E713CFBEFF9F1" box="[950,1028,1607,1633]" metricMagnitude="0" metricUnit="m" metricValue="5.0292" pageId="3" pageNumber="30" unit="in" value="198.0">198 in</quantity>
<taxonomicName id="4C764D33C37A9770FB96713CFB94F9F0" box="[1038,1151,1607,1632]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37A9770FB96713CFB94F9F0" box="[1038,1151,1607,1632]" italics="true" pageId="3" pageNumber="30">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37A9770FB12713CFB26F9F1" box="[1162,1229,1607,1633]" pageId="3" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
), fewer subcaudal scales (13–14 vs. 21), preocular present, rostral higher than wide (vs. rostral wider than high), its mental touching the anterior chin shields (which is not the case in
<taxonomicName id="4C764D33C37A9770FA8871ECFEB8F948" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="30" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37A9770FA8871ECFEB8F948" italics="true" pageId="3" pageNumber="30">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37A9770FEC571C4FE4BF949" box="[349,416,1727,1753]" pageId="3" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
), only five (vs. six) scales touch the paraparietal, dorsal scale reduction to four rows on tail opposite the third to fourth subcaudal anterior to terminal scute, and a ratio of tail to total length of 0.039 to 0.041 (vs. 0.068) in females.
</paragraph>
<caption id="DF096638C37D9777FE907489FEB0FBBB" httpUri="https://zenodo.org/record/169840/files/figure.png" pageId="4" pageNumber="31" targetBox="[264,1322,323,1002]" targetPageId="4">
<paragraph id="8BC936B0C37D9777FE907489FEB0FBBB" blockId="4.[264,1323,1010,1067]" pageId="4" pageNumber="31">
<emphasis id="B902EAA2C37D9777FE907489FE65FB9A" bold="true" box="[264,398,1010,1034]" pageId="4" pageNumber="31">FIGURE 1.</emphasis>
Dorsal view of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37D9777FC5C7489FB6DFB99" box="[964,1158,1010,1033]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37D9777FC5C7489FB6DFB99" box="[964,1158,1010,1033]" italics="true" pageId="4" pageNumber="31">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37D9777FB097489FB25FB99" box="[1169,1230,1010,1033]" pageId="4" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920).
</paragraph>
</caption>
<caption id="DF096638C37D9777FE907060FEB0F8C5" httpUri="https://zenodo.org/record/169841/files/figure.png" pageId="4" pageNumber="31" targetBox="[264,1322,1140,1801]" targetPageId="4">
<paragraph id="8BC936B0C37D9777FE907060FEB0F8C5" blockId="4.[264,1323,1819,1877]" pageId="4" pageNumber="31">
<emphasis id="B902EAA2C37D9777FE907060FE66F8A3" bold="true" box="[264,397,1819,1843]" pageId="4" pageNumber="31">FIGURE 2.</emphasis>
Ventral view of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37D9777FC5E7067FB6CF8A3" box="[966,1159,1820,1843]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="31" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37D9777FC5E7067FB6CF8A3" box="[966,1159,1820,1843]" italics="true" pageId="4" pageNumber="31">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37D9777FB0A7067FB25F8A3" box="[1170,1230,1820,1843]" pageId="4" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920).
</paragraph>
</caption>
<caption id="DF096638C37C9776FE90736FFD52FBDE" httpUri="https://zenodo.org/record/169842/files/figure.png" pageId="5" pageNumber="32" targetBox="[264,1323,326,1020]" targetPageId="5">
<paragraph id="8BC936B0C37C9776FE90736FFD52FBDE" blockId="5.[264,1322,1044,1102]" pageId="5" pageNumber="32">
<emphasis id="B902EAA2C37C9776FE90736FFE7BFBBC" bold="true" box="[264,400,1044,1068]" pageId="5" pageNumber="32">FIGURE 3.</emphasis>
Oblique view of the head and neck region of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37C9776FE90734CFE2DFBDE" box="[264,454,1079,1102]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37C9776FE90734CFE2DFBDE" box="[264,454,1079,1102]" italics="true" pageId="5" pageNumber="32">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37C9776FE55734CFDEDFBDE" box="[461,518,1079,1102]" pageId="5" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920).
</paragraph>
</caption>
<caption id="DF096638C37C9776FE90719EFE5FF8B0" httpUri="https://zenodo.org/record/169843/files/figure.png" pageId="5" pageNumber="32" targetBox="[264,1322,1140,1758]" targetPageId="5">
<paragraph id="8BC936B0C37C9776FE90719EFE5FF8B0" blockId="5.[264,1323,1765,1824]" pageId="5" pageNumber="32">
<emphasis id="B902EAA2C37C9776FE90719EFE61F96D" bold="true" box="[264,394,1765,1789]" pageId="5" pageNumber="32">FIGURE 4.</emphasis>
Lateral view of the head of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37C9776FBB3719DFB02F96D" box="[1067,1257,1766,1789]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="32" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37C9776FBB3719DFB02F96D" box="[1067,1257,1766,1789]" italics="true" pageId="5" pageNumber="32">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37C9776FB69719DFAC0F96D" box="[1265,1323,1766,1789]" pageId="5" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920).
</paragraph>
</caption>
<caption id="DF096638C37F9775FE907374FE5FFBDA" httpUri="https://zenodo.org/record/169844/files/figure.png" pageId="6" pageNumber="33" targetBox="[264,1323,326,1014]" targetPageId="6">
<paragraph id="8BC936B0C37F9775FE907374FE5FFBDA" blockId="6.[264,1323,1039,1098]" pageId="6" pageNumber="33">
<emphasis id="B902EAA2C37F9775FE907374FE61FBB7" bold="true" box="[264,394,1039,1063]" pageId="6" pageNumber="33">FIGURE 5.</emphasis>
Ventral view of the head of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37F9775FBB3736BFB01FBB7" box="[1067,1258,1040,1063]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="33" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37F9775FBB3736BFB01FBB7" box="[1067,1258,1040,1063]" italics="true" pageId="6" pageNumber="33">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37F9775FB69736BFAC0FBB7" box="[1265,1323,1040,1063]" pageId="6" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920).
</paragraph>
</caption>
<caption id="DF096638C37F9775FE907068FD31F8C1" httpUri="https://zenodo.org/record/169845/files/figure.png" pageId="6" pageNumber="33" targetBox="[264,1322,1135,1787]" targetPageId="6">
<paragraph id="8BC936B0C37F9775FE907068FD31F8C1" blockId="6.[264,1323,1811,1873]" pageId="6" pageNumber="33">
<emphasis id="B902EAA2C37F9775FE907068FE66F8BB" bold="true" box="[264,397,1811,1835]" pageId="6" pageNumber="33">FIGURE 6.</emphasis>
Dorsal view of the tail of the ethanol­preserved holotype of
<taxonomicName id="4C764D33C37F9775FBBF7068FB0CF8BA" box="[1063,1255,1811,1834]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="33" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37F9775FBBF7068FB0CF8BA" box="[1063,1255,1811,1834]" italics="true" pageId="6" pageNumber="33">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37F9775FB687068FAC0F8BA" box="[1264,1323,1811,1834]" pageId="6" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
(ZFMK 82920). Photos by Thomas Ziegler.
</paragraph>
</caption>
<paragraph id="8BC936B0C37E9774FEA07667FBB4FDE6" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="34">
<taxonomicName id="4C764D33C37E9774FEA07667FE1BFEA5" box="[312,496,284,309]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="pavimentata">
<emphasis id="B902EAA2C37E9774FEA07667FE1BFEA5" box="[312,496,284,309]" italics="true" pageId="7" pageNumber="34">C. pavimentata</emphasis>
</taxonomicName>
usually has narrow, dark, longitudinal stripes, a solid black collar immediately behind the neck, ventrals with dark lateral tips, a slightly smaller first supralabial than the second (vs. a first supralabial that is about half as long as second in
<taxonomicName id="4C764D33C37E9774FA887617FEB8FE3D" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FA887617FEB8FE3D" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FEC376EFFE70FE3E" box="[347,411,404,430]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
), the second pair of chin shields meeting in midline (which is not the case in
<taxonomicName id="4C764D33C37E9774FE9076C7FE9EFE45" box="[264,373,444,469]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FE9076C7FE9EFE45" box="[264,373,444,469]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FEE576C7FE55FE46" box="[381,446,444,470]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
, where the second pair diverges and only touches anteriorly), a rostral that is as broad as high (or slightly higher vs. a rostral that is wider than high), dorsal scale row reduction usually to four (vs. five) on tail, a rostral portion visible from above only onehalf to equal entire length of prefrontal suture (vs. a rostral portion visible from above longer than prefrontal suture in
<taxonomicName id="4C764D33C37E9774FDE47527FD02FDE5" box="[636,745,604,629]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FDE47527FD02FDE5" box="[636,745,604,629]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FD687527FCDBFDE6" box="[752,816,604,630]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
), and a preocular present.
</paragraph>
<paragraph id="8BC936B0C37E9774FEA075FFFC3EFC1E" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="34">
<taxonomicName id="4C764D33C37E9774FEA075FFFDE7FD0D" box="[312,524,644,669]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="B902EAA2C37E9774FEA075FFFDE7FD0D" box="[312,524,644,669]" italics="true" pageId="7" pageNumber="34">C. septentrionalis</emphasis>
</taxonomicName>
has dorsal scales with many small light dots forming a network, lower counts of ventral and subcaudal scales in females (168–188 vs. 198 and 6–11 vs. 21), rostral portion visible from above less than one­third length of prefrontal suture (vs. a rostral portion visible from above longer than prefrontal suture in
<taxonomicName id="4C764D33C37E9774FBCA7587FB2DFC85" box="[1106,1222,764,789]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FBCA7587FB2DFC85" box="[1106,1222,764,789]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FB4C7587FAF1FC86" box="[1236,1306,764,790]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
), preocular present, the tail being as thick as body, not tapering, with a broadly rounded tip (vs. a relatively thin tail in
<taxonomicName id="4C764D33C37E9774FDDC7437FD5AFCF5" box="[580,689,844,869]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FDDC7437FD5AFCF5" box="[580,689,844,869]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FD207437FD1DFCF6" box="[696,758,844,870]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
that is tapering gradually to a point), and a ratio of tail to total length of 0.026 to 0.043 (vs. 0.068) in females.
</paragraph>
<paragraph id="8BC936B0C37E9774FEA074E7FC60FB5E" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="34">
<taxonomicName id="4C764D33C37E9774FEA074E7FDBDFC25" box="[312,598,924,949]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="subSpecies" species="lovii" subSpecies="ingermarxorum">
<emphasis id="B902EAA2C37E9774FEA074E7FDBDFC25" box="[312,598,924,949]" italics="true" pageId="7" pageNumber="34">C. lovii ingermarxorum</emphasis>
</taxonomicName>
has an immaculate grey­bluish dorsum with light spots on each side of the neck covering four scales, uniformly dark grey ventrals with light posterior edges, prefrontals being longer than frontal (vs. prefrontals shorter than frontal in
<taxonomicName id="4C764D33C37E9774FE90736FFE92FBBD" box="[264,377,1044,1069]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FE90736FFE92FBBD" box="[264,377,1044,1069]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FE1C736FFE2DFBBE" box="[388,454,1044,1070]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
), rostral portion visible from above more than half length of prefrontal suture (vs. a rostral portion visible from above longer than prefrontal suture), its mental touching the anterior chin shields, dorsal scale reduction to four rows on tail opposite first to fifth subcaudal anterior to terminal scute, and a short, blunt tail that is not tapering (vs. a relatively thin tail that is tapering gradually to a point).
</paragraph>
<paragraph id="8BC936B0C37E9774FEA073A7FE41F916" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="34">
Our new
<taxonomicName id="4C764D33C37E9774FE3C73A7FDC8FB65" box="[420,547,1244,1269]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B902EAA2C37E9774FE3C73A7FDC8FB65" box="[420,547,1244,1269]" italics="true" pageId="7" pageNumber="34">Calamaria</emphasis>
</taxonomicName>
species can be distinguished from the other subspecies of
<taxonomicName id="4C764D33C37E9774FB5573A7FAC8FB65" box="[1229,1315,1244,1269]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="lovii">
<emphasis id="B902EAA2C37E9774FB5573A7FAC8FB65" box="[1229,1315,1244,1269]" italics="true" pageId="7" pageNumber="34">C. lovii</emphasis>
</taxonomicName>
, that do not occur in
<collectingCountry id="F3617620C37E9774FE6A727FFDB2FA8E" box="[498,601,1284,1310]" name="Vietnam" pageId="7" pageNumber="34">Vietnam</collectingCountry>
, according to
<bibRefCitation id="EFE74B41C37E9774FD64727FFC1FFA8E" author="Inger" box="[764,1012,1284,1310]" pageId="7" pageNumber="38" refString="Inger, R. F. &amp; Marx, H. (1965) The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria. Fieldiana Zoology, 49, 1 - 304." type="journal article" year="1965">Inger &amp; Marx (1965)</bibRefCitation>
e.g. by its quite large total length (
<quantity id="4C8E9B55C37E9774FEFB7257FE23FAD6" box="[355,456,1324,1350]" metricMagnitude="-1" metricUnit="m" metricValue="4.55" pageId="7" pageNumber="34" unit="mm" value="455.0">455 mm</quantity>
vs.
<quantity id="4C8E9B55C37E9774FE637257FDB4FAD6" box="[507,607,1324,1350]" metricMagnitude="-1" metricUnit="m" metricValue="3.16" pageId="7" pageNumber="34" unit="mm" value="316.0">316 mm</quantity>
in
<emphasis id="B902EAA2C37E9774FD1F7257FD13FAD5" box="[647,760,1324,1349]" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FD5C7257FD13FAD5" authority="Boulenger" authorityName="Boulenger" box="[708,760,1324,1349]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="lovii">lovii</taxonomicName>
</emphasis>
Boulenger,
<quantity id="4C8E9B55C37E9774FC147257FC04FAD6" box="[908,1007,1324,1350]" metricMagnitude="-1" metricUnit="m" metricValue="3.15" pageId="7" pageNumber="34" unit="mm" value="315.0">315 mm</quantity>
in
<emphasis id="B902EAA2C37E9774FB807257FB2FFAD5" box="[1048,1220,1324,1349]" italics="true" pageId="7" pageNumber="34">C. l. wermuthi</emphasis>
Inger &amp; Marx, and
<quantity id="4C8E9B55C37E9774FE10722FFE02FAFE" box="[392,489,1364,1390]" metricMagnitude="-1" metricUnit="m" metricValue="2.69" pageId="7" pageNumber="34" unit="mm" value="269.0">269 mm</quantity>
in
<emphasis id="B902EAA2C37E9774FD97722FFD49FAFD" box="[527,674,1364,1389]" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FDD1722FFD49FAFD" authority="Boulenger" authorityName="Boulenger" box="[585,674,1364,1389]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="gimletti">gimletti</taxonomicName>
</emphasis>
Boulenger), the ventral and subcaudal counts (198 and 21 vs. 218–254 and
<quantity id="4C8E9B55C37E9774FE6A7207FDBCFA06" box="[498,599,1404,1430]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="4.572" metricValueMin="2.794" pageId="7" pageNumber="34" unit="in" value="14.5" valueMax="18.0" valueMin="11.0">11–18 in</quantity>
<emphasis id="B902EAA2C37E9774FDC57207FD22FA05" box="[605,713,1404,1429]" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FD0D7207FD22FA05" box="[661,713,1404,1429]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="lovii">lovii</taxonomicName>
</emphasis>
, 256 and
<quantity id="4C8E9B55C37E9774FCA37207FC9DFA06" box="[827,886,1404,1430]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="7" pageNumber="34" unit="in" value="11.0">11 in</quantity>
<emphasis id="B902EAA2C37E9774FCE47207FBC8FA05" box="[892,1059,1404,1429]" italics="true" pageId="7" pageNumber="34">C. l. wermuthi</emphasis>
, and 215–249 and
<quantity id="4C8E9B55C37E9774FA987207FEAFFA2E" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="7" pageNumber="34" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10– 12 in</quantity>
<emphasis id="B902EAA2C37E9774FED472DFFE34FA2D" box="[332,479,1444,1469]" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FE1E72DFFE34FA2D" box="[390,479,1444,1469]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="gimletti">gimletti</taxonomicName>
</emphasis>
), by its eye diameter being larger than the eye­mouth distance and by a differing coloration and pattern; in addition, contrary to the situation in
<taxonomicName id="4C764D33C37E9774FBC972B7FB54FA75" box="[1105,1215,1484,1509]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FBC972B7FB54FA75" box="[1105,1215,1484,1509]" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FB5E72B7FAEDFA76" box="[1222,1286,1484,1510]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
, in
<emphasis id="B902EAA2C37E9774FE90728FFE96F99D" box="[264,381,1524,1549]" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FED1728FFE96F99D" box="[329,381,1524,1549]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="lovii">lovii</taxonomicName>
</emphasis>
and
<emphasis id="B902EAA2C37E9774FE27728FFD9BF99D" box="[447,624,1524,1549]" italics="true" pageId="7" pageNumber="34">C. l. wermuthi</emphasis>
the mental touches the anterior chin shields and in
<emphasis id="B902EAA2C37E9774FB6C728FFE8AF9A5" italics="true" pageId="7" pageNumber="34">
C. l.
<taxonomicName id="4C764D33C37E9774FE907167FE8AF9A5" box="[264,353,1564,1589]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="gimletti">gimletti</taxonomicName>
</emphasis>
the rostral portion visible from above is distinctly smaller than the prefrontal suture (vs. a rostral portion visible from above being longer than prefrontal suture in
<taxonomicName id="4C764D33C37E9774FA88713FFEB8F915" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FA88713FFEB8F915" italics="true" pageId="7" pageNumber="34">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FEC27117FE70F916" box="[346,411,1644,1670]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC37E977BFEA071EFFB14FD56" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34" type="reference_group">
<paragraph id="8BC936B0C37E9774FEA071EFFD75F8DE" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="34">
<taxonomicName id="4C764D33C37E9774FEA071EFFDE2F93D" box="[312,521,1684,1709]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C37E9774FEA071EFFDE2F93D" box="[312,521,1684,1709]" italics="true" pageId="7" pageNumber="34">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C37E9774FD9771EFFDA5F93E" box="[527,590,1684,1710]" pageId="7" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
differs from the Chinese species
<taxonomicName id="4C764D33C37E9774FC4C71EFFB18F93E" authority="Chernov" authorityName="Chernov" box="[980,1267,1684,1710]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="B902EAA2C37E9774FC4C71EFFB6CF93D" box="[980,1159,1684,1709]" italics="true" pageId="7" pageNumber="34">C. yunnanensis</emphasis>
Chernov
</taxonomicName>
, that was judged as doubtful form by
<bibRefCitation id="EFE74B41C37E9774FD1D71C7FC6AF946" author="Inger" box="[645,897,1724,1750]" pageId="7" pageNumber="38" refString="Inger, R. F. &amp; Marx, H. (1965) The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria. Fieldiana Zoology, 49, 1 - 304." type="journal article" year="1965">Inger &amp; Marx (1965)</bibRefCitation>
, but listed as valid by
<bibRefCitation id="EFE74B41C37E9774FB1171C7FEBDF96E" author="Yang" pageId="7" pageNumber="38" refString="Yang, D. &amp; Inger, R. F. (1986) Key to the snakes and lizards of China. Smithsonian Herpetological Information Service, 71, 1 - 21." type="journal article" year="1986">Yang &amp; Inger (1986)</bibRefCitation>
or
<bibRefCitation id="EFE74B41C37E9774FEE6719FFD90F96E" author="Zhao" box="[382,635,1764,1790]" pageId="7" pageNumber="38" refString="Zhao, E. &amp; K. Adler (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Contribution to Herpetology, 10, 522 pp." type="book" year="1993">Zhao &amp; Adler (1993)</bibRefCitation>
, in lacking narrow, dark, elongated stripes along the body, dark edges on the ventrals, and in having a higher ventral count (198 vs. 173) as well as in a lower tail ratio (0.068 vs. 0.082).
</paragraph>
<paragraph id="8BC936B0C371977BFEA07667FB14FD56" blockId="8.[264,1323,284,1870]" pageId="8" pageNumber="35">
From the four
<taxonomicName id="4C764D33C371977BFE7D7667FD8FFEA5" box="[485,612,284,309]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B902EAA2C371977BFE7D7667FD8FFEA5" box="[485,612,284,309]" italics="true" pageId="8" pageNumber="35">Calamaria</emphasis>
</taxonomicName>
species, that reach Malay Peninsula (
<taxonomicName id="4C764D33C371977BFBBC7667FAE8FEA6" authority="Gray" authorityName="Gray" box="[1060,1283,284,310]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="albiventer">
<emphasis id="B902EAA2C371977BFBBC7667FB54FEA5" box="[1060,1215,284,309]" italics="true" pageId="8" pageNumber="35">C. albiventer</emphasis>
Gray
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFA887667FD41FECE" authority="Grismer, Kaiser &amp; Yaakob" authorityName="Grismer, Kaiser &amp; Yaakob" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B902EAA2C371977BFA887667FEA5FECD" italics="true" pageId="8" pageNumber="35">C. ingeri</emphasis>
Grismer, Kaiser &amp; Yaakob
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFD24763FFBA9FECE" authority="Dumeril &amp; Bibron" authorityName="Dumeril &amp; Bibron" box="[700,1090,324,350]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="schlegeli">
<emphasis id="B902EAA2C371977BFD24763FFCA6FECD" box="[700,845,324,349]" italics="true" pageId="8" pageNumber="35">C. schlegeli</emphasis>
Duméril &amp; Bibron
</taxonomicName>
) and
<collectingCountry id="F3617620C371977BFB08763FFB13FECE" box="[1168,1272,324,350]" name="Thailand" pageId="8" pageNumber="35">Thailand</collectingCountry>
(
<taxonomicName id="4C764D33C371977BFA88763FFE06FE16" authority="Boie" authorityName="Boie" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="B902EAA2C371977BFA88763FFE4FFE15" italics="true" pageId="8" pageNumber="35">C. lumbricoidea</emphasis>
Boie
</taxonomicName>
) in the north,
<taxonomicName id="4C764D33C371977BFD2A7617FCC3FE15" box="[690,808,364,389]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFD2A7617FCC3FE15" box="[690,808,364,389]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFCAF7617FC95FE16" box="[823,894,364,390]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
is distinguishable in lacking five supralabials, of which the third and fourth entering the orbit (namely four supralabials with second and third entering orbit in
<taxonomicName id="4C764D33C371977BFD0576C7FCE7FE45" box="[669,780,444,469]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFD0576C7FCE7FE45" box="[669,780,444,469]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFC8C76C7FCBEFE46" box="[788,853,444,470]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
), and in lacking a preocular (present in
<taxonomicName id="4C764D33C371977BFE90769FFE4BFE6D" box="[264,416,484,509]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="albiventer">
<emphasis id="B902EAA2C371977BFE90769FFE4BFE6D" box="[264,416,484,509]" italics="true" pageId="8" pageNumber="35">C. albiventer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFE37769FFDFCFE6D" box="[431,535,484,509]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B902EAA2C371977BFE37769FFDFCFE6D" box="[431,535,484,509]" italics="true" pageId="8" pageNumber="35">C. ingeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D33C371977BFDC9769FFCE4FE6D" box="[593,783,484,509]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="B902EAA2C371977BFDC9769FFCE4FE6D" box="[593,783,484,509]" italics="true" pageId="8" pageNumber="35">C. lumbricoidea</emphasis>
</taxonomicName>
), in lacking a mental that touches the anterior chin shields (as is present in
<taxonomicName id="4C764D33C371977BFDCF7577FD1BFDB5" box="[599,752,524,549]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="albiventer">
<emphasis id="B902EAA2C371977BFDCF7577FD1BFDB5" box="[599,752,524,549]" italics="true" pageId="8" pageNumber="35">C. albiventer</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D33C371977BFCB07577FC0DFDB5" box="[808,998,524,549]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="B902EAA2C371977BFCB07577FC0DFDB5" box="[808,998,524,549]" italics="true" pageId="8" pageNumber="35">C. lumbricoidea</emphasis>
</taxonomicName>
), in having a higher ventral count in females (198 vs.
<quantity id="4C8E9B55C371977BFDDE754FFD23FDDE" box="[582,712,564,590]" metricMagnitude="0" metricUnit="m" metricValue="4.013199999999999" metricValueMax="4.572" metricValueMin="3.4543999999999997" pageId="8" pageNumber="35" unit="in" value="158.0" valueMax="180.0" valueMin="136.0">136­180 in</quantity>
<taxonomicName id="4C764D33C371977BFD4A754FFCB4FDDD" box="[722,863,564,589]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="schlegeli">
<emphasis id="B902EAA2C371977BFD4A754FFCB4FDDD" box="[722,863,564,589]" italics="true" pageId="8" pageNumber="35">C. schlegeli</emphasis>
</taxonomicName>
and vs.
<quantity id="4C8E9B55C371977BFC51754FFBA7FDDE" box="[969,1100,564,590]" metricMagnitude="0" metricUnit="m" metricValue="3.9243" metricValueMax="4.1148" metricValueMin="3.7338" pageId="8" pageNumber="35" unit="in" value="154.5" valueMax="162.0" valueMin="147.0">147­162 in</quantity>
<taxonomicName id="4C764D33C371977BFBCE754FFB1AFDDD" box="[1110,1265,564,589]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="albiventer">
<emphasis id="B902EAA2C371977BFBCE754FFB1AFDDD" box="[1110,1265,564,589]" italics="true" pageId="8" pageNumber="35">C. albiventer</emphasis>
</taxonomicName>
), by having a paraparietal that is surrounded by six and not only by four or five scales or shields as in
<taxonomicName id="4C764D33C371977BFED475FFFDE5FD0D" box="[332,526,644,669]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="B902EAA2C371977BFED475FFFDE5FD0D" box="[332,526,644,669]" italics="true" pageId="8" pageNumber="35">C. lumbricoidea</emphasis>
</taxonomicName>
, as well as in pattern (
<taxonomicName id="4C764D33C371977BFCB175FFFC2DFD0D" box="[809,966,644,669]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="albiventer">
<emphasis id="B902EAA2C371977BFCB175FFFC2DFD0D" box="[809,966,644,669]" italics="true" pageId="8" pageNumber="35">C. albiventer</emphasis>
</taxonomicName>
: body with four narrow light stripes,
<taxonomicName id="4C764D33C371977BFEFB75D7FDC9FD55" box="[355,546,684,709]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumbricoidea">
<emphasis id="B902EAA2C371977BFEFB75D7FDC9FD55" box="[355,546,684,709]" italics="true" pageId="8" pageNumber="35">C. lumbricoidea</emphasis>
</taxonomicName>
: belly with black crossbars,
<taxonomicName id="4C764D33C371977BFCE975D7FC10FD55" box="[881,1019,684,709]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="schlegeli">
<emphasis id="B902EAA2C371977BFCE975D7FC10FD55" box="[881,1019,684,709]" italics="true" pageId="8" pageNumber="35">C. schlegeli</emphasis>
</taxonomicName>
: distinctly bicolored).
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC371977BFEA075AFFD69F9A6" pageId="8" pageNumber="35" type="description">
<paragraph id="8BC936B0C371977BFEA075AFFD69F9A6" blockId="8.[264,1323,284,1870]" pageId="8" pageNumber="35">
Most of the remaining, geographically quite distant
<taxonomicName id="4C764D33C371977BFC0575AFFBF7FD7D" box="[925,1052,724,749]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B902EAA2C371977BFC0575AFFBF7FD7D" box="[925,1052,724,749]" italics="true" pageId="8" pageNumber="35">Calamaria</emphasis>
</taxonomicName>
species from Sumatra, Borneo, Java, the
<collectingCountry id="F3617620C371977BFE7B7587FD81FC86" box="[483,618,764,790]" name="Philippines" pageId="8" pageNumber="35">Philippines</collectingCountry>
, Sulawesi and the
<collectingCountry id="F3617620C371977BFCD77587FC2AFC86" box="[847,961,764,790]" name="Indonesia" pageId="8" pageNumber="35">Moluccas</collectingCountry>
can be distinguished from
<taxonomicName id="4C764D33C371977BFA887587FEB8FCAD" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFA887587FEB8FCAD" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFEC3745FFE70FCAE" box="[347,411,804,830]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
not only by coloration and pattern but also by the presence of the (due to its limited intraspecific variation) stable character of five supralabials (four in
<taxonomicName id="4C764D33C371977BFB087437FAEBFCF5" box="[1168,1280,844,869]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFB087437FAEBFCF5" box="[1168,1280,844,869]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFA917437FEF4FC1E" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
), of which, in addition, the third and fourth (and not the second and third as in
<taxonomicName id="4C764D33C371977BFB25740FFAC0FC1D" box="[1213,1323,884,909]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFB25740FFAC0FC1D" box="[1213,1323,884,909]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFE9074E7FEA0FC26" box="[264,331,924,950]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
) enter the orbit (
<taxonomicName id="4C764D33C371977BFDBA74E7FC83FC26" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[546,872,924,950]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="abstrusa">
<emphasis id="B902EAA2C371977BFDBA74E7FD45FC25" box="[546,686,924,949]" italics="true" pageId="8" pageNumber="35">C. abstrusa</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFCEF74E7FB40FC26" authority="Boulenger" authorityName="Boulenger" box="[887,1195,924,950]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="acutirostris">
<emphasis id="B902EAA2C371977BFCEF74E7FBCFFC25" box="[887,1060,924,949]" italics="true" pageId="8" pageNumber="35">C. acutirostris</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFB2374E7FE6FFC4E" authority="Boulenger" authorityName="Boulenger" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="alidae">
<emphasis id="B902EAA2C371977BFB2374E7FAC0FC25" box="[1211,1323,924,949]" italics="true" pageId="8" pageNumber="35">C. alidae</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFE0974BFFD42FC4E" authority="Smith" authorityName="Smith" box="[401,681,964,990]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="apraeocularis">
<emphasis id="B902EAA2C371977BFE0974BFFDB2FC4D" box="[401,601,964,989]" italics="true" pageId="8" pageNumber="35">C. apraeocularis</emphasis>
Smith
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFD2D74BFFBF8FC4E" authority="Dumeril &amp; Bibron" authorityName="Dumeril &amp; Bibron" box="[693,1043,964,990]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B902EAA2C371977BFD2D74BFFCC5FC4D" box="[693,814,964,989]" italics="true" pageId="8" pageNumber="35">C. bicolor</emphasis>
Duméril &amp; Bibron
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFBB874BFFAE8FC4E" authority="Peters" authorityName="Peters" box="[1056,1283,964,990]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="bitorques">
<emphasis id="B902EAA2C371977BFBB874BFFB5AFC4D" box="[1056,1201,964,989]" italics="true" pageId="8" pageNumber="35">C. bitorques</emphasis>
Peters
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFA8874BFFDAAFB96" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="boesemani">
<emphasis id="B902EAA2C371977BFA8874BFFE6EFB95" italics="true" pageId="8" pageNumber="35">C. boesemani</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFDCB7497FC20FB96" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[595,971,1004,1030]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="brongersmai">
<emphasis id="B902EAA2C371977BFDCB7497FCE4FB95" box="[595,783,1004,1029]" italics="true" pageId="8" pageNumber="35">C. brongersmai</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFC437497FB6DFB95" box="[987,1158,1004,1029]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="ceramensis">
<emphasis id="B902EAA2C371977BFC437497FB6DFB95" box="[987,1158,1004,1029]" italics="true" pageId="8" pageNumber="35">C. ceramensis</emphasis>
</taxonomicName>
de Rooij,
<taxonomicName id="4C764D33C371977BFA887497FE70FBBE" authority="Lidth" authorityName="Lidth" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="crassa">
<emphasis id="B902EAA2C371977BFA887497FEBFFBBD" italics="true" pageId="8" pageNumber="35">C. crassa</emphasis>
Lidth
</taxonomicName>
de Jeude,
<taxonomicName id="4C764D33C371977BFD8E736FFD12FBBE" authority="Boulenger" authorityName="Boulenger" box="[534,761,1044,1070]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="curta">
<emphasis id="B902EAA2C371977BFD8E736FFD9DFBBD" box="[534,630,1044,1069]" italics="true" pageId="8" pageNumber="35">C. curta</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFC9D736FFBECFBBE" authority="Gough" authorityName="Gough" box="[773,1031,1044,1070]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="doederleini">
<emphasis id="B902EAA2C371977BFC9D736FFC47FBBD" box="[773,940,1044,1069]" italics="true" pageId="8" pageNumber="35">C. doederleini</emphasis>
Gough
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFB8A736FFACDFBBE" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[1042,1318,1044,1070]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="eiselti">
<emphasis id="B902EAA2C371977BFB8A736FFB91FBBD" box="[1042,1146,1044,1069]" italics="true" pageId="8" pageNumber="35">C. eiselti</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFE907347FDF8FBC6" authority="Boulenger" authorityName="Boulenger" box="[264,531,1084,1110]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="everetti">
<emphasis id="B902EAA2C371977BFE907347FE63FBC5" box="[264,392,1084,1109]" italics="true" pageId="8" pageNumber="35">C. everetti</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFDBD7347FC80FBC6" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[549,875,1084,1110]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="forcarti">
<emphasis id="B902EAA2C371977BFDBD7347FD4CFBC5" box="[549,679,1084,1109]" italics="true" pageId="8" pageNumber="35">C. forcarti</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFCE57347FB15FBC6" authority="Dumeril &amp; Bibron" authorityName="Dumeril &amp; Bibron" box="[893,1278,1084,1110]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="gervaisi">
<emphasis id="B902EAA2C371977BFCE57347FBEEFBC5" box="[893,1029,1084,1109]" italics="true" pageId="8" pageNumber="35">C. gervaisi</emphasis>
Duméril &amp; Bibron
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFA977347FE1FFBEE" authority="Fischer" authorityName="Fischer" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="grabowskyi">
<emphasis id="B902EAA2C371977BFA977347FE64FBED" italics="true" pageId="8" pageNumber="35">C. grabowskyi</emphasis>
Fischer
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFD9E731FFCCEFBEE" authority="Loveridge" authorityName="Loveridge" box="[518,805,1124,1150]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="griswoldi">
<emphasis id="B902EAA2C371977BFD9E731FFD76FBED" box="[518,669,1124,1149]" italics="true" pageId="8" pageNumber="35">C. griswoldi</emphasis>
Loveridge
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFCAE731FFB6DFBEE" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[822,1158,1124,1150]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="hilleniusi">
<emphasis id="B902EAA2C371977BFCAE731FFC20FBED" box="[822,971,1124,1149]" italics="true" pageId="8" pageNumber="35">C. hilleniusi</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFB03731FFEBDFB36" authority="Taylor" authorityName="Taylor" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="joloensis">
<emphasis id="B902EAA2C371977BFB03731FFAC1FBED" box="[1179,1322,1124,1149]" italics="true" pageId="8" pageNumber="35">C. joloensis</emphasis>
Taylor
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFEFA73F7FD80FB36" authority="Mocquard" authorityName="Mocquard" box="[354,619,1164,1190]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B902EAA2C371977BFEFA73F7FE0CFB35" box="[354,487,1164,1189]" italics="true" pageId="8" pageNumber="35">C. lateralis</emphasis>
Mocquard
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFDEF73F7FCE9FB35" box="[631,770,1164,1189]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lautensis">
<emphasis id="B902EAA2C371977BFDEF73F7FCE9FB35" box="[631,770,1164,1189]" italics="true" pageId="8" pageNumber="35">C. lautensis</emphasis>
</taxonomicName>
de Rooij,
<taxonomicName id="4C764D33C371977BFCE673F7FB65FB36" authority="Bleeker" authorityName="Bleeker" box="[894,1166,1164,1190]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="leucogaster">
<emphasis id="B902EAA2C371977BFCE673F7FBC1FB35" box="[894,1066,1164,1189]" italics="true" pageId="8" pageNumber="35">C. leucogaster</emphasis>
Bleeker
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFB0373F7FE63FB5E" authority="Andersson" authorityName="Andersson" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lumholtzi">
<emphasis id="B902EAA2C371977BFB0373F7FAC0FB35" box="[1179,1323,1164,1189]" italics="true" pageId="8" pageNumber="35">C. lumholtzi</emphasis>
Andersson
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFE0373CFFD02FB5E" authority="Bleeker" authorityName="Bleeker" box="[411,745,1204,1230]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="margaritophora">
<emphasis id="B902EAA2C371977BFE0373CFFD94FB5D" box="[411,639,1204,1229]" italics="true" pageId="8" pageNumber="35">C. margaritophora</emphasis>
Bleeker
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFD6373CFFC11FB5E" authority="Schenkel" authorityName="Schenkel" box="[763,1018,1204,1230]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="mecheli">
<emphasis id="B902EAA2C371977BFD6373CFFC94FB5D" box="[763,895,1204,1229]" italics="true" pageId="8" pageNumber="35">C. mecheli</emphasis>
Schenkel
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFB9573CFFEB7FB66" authority="Dumeril &amp; Bibron" authorityName="Dumeril &amp; Bibron" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="modesta">
<emphasis id="B902EAA2C371977BFB9573CFFB7CFB5D" box="[1037,1175,1204,1229]" italics="true" pageId="8" pageNumber="35">C. modesta</emphasis>
Duméril &amp; Bibron
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFEF173A7FD85FB66" authority="Boulenger" authorityName="Boulenger" box="[361,622,1244,1270]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="B902EAA2C371977BFEF173A7FE04FB65" box="[361,495,1244,1269]" italics="true" pageId="8" pageNumber="35">C. muelleri</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFDE673A7FC62FB66" authority="Boulenger" authorityName="Boulenger" box="[638,905,1244,1270]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="nuchalis">
<emphasis id="B902EAA2C371977BFDE673A7FCEEFB65" box="[638,773,1244,1269]" italics="true" pageId="8" pageNumber="35">C. nuchalis</emphasis>
Boulenger
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFC0E73A7FAE9FB66" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" box="[918,1282,1244,1270]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="palavanensis">
<emphasis id="B902EAA2C371977BFC0E73A7FBB9FB65" box="[918,1106,1244,1269]" italics="true" pageId="8" pageNumber="35">C. palavanensis</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFA8873A7FE4CFA8E" authority="Lidth" authorityName="Lidth" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="prakkei">
<emphasis id="B902EAA2C371977BFA8873A7FE8BFA8D" italics="true" pageId="8" pageNumber="35">C. prakkei</emphasis>
Lidth
</taxonomicName>
de Jeude,
<taxonomicName id="4C764D33C371977BFDB9727FFCC2FA8E" authority="Bleeker" authorityName="Bleeker" box="[545,809,1284,1310]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="rebentischi">
<emphasis id="B902EAA2C371977BFDB9727FFD2FFA8D" box="[545,708,1284,1309]" italics="true" pageId="8" pageNumber="35">C. rebentischi</emphasis>
Bleeker
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFCAC727FFBFFFA8E" authority="Taylor" authorityName="Taylor" box="[820,1044,1284,1310]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="suluensis">
<emphasis id="B902EAA2C371977BFCAC727FFC29FA8D" box="[820,962,1284,1309]" italics="true" pageId="8" pageNumber="35">C. suluensis</emphasis>
Taylor
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFBBA727FFACDFA8E" authority="Edeling" authorityName="Edeling" box="[1058,1318,1284,1310]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="sumatrana">
<emphasis id="B902EAA2C371977BFBBA727FFB2AFA8D" box="[1058,1217,1284,1309]" italics="true" pageId="8" pageNumber="35">C. sumatrana</emphasis>
Edeling
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFE907257FE3CFAD6" authority="Sackett" authorityName="Sackett" box="[264,471,1324,1350]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="ulmeri">
<emphasis id="B902EAA2C371977BFE907257FE9CFAD5" box="[264,375,1324,1349]" italics="true" pageId="8" pageNumber="35">C. ulmeri</emphasis>
Sackett
</taxonomicName>
, and
<taxonomicName id="4C764D33C371977BFD8E7257FD0CFAD6" authority="Boie" authorityName="Boie" box="[534,743,1324,1350]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="virgulata">
<emphasis id="B902EAA2C371977BFD8E7257FD48FAD5" box="[534,675,1324,1349]" italics="true" pageId="8" pageNumber="35">C. virgulata</emphasis>
Boie
</taxonomicName>
). Of the remaining species,
<taxonomicName id="4C764D33C371977BFBB77257FEA1FAFE" authority="Inger &amp; Marx" authorityName="Inger &amp; Marx" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="battersbyi">
<emphasis id="B902EAA2C371977BFBB77257FB2DFAD5" box="[1071,1222,1324,1349]" italics="true" pageId="8" pageNumber="35">C. battersbyi</emphasis>
Inger &amp; Marx
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFECE722FFDB7FAFE" authority="Bleeker" authorityName="Bleeker" box="[342,604,1364,1390]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="B902EAA2C371977BFECE722FFE13FAFD" box="[342,504,1364,1389]" italics="true" pageId="8" pageNumber="35">C. borneensis</emphasis>
Bleeker
</taxonomicName>
,
<taxonomicName id="4C764D33C371977BFDF0722FFCF6FAFE" authority="Boie" authorityName="Boie" box="[616,797,1364,1390]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="linnaei">
<emphasis id="B902EAA2C371977BFDF0722FFD30FAFD" box="[616,731,1364,1389]" italics="true" pageId="8" pageNumber="35">C. linnaei</emphasis>
Boie
</taxonomicName>
, and
<taxonomicName id="4C764D33C371977BFCC3722FFBF3FAFE" authority="Jan" authorityName="Jan" box="[859,1048,1364,1390]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="melanota">
<emphasis id="B902EAA2C371977BFCC3722FFC01FAFD" box="[859,1002,1364,1389]" italics="true" pageId="8" pageNumber="35">C. melanota</emphasis>
Jan
</taxonomicName>
possess a preocular,
<taxonomicName id="4C764D33C371977BFA88722FFDF3FA06" authority="Marx &amp; Inger" authorityName="Marx &amp; Inger" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B902EAA2C371977BFA88722FFE80FA05" italics="true" pageId="8" pageNumber="35">C. schmidti</emphasis>
Marx &amp; Inger
</taxonomicName>
bears unmodified maxillary teeth,
<taxonomicName id="4C764D33C371977BFC2E7207FB3BFA06" authority="Gunther" authorityName="Gunther" box="[950,1232,1404,1430]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="gracillima">
<emphasis id="B902EAA2C371977BFC2E7207FBBFFA05" box="[950,1108,1404,1429]" italics="true" pageId="8" pageNumber="35">C. gracillima</emphasis>
(Günther)
</taxonomicName>
has the supraocular fused with the postocular, and
<taxonomicName id="4C764D33C371977BFC9672DFFBF0FA2E" authority="Boulenger" authorityName="Boulenger" box="[782,1051,1444,1470]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="javanica">
<emphasis id="B902EAA2C371977BFC9672DFFC73FA2D" box="[782,920,1444,1469]" italics="true" pageId="8" pageNumber="35">C. javanica</emphasis>
Boulenger
</taxonomicName>
has, besides distinctly lower subcaudal counts in females (10–12 vs. 21), four gulars (vs. three gulars in
<taxonomicName id="4C764D33C371977BFB2672B7FAC0FA75" box="[1214,1323,1484,1509]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFB2672B7FAC0FA75" box="[1214,1323,1484,1509]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFE90728FFEADF99E" box="[264,326,1524,1550]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
) in midline between posterior chin shields and first ventral, which clearly distinguish all of them from
<taxonomicName id="4C764D33C371977BFE577167FDD7F9A5" box="[463,572,1564,1589]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFE577167FDD7F9A5" box="[463,572,1564,1589]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFDDB7167FD69F9A6" box="[579,642,1564,1590]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36C653BC371977AFEA0713FFD65FE16" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="distribution">
<paragraph id="8BC936B0C371977BFEA0713FFD02F93E" blockId="8.[264,1323,284,1870]" pageId="8" pageNumber="35">
<emphasis id="B902EAA2C371977BFEA0713FFD12F9CE" bold="true" box="[312,761,1604,1630]" pageId="8" pageNumber="35">Distribution and Natural History.—</emphasis>
<taxonomicName id="4C764D33C371977BFD61713FFC20F9CD" box="[761,971,1604,1629]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFD61713FFC20F9CD" box="[761,971,1604,1629]" italics="true" pageId="8" pageNumber="35">Calamaria thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFC4A713FFBFAF9CE" box="[978,1041,1604,1630]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
is only known from the
<typeStatus id="54CD8812C371977BFE907117FED1F916" box="[264,314,1644,1670]" pageId="8" pageNumber="35">type</typeStatus>
locality. Records for
<collectingCountry id="F3617620C371977BFDA17117FD99F916" box="[569,626,1644,1670]" name="Laos" pageId="8" pageNumber="35">Laos</collectingCountry>
are to be expected in the future, due to the close proximity of the
<typeStatus id="54CD8812C371977BFECB71EFFE6DF93E" box="[339,390,1684,1710]" pageId="8" pageNumber="35">type</typeStatus>
locality to the Laotian border.
</paragraph>
<paragraph id="8BC936B0C371977AFEA071C7FD65FE16" blockId="8.[264,1323,284,1870]" lastBlockId="9.[264,1325,284,390]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
The snake was found heavily injured on the ground of a large limestone cave under a mist net. Further records of that species must confirm, whether
<taxonomicName id="4C764D33C371977BFC6F719FFB8EF96D" box="[1015,1125,1764,1789]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="thanhi" status="sp. nov.">
<emphasis id="B902EAA2C371977BFC6F719FFB8EF96D" box="[1015,1125,1764,1789]" italics="true" pageId="8" pageNumber="35">C. thanhi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157D9C371977BFBF4719FFB47F96E" box="[1132,1196,1764,1790]" pageId="8" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
is adapted to primary limestone forest caves. The adult female, which was collected during the dry season, contained at least seven large eggs besides smaller ova in the ovaries; folded oviducts indicated, that eggs had already been laid. Concerning the larger eggs, mean egg length measured
<quantity id="4C8E9B55C370977AFE4B763FFD7EFECE" box="[467,661,324,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.22" metricValueMin="0.9199999999999999" pageId="9" pageNumber="36" unit="mm" value="10.7" valueMax="12.2" valueMin="9.2">10.7 +/­ 1.5 mm</quantity>
(min. 8.0, max.
<quantity id="4C8E9B55C370977AFCC4763FFC21FECE" box="[860,970,324,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="36" unit="mm" value="12.2">12.2 mm</quantity>
), mean egg width was
<quantity id="4C8E9B55C370977AFB45763FFE89FE16" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="4.3" metricValueMin="2.5" pageId="9" pageNumber="36" unit="mm" value="3.4" valueMax="4.3" valueMin="2.5">3.4 +/­ 0.9 mm</quantity>
(min. 2.3, max.
<quantity id="4C8E9B55C370977AFDBB7617FD69FE16" box="[547,642,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="36" unit="mm" value="4.5">4.5 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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