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<document id="760374E40E76323ACBE7E0678E4E274D" ID-DOI="10.25221/fee.451.4" ID-ISSN="2713-2196" ID-Zenodo-Dep="7166599" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665255292770" checkinUser="felipe" docAuthor="Lin, J. - Z." docDate="2022" docId="F304878DFF9AFFFE890B3175FBBE300E" docLanguage="en" docName="FarEastEntomol.451.19-28.pdf" docOrigin="Far Eastern Entomologist 451" docSource="http://dx.doi.org/10.25221/fee.451.4" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.2:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Nigidius distinctus Parry 1873" docType="key" docVersion="4" lastPageNumber="26" masterDocId="0F3DFFF5FF9DFFF98838331BFFDA357E" masterDocTitle="A new species of the genus Nigidius MacLeay (Coleoptera: Lucanidae) from the Green Island, Eastern Taiwan" masterLastPageNumber="28" masterPageNumber="19" pageNumber="26" updateTime="1699372226084" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="981CE35A7A8B637AF2FAF27A38865331">
<mods:title id="5F3ECAA23C234322D65A60D3CBC61E64">A new species of the genus Nigidius MacLeay (Coleoptera: Lucanidae) from the Green Island, Eastern Taiwan</mods:title>
</mods:titleInfo>
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<mods:namePart id="E91609F2261E309DE434FC70CDD4E6E7">Lin, J. - Z.</mods:namePart>
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<mods:typeOfResource id="44C1D118EC7CE7F22E2380A6D29773DF">text</mods:typeOfResource>
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<mods:title id="A4263C3905CA7F2BBC20C1011D9CB51F">Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part id="E215DD921F06504B836A2EAB287FB2EE">
<mods:date id="770BA6E49992CE74212F573A67BD59EB">2022</mods:date>
<mods:detail id="6A67D278E3D67227DC5DCF77330AF254" type="pubDate">
<mods:number id="5E15516445B5BBB5BE40E6CC1D76C03F">2022-03-15</mods:number>
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<mods:detail id="33149A5CFBE6D1150DE367469FAF3E41" type="volume">
<mods:number id="542C69310F9444A1A194693FE5AE1018">451</mods:number>
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<mods:start id="E4538EAD78B216F815FFA75DE6EDFF66">19</mods:start>
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<mods:url id="91BF58671CC3CB36E4D76BE19C55E35C">http://dx.doi.org/10.25221/fee.451.4</mods:url>
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<mods:classification id="FB1A867BBDB091F2DF319900D9B172A8">journal article</mods:classification>
<mods:identifier id="C44029E8616C5CAA1805CD3BA43BADB9" type="DOI">10.25221/fee.451.4</mods:identifier>
<mods:identifier id="159591A159AE67E5DE74331017A5B216" type="ISSN">2713-2196</mods:identifier>
<mods:identifier id="3702B59433552750CD5B7ADFC48074DD" type="Zenodo-Dep">7166599</mods:identifier>
</mods:mods>
<treatment id="F304878DFF9AFFFE890B3175FBBE300E" ID-DOI="http://doi.org/10.5281/zenodo.7168552" ID-Zenodo-Dep="7168552" LSID="urn:lsid:plazi:treatment:F304878DFF9AFFFE890B3175FBBE300E" httpUri="http://treatment.plazi.org/id/F304878DFF9AFFFE890B3175FBBE300E" lastPageNumber="26" pageId="7" pageNumber="26">
<subSubSection id="33B76510FF9AFFFE890B3175FBCC37FA" box="[307,1046,622,644]" pageId="7" pageNumber="26" type="nomenclature">
<paragraph id="7B12369BFF9AFFFE890B3175FBCC37FA" blockId="7.[307,1046,622,644]" box="[307,1046,622,644]" pageId="7" pageNumber="26">
<heading id="205A81F7FF9AFFFE890B3175FBCC37FA" box="[307,1046,622,644]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="26" reason="2">
<emphasis id="49D9EA89FF9AFFFE890B3175FBCC37FA" bold="true" box="[307,1046,622,644]" pageId="7" pageNumber="26">
Key to the species of the
<taxonomicName id="BCAD4D18FF9AFFFE8A173175FD3237FA" ID-CoL="6RY86" authorityName="Parry" authorityYear="1873" box="[559,744,622,644]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="49D9EA89FF9AFFFE8A173175FD3237FA" bold="true" box="[559,744,622,644]" italics="true" pageId="7" pageNumber="26">Nigidius distinctus</emphasis>
</taxonomicName>
species group from
<collectingCountry id="03BA760BFF9AFFFE8B8E3175FBCC37FA" box="[950,1046,622,644]" name="China" pageId="7" pageNumber="26">East Asia</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="33B76510FF9AFFFE88DB31B2FBBE300E" pageId="7" pageNumber="26" type="key">
<keyStep id="C0592E3EFF9AFFFE88DB31B2FBBE3652" pageId="7" pageNumber="26">
<paragraph id="7B12369BFF9AFFFE88DB31B2FBBA3788" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB31B2FBBA3788" pageId="7" pageNumber="26">
1. Canthus with outer margin entirely outside of the eye laterally. Pronotum without a minute central process at posterior carina.
<collectingCountry id="03BA760BFF9AFFFE8A5131DEFD1337A4" box="[617,713,709,730]" name="Myanmar" pageId="7" pageNumber="26">Myanmar</collectingCountry>
,
<collectingCountry id="03BA760BFF9AFFFE8AEF31DEFCCE37A5" box="[727,788,709,731]" name="China" pageId="7" pageNumber="26">China</collectingCountry>
:
<collectingRegion id="B969F879FF9AFFFE8B1831DEFCB537A4" box="[800,879,709,730]" country="China" name="Yunnan" pageId="7" pageNumber="26">Yunnan</collectingRegion>
. (
<figureCitation id="E3962A1EFF9AFFFE8BBB31DEFC1637A4" box="[899,972,709,731]" captionStart="Figs 13–23" captionStartId="4.[265,306,1552,1574]" captionTargetBox="[251,1105,282,1501]" captionTargetId="figure-0@4.[227,1120,256,1519]" captionTargetPageId="4" captionText="Figs 13–23. Habitus of Nigidius spp., dorsal view. 13–15 – N. wushuae sp. n.; 16–18 – N." figureDoi="http://doi.org/10.5281/zenodo.7166607" httpUri="https://zenodo.org/record/7166607/files/figure.png" pageId="7" pageNumber="26">Fig. 22</figureCitation>
) ....................... .........................................................................................................................
<taxonomicName id="BCAD4D18FF9AFFFE8BDD31FBFBBA3788" ID-CoL="6RY86" authorityName="Parry" authorityYear="1873" box="[997,1120,736,758]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="49D9EA89FF9AFFFE8BDD31FBFBBA3788" bold="true" box="[997,1120,736,758]" italics="true" pageId="7" pageNumber="26">N. distinctus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B12369BFF9AFFFE88DB31E7FBBE3652" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB31E7FBBE3652" pageId="7" pageNumber="26">– Bottom of the emargination of the canthus inside of the outer limit of the eye. Pronotum have a minute central process at posterior carina .............................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0592E3EFF9AFFFE88DB3028FBB9315B" pageId="7" pageNumber="26">
<paragraph id="7B12369BFF9AFFFE88DB3028FBBA36C8" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB3028FBBA36C8" pageId="7" pageNumber="26">
2. Canthus posterior angles more horizontal, with the processes at anterior and posterior angles more subequal in length. Corner lobe of the pronotum is placed more anteriorly than laterally; pronotum distinctly serrate in front. Female genitalia with stylus noticeably stouter and situated at a little prior to the gently rounded distal end.
<collectingCountry id="03BA760BFF9AFFFE8B9F309DFC3E36E2" box="[935,996,902,924]" name="China" pageId="7" pageNumber="26">China</collectingCountry>
:
<collectingCountry id="03BA760BFF9AFFFE8BD6309DFBB936E5" box="[1006,1123,902,923]" name="Hong Kong" pageId="7" pageNumber="26">Hong Kong</collectingCountry>
, Guangdong, Hainan. (
<figureCitation id="E3962A1EFF9AFFFE89E530BAFDF936C9" box="[477,547,929,951]" captionStart="Figs 13–23" captionStartId="4.[265,306,1552,1574]" captionTargetBox="[251,1105,282,1501]" captionTargetId="figure-0@4.[227,1120,256,1519]" captionTargetPageId="4" captionText="Figs 13–23. Habitus of Nigidius spp., dorsal view. 13–15 – N. wushuae sp. n.; 16–18 – N." figureDoi="http://doi.org/10.5281/zenodo.7166607" httpUri="https://zenodo.org/record/7166607/files/figure.png" pageId="7" pageNumber="26">Fig. 23</figureCitation>
) ............................................................................
<taxonomicName id="BCAD4D18FF9AFFFE8BC630B9FBBA36C8" ID-CoL="749VM" authorityName="Schenk" authorityYear="2011" box="[1022,1120,929,951]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="49D9EA89FF9AFFFE8BC630B9FBBA36C8" bold="true" box="[1022,1120,929,951]" italics="true" pageId="7" pageNumber="26">N. sinicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B12369BFF9AFFFE88DB30A6FBB9315B" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB30A6FBB9315B" pageId="7" pageNumber="26">– Canthus with the process at posterior angle much longer than that at anterior angle. Corner lobe of the pronotum is placed more laterally than anteriorly, with a straight, convex or waved inner border on the pronotum. Stylus of female genitalia placed at the inner apex of the hemisternite ............................................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0592E3EFF9AFFFE88DB3730FBB831D1" pageId="7" pageNumber="26">
<paragraph id="7B12369BFF9AFFFE88DB3730FBBA31EA" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB3730FBBA31EA" pageId="7" pageNumber="26">
3. Canthus less developed, with posterior margin more weakly produced. Pronotum with lateral margin distinctly to sinuous in middle. Body relatively shorter and wider. Female genitalia with clearly slender and very sharp stylus.
<collectingCountry id="03BA760BFF9AFFFE8B373778FC933106" box="[783,841,1123,1144]" name="Japan" pageId="7" pageNumber="26">Japan</collectingCountry>
. (
<figureCitation id="E3962A1EFF9AFFFE8B633778FC113106" box="[859,971,1123,1145]" captionStart="Figs 10–12" captionStartId="3.[265,306,1172,1194]" captionTargetBox="[304,1049,663,1158]" captionTargetId="figure-206@3.[272,1082,652,1167]" captionTargetPageId="3" captionText="Figs 10–12. Nigidius lewisi, syntype. 10 – habitus, dorsal view; 11 – the same, ventral" figureDoi="http://doi.org/10.5281/zenodo.7166605" httpUri="https://zenodo.org/record/7166605/files/figure.png" pageId="7" pageNumber="26">Figs 10–12</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE8BE03779FBCC3106" box="[984,1046,1122,1144]" captionStart="Figs 13–23" captionStartId="4.[265,306,1552,1574]" captionTargetBox="[251,1105,282,1501]" captionTargetId="figure-0@4.[227,1120,256,1519]" captionTargetPageId="4" captionText="Figs 13–23. Habitus of Nigidius spp., dorsal view. 13–15 – N. wushuae sp. n.; 16–18 – N." figureDoi="http://doi.org/10.5281/zenodo.7166607" httpUri="https://zenodo.org/record/7166607/files/figure.png" pageId="7" pageNumber="26">16–18</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE8C1A3779FBB93106" box="[1058,1123,1122,1144]" captionStart="Figs 24–29" captionStartId="5.[265,306,1255,1277]" captionTargetBox="[227,1118,566,1217]" captionTargetId="figure-127@5.[227,1122,564,1222]" captionTargetPageId="5" captionText="Figs 24–29. Male (24–26) and female (27–29) genitalia of Nigidius spp. 24, 27 – N." figureDoi="http://doi.org/10.5281/zenodo.7166609" httpUri="https://zenodo.org/record/7166609/files/figure.png" pageId="7" pageNumber="26">25, 28</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE89313765FEF831ED" box="[265,290,1150,1171]" captionStart="Figs 30–33" captionStartId="6.[265,306,1320,1342]" captionTargetBox="[259,1071,702,1295]" captionTargetId="figure-226@6.[248,1106,699,1315]" captionTargetPageId="6" captionText="Figs 30–33. Head and pronotum of Nigidius spp. 30, 31 – N. wushuae sp. n.: 30 – cantus;" figureDoi="http://doi.org/10.5281/zenodo.7166611" httpUri="https://zenodo.org/record/7166611/files/figure.png" pageId="7" pageNumber="26">32</figureCitation>
) .........................................................................................................................
<taxonomicName id="BCAD4D18FF9AFFFE8C333764FBBA31EA" ID-CoL="73Y7R" authorityName="Boileau" authorityYear="1905" box="[1035,1120,1150,1172]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="49D9EA89FF9AFFFE8C333764FBBA31EA" bold="true" box="[1035,1120,1150,1172]" italics="true" pageId="7" pageNumber="26">N. lewisi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B12369BFF9AFFFE88DB3781FBB831D1" blockId="7.[227,1128,681,1393]" box="[227,1122,1178,1200]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB3781FBB831D1" box="[227,1122,1178,1200]" pageId="7" pageNumber="26">– Canthus well developed. Pronotum with lateral weak sinuous in middle ............................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0592E3EFF9AFFFE88DB37ADFBBE300E" pageId="7" pageNumber="26">
<paragraph id="7B12369BFF9AFFFE88DB37ADFB86307C" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB37ADFB86307C" pageId="7" pageNumber="26">
4. Pronotum surface variable with almost impunctate and coarsely punctate areas. Elytra shorter. Female genitalia with strongly hooked and short stylus.
<collectingCountry id="03BA760BFF9AFFFE8BAD37CAFC3B3199" box="[917,993,1233,1255]" name="Taiwan" pageId="7" pageNumber="26">Taiwan</collectingCountry>
: main island. (
<figureCitation id="E3962A1EFF9AFFFE892837F7FEBC307C" box="[272,358,1260,1282]" captionStart="Figs 6–9" captionStartId="2.[265,306,1582,1604]" captionTargetBox="[249,1092,1061,1561]" captionTargetId="figure-97@2.[227,1120,1050,1576]" captionTargetPageId="2" captionText="Figs 6–9. Nigidius lohi, holotype. 6 – habitus, dorsal view; 7 – the same, ventral view; 8 –" figureDoi="http://doi.org/10.5281/zenodo.7166603" httpUri="https://zenodo.org/record/7166603/files/figure.png" pageId="7" pageNumber="26">Figs 6–9</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE894837F6FE74307C" box="[368,430,1261,1282]" captionStart="Figs 13–23" captionStartId="4.[265,306,1552,1574]" captionTargetBox="[251,1105,282,1501]" captionTargetId="figure-0@4.[227,1120,256,1519]" captionTargetPageId="4" captionText="Figs 13–23. Habitus of Nigidius spp., dorsal view. 13–15 – N. wushuae sp. n.; 16–18 – N." figureDoi="http://doi.org/10.5281/zenodo.7166607" httpUri="https://zenodo.org/record/7166607/files/figure.png" pageId="7" pageNumber="26">19–21</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE898037F7FE2C307C" box="[440,502,1260,1282]" captionStart="Figs 24–29" captionStartId="5.[265,306,1255,1277]" captionTargetBox="[227,1118,566,1217]" captionTargetId="figure-127@5.[227,1122,564,1222]" captionTargetPageId="5" captionText="Figs 24–29. Male (24–26) and female (27–29) genitalia of Nigidius spp. 24, 27 – N." figureDoi="http://doi.org/10.5281/zenodo.7166609" httpUri="https://zenodo.org/record/7166609/files/figure.png" pageId="7" pageNumber="26">26, 29</figureCitation>
,
<figureCitation id="E3962A1EFF9AFFFE8A3837F6FDC3307C" box="[512,537,1261,1282]" captionStart="Figs 30–33" captionStartId="6.[265,306,1320,1342]" captionTargetBox="[259,1071,702,1295]" captionTargetId="figure-226@6.[248,1106,699,1315]" captionTargetPageId="6" captionText="Figs 30–33. Head and pronotum of Nigidius spp. 30, 31 – N. wushuae sp. n.: 30 – cantus;" figureDoi="http://doi.org/10.5281/zenodo.7166611" httpUri="https://zenodo.org/record/7166611/files/figure.png" pageId="7" pageNumber="26">33</figureCitation>
) ..................................................................................
<taxonomicName id="BCAD4D18FF9AFFFE8C2037F6FB86307C" ID-CoL="749W3" authorityName="Kawahara, Toguchi et Ochi" authorityYear="2019" box="[1048,1116,1260,1282]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="lohi">
<emphasis id="49D9EA89FF9AFFFE8C2037F6FB86307C" bold="true" box="[1048,1116,1260,1282]" italics="true" pageId="7" pageNumber="26">N. lohi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B12369BFF9AFFFE88DB3612FBBE300E" blockId="7.[227,1128,681,1393]" pageId="7" pageNumber="26">
<keyLead id="C05C95AEFF9AFFFE88DB3612FBBE300E" pageId="7" pageNumber="26">
– Pronotum surface more uniformly punctate. Elytra distinctly longer and slender. Female genitalia with stylus distinctly longer and situated at triangular distal end. – Canthus with posterior margin very strongly produced outward as a fairly sharply pointed projection.
<collectingCountry id="03BA760BFF9AFFFE89313640FE8F300F" box="[265,341,1371,1393]" name="Taiwan" pageId="7" pageNumber="26">Taiwan</collectingCountry>
: Green Island. (
<figureCitation id="E3962A1EFF9AFFFE89D43640FDE8300E" box="[492,562,1371,1393]" captionStart="Fig 34" captionStartId="8.[408,440,1155,1177]" captionTargetBox="[227,1120,478,1131]" captionTargetId="figure-107@8.[227,1122,478,1134]" captionTargetPageId="8" captionText="Fig 34. Type locality and habitat of Nigidius wushuae sp. n." figureDoi="http://doi.org/10.5281/zenodo.7166613" httpUri="https://zenodo.org/record/7166613/files/figure.png" pageId="7" pageNumber="26">Fig. 34</figureCitation>
) .............................................................
<emphasis id="49D9EA89FF9AFFFE8B8B3640FBBE300E" bold="true" box="[947,1124,1370,1392]" pageId="7" pageNumber="26">
<taxonomicName id="BCAD4D18FF9AFFFE8B8B3640FBF2300E" ID-CoL="8W97V" authority="Lin, 2022" authorityName="Lin" authorityYear="2022" box="[947,1064,1370,1392]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="wushuae" status="sp. nov.">
<emphasis id="49D9EA89FF9AFFFE8B8B3640FBF2300E" bold="true" box="[947,1064,1370,1392]" italics="true" pageId="7" pageNumber="26">N. wushuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52EA57F2FF9AFFFE8C163640FBBE300E" box="[1070,1124,1371,1392]" pageId="7" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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