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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1012.57047" ID-GBIF-Dataset="96b35ac5-ed73-465c-9e88-76f169714a83" ID-PMC="PMC7854560" ID-Pensoft-Pub="1313-2970-1012-21" ID-Pensoft-UUID="1AC67CC9000A5FCF86DECCE264464FD8" ID-PubMed="33584107" ID-ZooBank="4B5ACAB053224893BC53B4A48F8DC20C" ModsDocID="1313-2970-1012-21" checkinTime="1611749546682" checkinUser="pensoft" docAuthor="Rivera-Quiroz, Francisco Andres, Petcharad, Booppa &amp; Miller, Jeremy A." docDate="2021" docId="6EFF0126E3795E8B8FBC5DF030F5AF05" docLanguage="en" docName="ZooKeys 1012: 21-53" docOrigin="ZooKeys 1012" docPubDate="2021-01-26" docSource="http://dx.doi.org/10.3897/zookeys.1012.57047" docTitle="Crassignatha seeliam Rivera-Quiroz &amp; Petcharad &amp; Miller 2021, sp. nov." docType="treatment" docUuid="DA61A955-A1D4-4B7D-A7A0-89AD024460A3" docUuidSource="ZooBank" docVersion="4" id="1AC67CC9000A5FCF86DECCE264464FD8" lastPageNumber="21" masterDocId="1AC67CC9000A5FCF86DECCE264464FD8" masterDocTitle="First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995" masterLastPageNumber="53" masterPageNumber="21" pageNumber="21" updateTime="1668149733872" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rivera-Quiroz, Francisco Andres</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8748-4930</mods:nameIdentifier>
<mods:affiliation>Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands &amp; Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">andres.riveraquiroz@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petcharad, Booppa</mods:namePart>
<mods:affiliation>Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333 BE Leiden, the Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8918-9775</mods:nameIdentifier>
<mods:affiliation>Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-26</mods:number>
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<mods:detail type="volume">
<mods:number>1012</mods:number>
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<mods:start>21</mods:start>
<mods:end>53</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1012.57047</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1012.57047</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1012-21</mods:identifier>
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<treatment ID-GBIF-Taxon="177132026" LSID="urn:lsid:zoobank.org:act:DA61A955-A1D4-4B7D-A7A0-89AD024460A3" httpUri="http://treatment.plazi.org/id/6EFF0126E3795E8B8FBC5DF030F5AF05" lastPageNumber="21" pageId="0" pageNumber="21">
<subSubSection pageId="0" pageNumber="21" type="nomenclature">
<paragraph pageId="0" pageNumber="21">
<taxonomicName LSID="http://zoobank.org/DA61A955-A1D4-4B7D-A7A0-89AD024460A3" authority="Rivera-Quiroz &amp; Petcharad &amp; Miller, 2021" authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seeliam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seeliam" status="sp. nov.">Crassignatha seeliam</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="21">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. 3 D reconstruction of the male palp of Crassignatha with detail in the spermatic ducts: a-c C. seeliam sp. nov. d-f C. danaugirangensis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1012.57047.figure4" httpUri="https://binary.pensoft.net/fig/501612" pageId="0" pageNumber="21">Figures 4a-c</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D reconstruction of some diagnostic characters of Crassignatha males: a, c, e C. danaugirangensis b, d C. seeliam sp. nov. a chelicerae, arrow pointing at the bifurcated tooth b, c detail of the carapace; cephalothorax tubercles (in the squares), and pore bearing sulcus (arrows) d, e male leg II clasper f whole male specimen of C. danaugirangensis prepared for micro-CT inside a modified 10 µl pipette tip and a 0.5 ml Eppendorf tube filled with 70 % Et-OH. Scale bars: 0.06 mm (a); 0.1 mm (b-e)." figureDoi="10.3897/zookeys.1012.57047.figure5" httpUri="https://binary.pensoft.net/fig/501613" pageId="0" pageNumber="21">, 5b, d</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D reconstruction of the habitus of Crassignatha males: a, b C. seeliam sp. nov. c, d C. danaugirangensis. Right pedipalp was dissected previous to the scanning. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1012.57047.figure6" httpUri="https://binary.pensoft.net/fig/501614" pageId="0" pageNumber="21">, 6a, b</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">, 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="materials_examined">
<paragraph pageId="0" pageNumber="21">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Holotype</emphasis>
: Thailand • ♂: Chiang Mai, Doi Inthanon National Park. Montane evergreen forest;
<geoCoordinate degrees="18" direction="north" minutes="30.454" orientation="latitude" precision="1" value="18.507566">18°30.454'N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="30.584" orientation="longitude" precision="1" value="98.509735">98°30.584'E</geoCoordinate>
. 1605 m; July 21-24, 2018; Booppa Petcharad, Jeremy Miller, F. Andres Rivera-Quiroz leg.; direct hand coll.; RMNH.ARA.18444.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Paratypes</emphasis>
: Thailand • ♀ allotype; same data as holotype • 8 ♀; same data as holotype; RMNH.5106641• ♂ and ♀ Chiang Mai, Doi Suthep National Park. Montane evergreen forest with pine;
<geoCoordinate degrees="18" direction="north" minutes="48.502" orientation="latitude" precision="1" value="18.808367">18°48.502'N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="53.528" orientation="longitude" precision="1" value="98.892136">98°53.528'E</geoCoordinate>
. 1409 m; July 24-28, 2018; same collectors as holotype; pitfall traps. RMNH.ARA.18445.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="etymology">
<paragraph pageId="0" pageNumber="21">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="21">The species epithet is a derivation of the Thai seeliam (square), in reference to the shape of the abdomen in dorsal view.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="diagnosis">
<paragraph pageId="0" pageNumber="21">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="21">
Distinguished from other
<taxonomicName class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="genus">Crassignatha</taxonomicName>
species except
<taxonomicName authorityName="Lin &amp; Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha quadriventris" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="quadriventris">Crassignatha quadriventris</taxonomicName>
(Lin &amp; Li, 2009) by the semi-squared posterior of the abdomen in dorsal view (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">10b</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11b</figureCitation>
). Female can be separated from
<taxonomicName lsidName="C. quadriventris" pageId="0" pageNumber="21" rank="species" species="quadriventris">C. quadriventris</taxonomicName>
by the coiling of the copulatory ducts in the epigynum (compare Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11d</figureCitation>
and
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12c, d</figureCitation>
to
<bibRefCitation author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="47 - 68" refId="B23" refString="Lin, Y, Li, S, 2009. First described Patu spiders (Araneae, Symphytognathidae) from Asia. Zootaxa 2154: 47 - 68" title="First described Patu spiders (Araneae, Symphytognathidae) from Asia." volume="2154" year="2009">Lin and Li 2009</bibRefCitation>
: fig. 10). Male differs on the size of tegular sclerites and the cymbial tooth being short and stout instead of hook-shaped (compare Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">10c, d</figureCitation>
and
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12a, b</figureCitation>
to
<bibRefCitation author="Lin, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="47 - 68" refId="B23" refString="Lin, Y, Li, S, 2009. First described Patu spiders (Araneae, Symphytognathidae) from Asia. Zootaxa 2154: 47 - 68" title="First described Patu spiders (Araneae, Symphytognathidae) from Asia." volume="2154" year="2009">Lin and Li 2009</bibRefCitation>
: fig. 8).
</paragraph>
<caption doi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Figure 10.</emphasis>
<taxonomicName authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seeliam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seeliam">Crassignatha seeliam</taxonomicName>
sp. nov., male: Habitus:
<emphasis bold="true" pageId="0" pageNumber="21">a</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="21">b</emphasis>
dorsal view. Palp:
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
retrolateral view. Prosoma:
<emphasis bold="true" pageId="0" pageNumber="21">e</emphasis>
anterior view. Scale bars: 0.3 mm (
<emphasis bold="true" pageId="0" pageNumber="21">a, b</emphasis>
); 0.15 mm (
<emphasis bold="true" pageId="0" pageNumber="21">c-e</emphasis>
). Arrow pointing at the cymbial tooth.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="description">
<paragraph pageId="0" pageNumber="21">Description.</paragraph>
<paragraph pageId="0" pageNumber="21">
Carapace coloration orange-brown covered by small tubercles (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. 3 D reconstruction of the habitus of Crassignatha males: a, b C. seeliam sp. nov. c, d C. danaugirangensis. Right pedipalp was dissected previous to the scanning. Scale bars: 0.3 mm." figureDoi="10.3897/zookeys.1012.57047.figure6" httpUri="https://binary.pensoft.net/fig/501614" pageId="0" pageNumber="21">6a, b</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">10a, b</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11a, b</figureCitation>
). Legs same color, slightly darker on distal portion its segments. Male Tibia II with two spines (mating claspers) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. 3 D reconstruction of some diagnostic characters of Crassignatha males: a, c, e C. danaugirangensis b, d C. seeliam sp. nov. a chelicerae, arrow pointing at the bifurcated tooth b, c detail of the carapace; cephalothorax tubercles (in the squares), and pore bearing sulcus (arrows) d, e male leg II clasper f whole male specimen of C. danaugirangensis prepared for micro-CT inside a modified 10 µl pipette tip and a 0.5 ml Eppendorf tube filled with 70 % Et-OH. Scale bars: 0.06 mm (a); 0.1 mm (b-e)." figureDoi="10.3897/zookeys.1012.57047.figure5" httpUri="https://binary.pensoft.net/fig/501613" pageId="0" pageNumber="21">5d</figureCitation>
). Abdomen black with light red patches; squared posteriorly, with sparse sclerotized patches, some bearing long setae (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">10b</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11b</figureCitation>
). Male with posterior scutum wrapping the abdomen. Male palp: slightly less sclerotized than carapace. Semicircular from ventral view (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Crassignatha seeliam sp. nov., male: Habitus: a ventral view b dorsal view. Palp: c ventral view d retrolateral view. Prosoma: e anterior view. Scale bars: 0.3 mm (a, b); 0.15 mm (c-e). Arrow pointing at the cymbial tooth." figureDoi="10.3897/zookeys.1012.57047.figure10" httpUri="https://binary.pensoft.net/fig/501618" pageId="0" pageNumber="21">10c</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12a</figureCitation>
). Cymbium with distal tooth. Median apophysis as big as Ct (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12a</figureCitation>
). Embolus filiform, exposed when palp is expanded (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12c</figureCitation>
). Spermatic duct very long and coiling 2
<normalizedToken originalValue="×">x</normalizedToken>
inside the bulb (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. 3 D reconstruction of the male palp of Crassignatha with detail in the spermatic ducts: a-c C. seeliam sp. nov. d-f C. danaugirangensis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1012.57047.figure4" httpUri="https://binary.pensoft.net/fig/501612" pageId="0" pageNumber="21">4b, c</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Figure 11.</emphasis>
<taxonomicName authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seeliam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seeliam">Crassignatha seeliam</taxonomicName>
sp. nov. female: Habitus:
<emphasis bold="true" pageId="0" pageNumber="21">a</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="21">b</emphasis>
dorsal view. Epigynum:
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
dorsal view, cleared. Scale bars: 0.4 mm (
<emphasis bold="true" pageId="0" pageNumber="21">a, b</emphasis>
); 0.15 mm (
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
); 0.07 mm (
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Figure 12.</emphasis>
<taxonomicName authorityName="Rivera-Quiroz &amp; Petcharad &amp; Miller" authorityYear="2021" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha seeliam" order="Araneae" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="seeliam">Crassignatha seeliam</taxonomicName>
sp. nov., genitalia. Palp:
<emphasis bold="true" pageId="0" pageNumber="21">a</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="21">b</emphasis>
dorsal view. Epigynum, cleared:
<emphasis bold="true" pageId="0" pageNumber="21">c</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="21">d</emphasis>
ventral view. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="21">a, b</emphasis>
); 0.07 mm (
<emphasis bold="true" pageId="0" pageNumber="21">c, d</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Vulva</emphasis>
: Epigynum with wide scape directed ventrally, heavily sclerotized at the tip (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11c</figureCitation>
). Copulatory opening at the tip of scape (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11d</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12c, d</figureCitation>
). Spermathecae spherical, slightly more sclerotized than epigynum, separated by ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
their diameter (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11d</figureCitation>
). Copulatory ducts very long, coiling over themselves before connecting to S. Fertilization ducts as long as S width, projecting dorsally (figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Crassignatha seeliam sp. nov. female: Habitus: a ventral view b dorsal view. Epigynum: c ventral view d dorsal view, cleared. Scale bars: 0.4 mm (a, b); 0.15 mm (c); 0.07 mm (d)." figureDoi="10.3897/zookeys.1012.57047.figure11" httpUri="https://binary.pensoft.net/fig/501619" pageId="0" pageNumber="21">11d</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Crassignatha seeliam sp. nov., genitalia. Palp: a ventral view b dorsal view. Epigynum, cleared: c dorsal view d ventral view. Scale bars: 0.1 mm (a, b); 0.07 mm (c, d)." figureDoi="10.3897/zookeys.1012.57047.figure12" httpUri="https://binary.pensoft.net/fig/501620" pageId="0" pageNumber="21">12c</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Male</emphasis>
: Total length 0.68; carapace 0.36 long, 0.30 wide; clypeus 0.13; Chelicera 0.1 long, 0.07 wide; Leg I: femur 0.28, patella 0.12, tibia 0.37, metatarsus 0.17, tarsus 0.22; leg formula I-II-IV-III; abdomen 0.42 long, 0.38 wide.
</paragraph>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">Female</emphasis>
: Total length 0.69, carapace 0.44 long, 0.39 wide; clypeus 0.12; Chelicera 0.15 long, 0.1 wide; Leg I: femur 0.42, patella 0.15, tibia 0.53, metatarsus 0.22, tarsus 0.27; leg formula I-II-IV-III abdomen 0.44 long, 0.43 wide.
</paragraph>
</subSubSection>
</treatment>
</document>