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<mods:title id="C52A0FD9383E6D00CFF1BC238D5618F5">Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan</mods:title>
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<mods:namePart id="15518C8D213B6B6D5B3DE6D512F2012C">Shimizu, So</mods:namePart>
<mods:affiliation id="9D0FB6937A07E67E00EC5DA539613FD1">Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan &amp; DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan &amp; Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="52353BEAB83A10D57115715A5ABA2B7D">Broad, Gavin R.</mods:namePart>
<mods:affiliation id="F5D8801D73628D858A3B550848B0AEA9">Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="482BA6CCA6F50E9C5BBEF2497796A34B">Maeto, Kaoru</mods:namePart>
<mods:affiliation id="ED882969E20B6D4DFCE59D1031DFE09B">Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan</mods:affiliation>
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<treatment id="B08ADC10635551BD90A3FDAB909B387C" ID-GBIF-Taxon="170296369" LSID="urn:lsid:plazi:treatment:B08ADC10635551BD90A3FDAB909B387C" httpUri="http://treatment.plazi.org/id/B08ADC10635551BD90A3FDAB909B387C" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Insecta" scope_family="Ichneumonidae" scope_order="Hymenoptera" scope_subFamily="Ophioninae" scope_superFamily="Ichneumonoidea">
<subSubSection id="67A0AC632B68E9C1DCAA0543D2E77A30" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="C954C0C5E517DF7B6E4E1107F27DC56D" pageId="0" pageNumber="1">
<taxonomicName id="416742974E66EC205B2F925933AAB6D8" ID-CoL="6FBCG" LSID="B08ADC10-6355-51BD-90A3-FDAB909B387C" authority="(Taschenberg, 1875)" baseAuthorityName="Taschenberg" baseAuthorityYear="1875" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus aciculatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aciculatus">Enicospilus aciculatus (Taschenberg, 1875)</taxonomicName>
<figureCitation id="480E7BDC3DD4F9D90540153330DECBB0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Enicospilus aciculatus (Taschenberg, 1875) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure10" httpUri="https://binary.pensoft.net/fig/473776" pageId="0" pageNumber="1">Figure 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F5527AAA9A9E1E54F5F143D18DC0DAA0" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="9ADC28E60C9C73CCA27C57503C0F7843" pageId="0" pageNumber="1">
<taxonomicName id="5F9202E5D0E221A0DB2C9ADAEF3A9AE4" class="Insecta" family="Ichneumonidae" genus="Ophion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophion aciculatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aciculatus">Ophion aciculatus</taxonomicName>
Taschenberg, 1875: 434; LCT ♀ from Java, designated by
<bibRefCitation id="9D5E99A239E80025E697D00807231E7F" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="1" pagination="1 - 522" refId="B178" refString="Townes, HK, Townes, M, Gupta, VK, 1961. A catalogue and reclassification of the Indo-Australian Ichneumonidae. Memoirs of the American Entomological Institute 1: 1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. (1961</bibRefCitation>
: 269), FZLU, not examined.
</paragraph>
<paragraph id="107868C90F750643A00EAB449C00E94C" pageId="0" pageNumber="1">
<taxonomicName id="F828DFD2791445B48120D3016F44E52C" authorityName="Brues" authorityYear="1918" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus malaitensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="malaitensis">Enicospilus malaitensis</taxonomicName>
Brues, 1918: 117; HT ♂ from Solomon Island, MCZ, not examined; synonymised by
<bibRefCitation id="385FC673140A22250B8745ACE9550814" author="Gauld, ID" journalOrPublisher="CAB: Slough / Commonwealth Institute of Entomology, London" pageId="0" pageNumber="1" refId="B41" refString="Gauld, ID, Mitchell, PA, 1981. The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae. CAB: Slough / Commonwealth Institute of Entomology, London" title="The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae." year="1981">Gauld and Mitchell (1981</bibRefCitation>
: 394).
</paragraph>
<paragraph id="0A09C7949F976314D7BB756679CEE895" pageId="0" pageNumber="1">
<taxonomicName id="27CCDA9E6A779EBCC9C5CD553F070046" class="Insecta" family="Ichneumonidae" genus="Henicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicospilus okinawensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="okinawensis">Henicospilus okinawensis</taxonomicName>
Matsumura and Uchida, 1926: 71; LCT ♀ from
<normalizedToken id="827A07346FA0AA350A8BCAAD23A57AD6" originalValue="Ryûkyûs">Ryukyus</normalizedToken>
, designated by
<bibRefCitation id="5AF386B476203CBA4A54789457D4839A" author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="1" pagination="1 - 522" refId="B178" refString="Townes, HK, Townes, M, Gupta, VK, 1961. A catalogue and reclassification of the Indo-Australian Ichneumonidae. Memoirs of the American Entomological Institute 1: 1 - 522" title="A catalogue and reclassification of the Indo-Australian Ichneumonidae." volume="1" year="1961">Townes et al. (1961</bibRefCitation>
: 285), SEHU, examined; synonymised by
<bibRefCitation id="EC014736BAE0B23EFB78F7C01F023B4C" author="Gauld, ID" journalOrPublisher="CAB: Slough / Commonwealth Institute of Entomology, London" pageId="0" pageNumber="1" refId="B41" refString="Gauld, ID, Mitchell, PA, 1981. The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae. CAB: Slough / Commonwealth Institute of Entomology, London" title="The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae." year="1981">Gauld and Mitchell (1981</bibRefCitation>
: 394).
</paragraph>
<paragraph id="3C834A19C9715AD5EC846EF7D6094653" pageId="0" pageNumber="1">
<taxonomicName id="5AC1B725D1E40AFF967F296C8ECC4F48" class="Insecta" family="Ichneumonidae" genus="Henicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicospilus neddenveli" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neddenveli">Henicospilus neddenveli</taxonomicName>
Cheesman, 1936: 185; HT ♂ from Vanuatu, NHMUK, examined; synonymised by
<bibRefCitation id="9C5C56E0665AE3B18CF40FEA6CECC51A" author="Gauld, ID" journalOrPublisher="CAB: Slough / Commonwealth Institute of Entomology, London" pageId="0" pageNumber="1" refId="B41" refString="Gauld, ID, Mitchell, PA, 1981. The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae. CAB: Slough / Commonwealth Institute of Entomology, London" title="The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae." year="1981">Gauld and Mitchell (1981</bibRefCitation>
: 394).
</paragraph>
<paragraph id="2F37102DFB6E61DECCBCEA49C1617952" pageId="0" pageNumber="1">
<taxonomicName id="9DFD680C57306F0C44CBD98E202AE2B3" authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus crucis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="crucis">Enicospilus crucis</taxonomicName>
Chiu, 1954: 70; HT ♀ from Taiwan, TARI, examined; synonymised by
<bibRefCitation id="C53801FA5632CA72C2DF05734B5C83DD" author="Gauld, ID" journalOrPublisher="CAB: Slough / Commonwealth Institute of Entomology, London" pageId="0" pageNumber="1" refId="B41" refString="Gauld, ID, Mitchell, PA, 1981. The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae. CAB: Slough / Commonwealth Institute of Entomology, London" title="The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae." year="1981">Gauld and Mitchell (1981</bibRefCitation>
: 394).
</paragraph>
</subSubSection>
<subSubSection id="9DAA6B4279F31B464DAB9C77F135BE7E" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="55E60DF5FD6C3FAC3E17BAB9BDD88EFB" pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph id="E6416C2655803BA5FDF4A41B7E39F4DF" pageId="0" pageNumber="1">Total of 54 specimens (29♀♀24♂♂ and 1 unsexed): India (17♀♀5♂♂), Japan (8♀♀17♂♂ and 1 unsexed), Solomon Islands (2♀♀), Sri Lanka (1♀1♂), Taiwan (1♀), Vanuatu (1♂).</paragraph>
<paragraph id="CE84058A1E11BA9E484164507CFB4BD8" pageId="0" pageNumber="1">
Type series: LCT ♀ of
<taxonomicName id="ADF8881CB74A09CC1F09DC383FFBF8C2" genus="Henicospilus" lsidName="Henicospilus okinawensis" pageId="0" pageNumber="1" rank="species" species="okinawensis">
<emphasis id="66904B54D46B040640E3022579A628B1" italics="true" pageId="0" pageNumber="1">Henicospilus okinawensis</emphasis>
</taxonomicName>
Matsumura &amp; Uchida, 1926, Okinawa,
<normalizedToken id="B00F621E382954406ECADDCA3626D8D6" originalValue="Ryûkyûs">Ryukyus</normalizedToken>
, JAPAN, 1925, S. Sakaguchi leg. (SEHU); HT ♀ of
<taxonomicName id="B76B1C8D955662CA24AFC3C010118595" authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus crucis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="crucis">
<emphasis id="9BA222EB01F16FD53DCDEF574DF2761F" italics="true" pageId="0" pageNumber="1">Enicospilus crucis</emphasis>
</taxonomicName>
Chiu, 1954,
<normalizedToken id="170CA1E81693BC4585BB2F3EC882271E" originalValue="Jûjiro">Jujiro</normalizedToken>
, TAIWAN, 26.IV.1931, T. Shiraki leg. (TARI); HT ♂ of
<taxonomicName id="FED718DFD46FBF432A2FDC071CB6D636" genus="Henicospilus" lsidName="Henicospilus neddenveli" pageId="0" pageNumber="1" rank="species" species="neddenveli">
<emphasis id="C2F80034AF2A4C7BC0B1B35FF8AFDC87" italics="true" pageId="0" pageNumber="1">Henicospilus neddenveli</emphasis>
</taxonomicName>
Cheesman, 1936, Malekula, New Hebrides [VANUATU], III.1930, L.E. Cheesman leg. (NHMUK, Type 3b.1242).
</paragraph>
</subSubSection>
<subSubSection id="8AD846254F17A70BEE74FB80A75419CC" pageId="0" pageNumber="1" type="distribution">
<paragraph id="0D4E722C7361D5D071BBE3708D5B647F" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="87BA70701E9F7C6ADD24E49987352C81" pageId="0" pageNumber="1">
Australasian, Eastern Palaearctic, Oceanic and Oriental regions (
<bibRefCitation id="FD69E2AF6B317C09B09867DC87F026B4" author="Yu, DSK" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" publicationUrl="www.taxapad.com" refId="B199" refString="Yu, DSK, van Achterberg, C, Horstmann, K, 2016. Taxapad 2016, Ichneumonoidea 2015 Database on flash-drive. Nepean, Ontario, Canada. www.taxapad.com" title="Taxapad 2016, Ichneumonoidea 2015 Database on flash-drive. Nepean, Ontario, Canada." url="www.taxapad.com" year="2016">Yu et al. 2016</bibRefCitation>
).
</paragraph>
<paragraph id="5BBFD6C1653894F8FD3C1F54BBEF9C25" pageId="0" pageNumber="1">
JAPAN: [
<normalizedToken id="6F4BA6C0B9E7157AE7647B101BC18996" originalValue="Kyûshû">Kyushu</normalizedToken>
] Fukuoka* and Kagoshima (
<bibRefCitation id="71A33591E645D854B87AA2BA113FBFD0" author="Momoi, S" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="327 - 399" refId="B97" refString="Momoi, S, 1970. Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12: 327 - 399" title="Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago." volume="12" year="1970">Momoi 1970</bibRefCitation>
; present study); [
<normalizedToken id="10D06102B733AEE6BF07E92DDEFC0E8A" originalValue="Ryûkyûs">Ryukyus</normalizedToken>
] Kagoshima (
<bibRefCitation id="2AC739BA7E9E95188B42A1058D958D6F" author="Uchida, T" journalOrPublisher="Insecta matsumurana" pageId="0" pageNumber="1" pagination="82 - 100" refId="B184" refString="Uchida, T, 1956. Die Ichneumoniden aus der Amami Inselgruppe. Insecta matsumurana 19: 82 - 100" title="Die Ichneumoniden aus der Amami Inselgruppe." volume="19" year="1956">Uchida 1956</bibRefCitation>
;
<bibRefCitation id="8845A986A9B2570265A11D2EAF4F5F0F" author="Momoi, S" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="327 - 399" refId="B97" refString="Momoi, S, 1970. Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12: 327 - 399" title="Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago." volume="12" year="1970">Momoi 1970</bibRefCitation>
; present study) and Okinawa (
<bibRefCitation id="B76D8655E49F0E263A1B462CC3E1AB4D" author="Matsumura, S" journalOrPublisher="Insecta matsumurana" pageId="0" pageNumber="1" pagination="63 - 77" refId="B94" refString="Matsumura, S, Uchida, T, 1926. Die Hymenopteran-Fauna von den Riukiu-Inseln. Insecta matsumurana 1: 63 - 77" title="Die Hymenopteran-Fauna von den Riukiu-Inseln." volume="1" year="1926">Matsumura and Uchida 1926</bibRefCitation>
;
<bibRefCitation id="36B561AF7ED5BE5187739C96336AEA84" author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido University" pageId="0" pageNumber="1" pagination="177 - 297" refId="B180" refString="Uchida, T, 1928. Zweiter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido University 21: 177 - 297" title="Zweiter Beitrag zur Ichneumoniden-Fauna Japans." volume="21" year="1928">Uchida 1928</bibRefCitation>
; present study). *New record.
<taxonomicName id="1019F3284175A0AF475384A290024AD1" baseAuthorityName="Taschenberg" baseAuthorityYear="1875" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus aciculatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aciculatus">
<emphasis id="30348E7B263B2571C2CE42853E727542" italics="true" pageId="0" pageNumber="1">Enicospilus aciculatus</emphasis>
</taxonomicName>
is a predominantly (sub-)tropical species and rare in the cooler zone.
</paragraph>
</subSubSection>
<subSubSection id="5948D1418F2B34158C4D93C6C2B7F46C" pageId="0" pageNumber="1" type="bionomics">
<paragraph id="FD1071020F41DC8BB4AC6CDF5622D335" pageId="0" pageNumber="1">Bionomics.</paragraph>
<paragraph id="69A201AEE717A180520C18084745CCC4" pageId="0" pageNumber="1">Although this is one of the most common species in the Oriental region, there are no host records.</paragraph>
</subSubSection>
<subSubSection id="3B899E47213123C26E0CD978E4820E13" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="7702FE6C745791A44C3CA9FE194FA843" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="1FFFF92ED245BFD855B17EF8F9643669" pageId="0" pageNumber="1">
This species is morphologically relatively close to
<taxonomicName id="1BE4C731B3CFF931C1538E6E0E35618A" lsidName="E. laqueatus" pageId="0" pageNumber="1" rank="species" species="laqueatus">
<emphasis id="CF4D3CE3FC21F04E02E04EB2E89D2EEA" italics="true" pageId="0" pageNumber="1">E. laqueatus</emphasis>
</taxonomicName>
and
<taxonomicName id="85BF1BBB417169B560F69019D6B2DB80" lsidName="E. yonezawanus" pageId="0" pageNumber="1" rank="species" species="yonezawanus">
<emphasis id="37185B72EF9B7EC8E3A6D971798F16F0" italics="true" pageId="0" pageNumber="1">E. yonezawanus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="C58F6BBF4C48F6B5E6115881AE92B9E1" lsidName="E. aciculatus" pageId="0" pageNumber="1" rank="species" species="aciculatus">
<emphasis id="39A5A90A54076B198008C9E7A3FC991E" italics="true" pageId="0" pageNumber="1">E. aciculatus</emphasis>
</taxonomicName>
is rather easily distinguished from all other species of Japanese
<taxonomicName id="5A8F2EAA7F480A209C1CA67343159340" authorityName="Stephens" authorityYear="1835" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="940F14311F1B97477D9BAEBA4E75AF6C" italics="true" pageId="0" pageNumber="1">Enicospilus</emphasis>
</taxonomicName>
by the following combination of character states: mandible with diagonal setose groove, and upper mandibular tooth more than 2.0
<normalizedToken id="5C483D67194C8345DF07E8E91E72BDD2" originalValue="×">x</normalizedToken>
as long as lower (Fig.
<figureCitation id="269D32AA097C71788E6447CB988B99D5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Enicospilus aciculatus (Taschenberg, 1875) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure10" httpUri="https://binary.pensoft.net/fig/473776" pageId="0" pageNumber="1">10B, D</figureCitation>
); central sclerite of fore wing fenestra moderately sized, ill-defined, very weakly pigmented, and positioned in anterodistal corner of fenestra (Fig.
<figureCitation id="21FCA0869BD31A4CEC87A7F355F369B6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Enicospilus aciculatus (Taschenberg, 1875) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure10" httpUri="https://binary.pensoft.net/fig/473776" pageId="0" pageNumber="1">10F</figureCitation>
); and proximal angle of proximal sclerite of fore wing fenestra ca. 45° (Fig.
<figureCitation id="68103BFB097BB7C27CD3FC1FF51A9B48" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Enicospilus aciculatus (Taschenberg, 1875) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure10" httpUri="https://binary.pensoft.net/fig/473776" pageId="0" pageNumber="1">10F</figureCitation>
). This species is morphologically stable but sometimes exhibits colour variation (i.e., usually the metasoma is entirely orange brown, but rarely the posterior segments are black).
</paragraph>
<caption id="838E0BAF379BF723FF535B7DDE687FE0" doi="10.3897/zookeys.990.55542.figure10" httpUri="https://binary.pensoft.net/fig/473776" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph id="1178BC1018517866C16477A3ACF0BDD2" pageId="0" pageNumber="1">
<emphasis id="A4A422087196A21E35315464AC921EEF" bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName id="72EFCACFF1BAB80D48A5DD0DEE2684E6" baseAuthorityName="Taschenberg" baseAuthorityYear="1875" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus aciculatus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aciculatus">
<emphasis id="93BCECC8AE628F95692416BD55180830" italics="true" pageId="0" pageNumber="1">Enicospilus aciculatus</emphasis>
</taxonomicName>
(Taschenberg, 1875) ♀ from Japan
<emphasis id="D0E4263D0B4A01141F14DF3024171927" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus
<emphasis id="FBCA532C6E3744B944704B6C4B8B9F30" bold="true" pageId="0" pageNumber="1">B</emphasis>
head, frontal view
<emphasis id="24DB0699C84A691EF837F6A5336138D5" bold="true" pageId="0" pageNumber="1">C</emphasis>
head, dorsal view
<emphasis id="4E16C8E4515E4E868B803B66F4A6678E" bold="true" pageId="0" pageNumber="1">D</emphasis>
head, lateral view
<emphasis id="98CCCD2091A7CF8B6CAF53A759AF3DCF" bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma, lateral view
<emphasis id="4247F75CAC0B5B844EDB301FC04D6E24" bold="true" pageId="0" pageNumber="1">F</emphasis>
central part of fore wing.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>