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<document ID-CLB-Dataset="7765" ID-DOI="http://dx.doi.org/10.3897/zookeys.1023.61478" ID-GBIF-Dataset="94123c77-d2c7-4aa7-9bdb-8367f7851b82" ID-PMC="PMC7973068" ID-Pensoft-Pub="1313-2970-1023-81" ID-Pensoft-UUID="8E4EE48C84345AC48B877220002D94B6" ID-PubMed="33776516" ID-ZooBank="F14F12AC47824643929A49941E24BB92" ModsDocID="1313-2970-1023-81" checkinTime="1615560751875" checkinUser="pensoft" docAuthor="Hajek, Jiri, Shaverdo, Helena, Hendrich, Lars &amp; Balke, Michael" docDate="2021" docId="8AF0CA7D9BD15945BF51A353D15580A2" docLanguage="en" docName="ZooKeys 1023: 81-118" docOrigin="ZooKeys 1023" docPubDate="2021-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1023.61478" docTitle="Copelatus portior Guignot 1956" docType="treatment" docVersion="7" id="8E4EE48C84345AC48B877220002D94B6" lastPageNumber="81" masterDocId="8E4EE48C84345AC48B877220002D94B6" masterDocTitle="A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="118" masterPageNumber="81" pageNumber="81" updateTime="1732937923291" updateUser="ExternalLinkService">
<mods:mods id="A5480DA9079568CF0CD075204E2A7448" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A050368B86190E8F621B1DB0B2D12971">
<mods:title id="F54C8011FAB98B3853D84DA7BC4C72D4">A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="3CCCB386B3AE93277D326C385201FCCE" type="personal">
<mods:role id="0D85EE2EC36CD79485DCB22AFE61D60D">
<mods:roleTerm id="7B26EF7D5F2F468DFB09B587E3E88143">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="11EFE68BC7222BAA422F9A3C81749A77">Hajek, Jiri</mods:namePart>
<mods:nameIdentifier id="3BCB0CD7BAE6445254DED3E25C6D3DE1" type="ORCID">https://orcid.org/0000-0001-5779-1542</mods:nameIdentifier>
<mods:affiliation id="0AF95A893EA8A2A5B87307063C796A97">Department of Entomology, National Museum, Cirkusova 1740, CZ- 193 00, Prague 9 - Horni Pocernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="1D9726C61E0FCA5C0F734293A1B2EF64" type="email">jiri.hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="AC8A94CB2BAACB2DB5D87C4483990C8D" type="personal">
<mods:role id="DF9944DCB4AA3C8D149FF53B3447D971">
<mods:roleTerm id="2FBFDE400211E6D4BC4EEE237887A92F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F804ADCE09A2962AE15CDC7F82E91DD4">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="545BC2D7A2340BB788A5250484EE55AC">Naturhistorisches Museum Wien, Burgring 7, 1010, Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name id="155EAA59A84642018D483923F6DEA2C5" type="personal">
<mods:role id="DEE926D780DD416FACAED8258BF488B3">
<mods:roleTerm id="7EDD0946521D925EF0941E7B05DF512D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="289A9DD61A21F24122440027BFD26418">Hendrich, Lars</mods:namePart>
<mods:affiliation id="88F9F59B860251B76D70C551D699B3F7">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany</mods:affiliation>
</mods:name>
<mods:name id="95CBB4590C0D498FFC4D22834879FC07" type="personal">
<mods:role id="9331EFEA55A0252721E3901D44BF4159">
<mods:roleTerm id="08491D14762858989AB196DC285404B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4409FBC0C63AB9A7871CE1B151B3D72A">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="A245F2D47AC51A6B064307300C178F8D" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="F835524439CBF50DC33427414EC2333C">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6EFB8005FF2FB031599DABFDF2A7C0F9">text</mods:typeOfResource>
<mods:relatedItem id="99FC2AF19D79A3D03267B6CC4DF270B6" type="host">
<mods:titleInfo id="C60F3CE007A56EA95BF20535DA13A791">
<mods:title id="55CF6B715A5B5C88ECBE62F1A668D370">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="7E9EF5424C2995F9B2D4112DDEA09DE5">
<mods:date id="E1144FA9459592434D2FE52299616A5D">2021</mods:date>
<mods:detail id="4700471CE327CCB34FEF425978EDDFBE" type="pubDate">
<mods:number id="48225ADB0CC40E8BBF2F7F4064B851ED">2021-03-10</mods:number>
</mods:detail>
<mods:detail id="B644EA911EEDB72AFFB0B4FDC82A3BEB" type="volume">
<mods:number id="787422F6DBC54C3C57BB7593500D9F7D">1023</mods:number>
</mods:detail>
<mods:extent id="EC5A74B3FAF644063DED65BD155E0B8E" unit="page">
<mods:start id="FC12C70438C96C1BFED2423081743EC3">81</mods:start>
<mods:end id="D5F385E4B60F8719B5786A0CFCC7D7EA">118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9B2821BC53BEBC2E2E67A2BCA6BE4A43">
<mods:url id="691594DD76483B830FA86B87EA4D57BF">http://dx.doi.org/10.3897/zookeys.1023.61478</mods:url>
</mods:location>
<mods:classification id="D8D5094146150ADF8BC1E194C056A7C7">journal article</mods:classification>
<mods:identifier id="6F65C75A13BD460CF8D5E72BD606AD1C" type="DOI">http://dx.doi.org/10.3897/zookeys.1023.61478</mods:identifier>
<mods:identifier id="80302CF5FFCC3ED9E052FB5C9AAFF1DA" type="Pensoft-Pub">1313-2970-1023-81</mods:identifier>
<mods:identifier id="B32389636738D0C87ECDF1796DCF0CC2" type="ZooBank">F14F12AC47824643929A49941E24BB92</mods:identifier>
<mods:identifier id="B67A6280DEDBECB3370681F0C349F748" type="Pensoft-UUID">8E4EE48C84345AC48B877220002D94B6</mods:identifier>
</mods:mods>
<treatment id="8AF0CA7D9BD15945BF51A353D15580A2" ID-GBIF-Taxon="180331288" LSID="urn:lsid:plazi:treatment:8AF0CA7D9BD15945BF51A353D15580A2" httpUri="http://treatment.plazi.org/id/8AF0CA7D9BD15945BF51A353D15580A2" lastPageNumber="81" pageId="0" pageNumber="81" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="4D83C4D3635783AED1841283346FF28B" pageId="0" pageNumber="81" type="nomenclature">
<paragraph id="C3DD4D47A6A8C733379653F03D13ED6A" pageId="0" pageNumber="81">
<taxonomicName id="F60A88BDFBF0A29A7C599198FC69D9E6" ID-CoL="5ZWX2" LSID="8AF0CA7D-9BD1-5945-BF51-A353D15580A2" authority="Guignot, 1956" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus portior" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="portior">Copelatus portior Guignot, 1956</taxonomicName>
<figureCitation id="0D901271C7CF6E654CFC23A515B94230" captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Habitus of Copelatus 5 C. portior Guignot, 1956 (male, Guadalcanal; TL: 5.2 mm) 6 C. portior (striolate female, Papua; TL: 4.7 mm)." figureDoi="10.3897/zookeys.1023.61478.figures5-6" httpUri="https://binary.pensoft.net/fig/519050" pageId="0" pageNumber="81">Figures 5-6</figureCitation>
<figureCitation id="0284DF13B2C3BD4D70016DC4C81EC317" captionStart="Figure 19" captionStartId="F11" captionText="Figure 19. Male genitalia of Copelatus portior Guignot, 1956 (Australia) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure19" httpUri="https://binary.pensoft.net/fig/519058" pageId="0" pageNumber="81">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ED13A45B5285F509200ADC5987DF0B5F" pageId="0" pageNumber="81" type="reference_group">
<paragraph id="F47E631A0BC3FB603DC0566B968A0039" pageId="0" pageNumber="81">
<taxonomicName id="8ACE1AA161DFC08DFA3A16DD54258269" authorityName="Guignot" authorityYear="1956" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus portior" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="portior">Copelatus portior</taxonomicName>
Guignot, 1956: 53 (
<typeStatus id="710539BFA8C0A12A002BDD21F9C7DBA1">type</typeStatus>
locality: &quot;New Hebrides [
<collectingCountry id="EF35D2D94AD1614F92DB93E7DDA5AFD9" name="Vanuatu">Vanuatu</collectingCountry>
], Malekula Island&quot;).
</paragraph>
<paragraph id="5F6BA44E13B5F64152710F59AF1720F7" pageId="0" pageNumber="81">
<taxonomicName id="19EEFDF93DC8495A887DFA2F1D6C9F93" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus divisus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="divisus">Copelatus divisus</taxonomicName>
Watts, 1978: 122 (&quot;Seleo, Berlinhafen&quot; [
<collectingCountry id="B0F46BCBBB908F6A204A24381CD5EFDA" name="Papua New Guinea">Papua New Guinea</collectingCountry>
,
<collectingRegion id="123E1074D6039CC6F215FC5116A34625" country="Papua New Guinea" name="West Sepik">Sandaun Province</collectingRegion>
, Seleo Island); synonymy by
<bibRefCitation id="6E5AB95DD36DEFAE4DC0745FAEE550BC" DOI="https://doi.org/10.3897/zookeys.889.39090" author="Hendrich, L" journalOrPublisher="ZooKeys" pageId="0" pageNumber="81" pagination="81 - 152" refId="B13" refString="Hendrich, L, Shaverdo, H, Hajek, J, Balke, M, 2019. Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 889: 81 - 152, DOI: https://doi.org/10.3897/zookeys.889.39090" title="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.889.39090" volume="889" year="2019">Hendrich et al. (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="577C198D8FAE1562583D6F7E0406CE7F" pageId="0" pageNumber="81" type="materials_examined">
<paragraph id="2E3A1A460B39E70F1177B638CDE0838A" pageId="0" pageNumber="81">Material examined.</paragraph>
<paragraph id="DE13E794063792AF1BE20D52340F624B" pageId="0" pageNumber="81">
<emphasis id="6D163DF6100B23DA071D773EB69ABFD3" bold="true" pageId="0" pageNumber="81">
<taxonomicName id="4E7E5922ADC95D150DD6BAAFB320E605" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">
<emphasis id="09B8CB025AF16C94A06B6095C7015445" italics="true" pageId="0" pageNumber="81">Guadalcanal</emphasis>
</taxonomicName>
:
</emphasis>
<materialsCitation id="4661D0233726C3892F3F94C2C5196CC7" ID-GBIF-Occurrence="3063630311" collectingDate="1953-09-08" collectingDateMax="1953-09-12" collectingDateMin="1953-09-08" location="Honiara" specimenCount="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="159E702B7B24BD4A0AC72ED0CA9C219A" type="male">1 ♂</specimenCount>
,
<collectingRegion id="181D8C1BC3B6974480B0F4EFBA00BCC7" country="Solomon Islands" name="Honiara">Honiara</collectingRegion>
, M.V. light,
<date id="5188C867207F491FE55C97E7FD8F88F4" value="1953-09-08" valueMax="1953-09-12" valueMin="1953-09-08">
<collectingDate id="095901EF8C1D7B1456BCAB374A25C175" value="1953-09-08" valueMax="1953-09-12" valueMin="1953-09-08">8.-12.ix.1953</collectingDate>
</date>
</materialsCitation>
, J.D. Bradley leg.;
<materialsCitation id="C79A7A839BE97F8920EC03D22FDB4C9C" ID-GBIF-Occurrence="3063630312" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. J. M. Greenslade" location="Kukum" specimenCount="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="2990FE14742F1E00BEEDDB892AE07936" type="male">1 ♂</specimenCount>
,
<collectingRegion id="2B990169F22B08592B27DBFB3EFA5327" country="Solomon Islands" name="Honiara">Honiara</collectingRegion>
,
<location id="5A841D2EDFD57053A0739C1AA59964F0" LSID="urn:lsid:plazi:treatment:8AF0CA7D9BD15945BF51A353D15580A2:5A841D2EDFD57053A0739C1AA59964F0" name="Kukum" stateProvince="Honiara">Kukum</location>
, 1962,
<collectorName id="33B93B24377ADD5BB3B5D4087A1B1869">P.J.M. Greenslade</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="23437AD6D4D4D3CD6F1679EB6192FCBC" ID-GBIF-Occurrence="3063630344" collectingDate="1962-05-19" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. J. M. Greenslade" location="Kukum" specimenCount="1" specimenCount-female="1" stateProvince="Honiara">
<specimenCount id="51B41FEA416F24AA247A60AEEB0430BC" type="female">1 ♀</specimenCount>
, same data, but
<date id="A2163E0880FEBDF84C192E7669FAB2F3" value="1962-05-19">
<collectingDate id="156D2035F3C6227903C2C1220560F732" value="1962-05-19">19.v.1962</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="1AB56FFDFEAFDBB7AF583831C05A74E0" ID-GBIF-Occurrence="3063630345" collectingDate="1963-05-20" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. J. M. Greenslade" location="Kukum" specimenCount="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="FFEB3B24470149490282E3D9C8A90DEB" type="male">1 ♂</specimenCount>
, same data, but
<date id="E956AB24C029E487FBC3CF418902D3D5" value="1963-05-20">
<collectingDate id="EF39654E3C2A7C99D95FFDE4C27163C5" value="1963-05-20">20.v.1963</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="955C51DB28BDBB21AE10DCEBE164A8B5" ID-GBIF-Occurrence="3063630309" collectingDate="1965-03-02" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. J. M. Greenslade" location="Kukum" specimenCount="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="4CA9539DEF114B99985B4BE05DF690E1" type="male">1♂</specimenCount>
, same data, but,
<date id="28079ACFDB9119943598386C7E7E64F8" value="1965-03-02">
<collectingDate id="A0AB8B171385BFDB5C5213B067BC0762" value="1965-03-02">2.iii.1965</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8F8DEA62FA3B096B4619469913AC475D" ID-GBIF-Occurrence="3063630325" collectingDate="1966-05-12" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMUK" collectorName="P. J. M. Greenslade" location="Kukum" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="338A0682CFF4FD4550F655FC95819D66" type="male">1 ♂</specimenCount>
,
<specimenCount id="2E2DE1C31D005AF1735E6CF386E01EF0" type="female">1 ♀</specimenCount>
, same data, but
<date id="3BEA4D6E1BF33F7069F2E10DF2E8E55F" value="1966-05-12">
<collectingDate id="03B28707D3A998F53CB5861DB1D79B5B" value="1966-05-12">12.v.1966</collectingDate>
</date>
(all
<collectionCode id="CE8AB2C54474D7A5039CD76D11FBA97B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DF88834B02D5BB26E58056C7297087C3" ID-GBIF-Occurrence="3063630322" collectingDate="2013-11-23" collectingDateMax="2013-12-08" collectingDateMin="2013-11-23" elevation="280" latitude="-9.466666" location="Barana" longLatPrecision="129" longitude="159.97333" municipality="Mt. Austine" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Honiara">
<specimenCount id="54691A226FCB817A0F20137FBA3068E9" type="male">1 ♂</specimenCount>
,
<specimenCount id="323D123342BB9719C085E76390190302" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="FC503F63D42EB91DE000462C706E8FDF">Mt. Austine</collectingMunicipality>
,
<location id="821F99E650E4229A1D5CE5BD60491E97" LSID="urn:lsid:plazi:treatment:8AF0CA7D9BD15945BF51A353D15580A2:821F99E650E4229A1D5CE5BD60491E97" latitude="-9.466666" longLatPrecision="129" longitude="159.97333" municipality="Mt. Austine" name="Barana" stateProvince="Honiara">Barana</location>
vill. env.,
<geoCoordinate id="A451674EB02C1E75BB0111131C094657" degrees="09" direction="south" minutes="28.0" orientation="latitude" precision="92" value="-9.466666">09°28.0'S</geoCoordinate>
,
<geoCoordinate id="C496AD09C0ECB97D569875F90B309AE3" degrees="159" direction="east" minutes="58.4" orientation="longitude" precision="92" value="159.97333">159°58.4'E</geoCoordinate>
,
<quantity id="CC6E7BA1BEEDB149DE8E72F86D47A880" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<elevation id="209F48E702DDECFC44D534C054797554" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</elevation>
</quantity>
,
<date id="B68B566A84198784E57FD59D21CBE90B" value="2013-11-23" valueMax="2013-12-08" valueMin="2013-11-23">
<collectingDate id="A63FF018000D56E7D74B427D2B4D6766" value="2013-11-23" valueMax="2013-12-08" valueMin="2013-11-23">23.xi.-8.xii.2013</collectingDate>
</date>
</materialsCitation>
,
<collectorName id="33A657429DD6C8B5E68DC01EDB6F362A">
J.
<normalizedToken id="1A09880B6ECDAC6F3AE007387C8425BD" originalValue="Hájek">Hajek</normalizedToken>
</collectorName>
leg. (
<collectionCode id="92EF1B61267CA3AD0E56369EB506F745" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
,
<collectionCode id="6C92B53503410493435F6CD418390572">ZSMG</collectionCode>
).
</paragraph>
<paragraph id="56119E11D1EFE3D049043B52ABE6D77A" pageId="0" pageNumber="81">
<emphasis id="B9DCF1FA1DA3AAB8080BB88A92B4EEF7" bold="true" pageId="0" pageNumber="81">
<emphasis id="67E8C4EB4F33BD94098890FA61A15F1D" italics="true" pageId="0" pageNumber="81">Ontong Java</emphasis>
:
</emphasis>
<materialsCitation id="83ECAF78979841A9D7143A031481990C" ID-GBIF-Occurrence="3063630348" collectingDate="1955-01-31" collectionCode="NHMUK" collectorName="Keila &amp; E. S. Brown" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="750D5CD3E90C523EADECBEE3F3110783" type="male">1 ♂</specimenCount>
,
<specimenCount id="6B10E5FA6BBE624B4CAC268985FA37EA" type="female">1 ♀</specimenCount>
,
<collectorName id="0DE62E07BCDD1C315C8C42108E60E551">Keila</collectorName>
,
<date id="85C3C88C38386010C5F7723A36351DFC" value="1955-01-31">
<collectingDate id="8D390EE5F7C37F2F59FD8894253875F1" value="1955-01-31">31.i.1955</collectingDate>
</date>
,
<collectorName id="AE50174559EECF8A80D5D4F5CEC17047">E.S. Brown</collectorName>
leg. (
<collectionCode id="614BDB50B417F46F2D45AC899CA7D167" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="271EFFE2B330DE00ECEAD33346FD8A95" ID-GBIF-Occurrence="3063630340" collectingDate="1955-01-27" collectionCode="NHMUK" collectorName="Leuaniua &amp; E. S. Brown" specimenCount="1" specimenCount-female="1">
<specimenCount id="10FBB4E3760EA093114791753B587250" type="female">1 ♀</specimenCount>
,
<collectorName id="3DBC5A76DA432AC2C3DE3C8FBD104288">Leuaniua</collectorName>
,
<date id="D049B6DD72BCAF0EDA9D25FB715AA26F" value="1955-01-27">
<collectingDate id="06B2C9EC5B9DE4EE2631C3BACF4FB108" value="1955-01-27">27.i.1955</collectingDate>
</date>
,
<collectorName id="145F0EA898682E0FF2513B9C747F5A95">E.S. Brown</collectorName>
leg. (
<collectionCode id="4A58159C17613B47009C51E8F62F37E9" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9E3E832470A467BCF134DFAFA3B33B6" pageId="0" pageNumber="81" type="diagnosis">
<paragraph id="978E0716BEC6438376A08ACF7195D66F" pageId="0" pageNumber="81">Diagnosis.</paragraph>
<paragraph id="CAD358ECF17205B4E5131BF75BE411DF" pageId="0" pageNumber="81">
For complete description, see
<bibRefCitation id="75FD82545CEA31A069E77048C4271289" DOI="https://doi.org/10.3897/zookeys.889.39090" author="Hendrich, L" journalOrPublisher="ZooKeys" pageId="0" pageNumber="81" pagination="81 - 152" refId="B13" refString="Hendrich, L, Shaverdo, H, Hajek, J, Balke, M, 2019. Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 889: 81 - 152, DOI: https://doi.org/10.3897/zookeys.889.39090" title="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.889.39090" volume="889" year="2019">Hendrich et al. (2019)</bibRefCitation>
. Medium sized (TL: 5.0-
<quantity id="3EC0F21E7A313F1A01662A8DC2AD4E36" metricMagnitude="-3" metricUnit="m" metricValue="5.7" unit="mm" value="5.7">5.7 mm</quantity>
), oblong-oval species. Species variable in elytral colouration: elytra from almost uniformly dark brown to black with only base and lateral sides yellowish, to almost yellowish-orange coloured with dark stripes along elytral striae (Fig.
<figureCitation id="D7E0A7C576C07BA3F4DA048CCC42E79B" captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Habitus of Copelatus 5 C. portior Guignot, 1956 (male, Guadalcanal; TL: 5.2 mm) 6 C. portior (striolate female, Papua; TL: 4.7 mm)." figureDoi="10.3897/zookeys.1023.61478.figures5-6" httpUri="https://binary.pensoft.net/fig/519050" pageId="0" pageNumber="81">5</figureCitation>
). Elytron with well-impressed six discal striae and a submarginal stria. Female dimorphic; striolate form with dorsal surface almost black, matt and with coarse microreticulation and numerous strioles on elytra and pronotum (Fig.
<figureCitation id="56462A977516C34047046D8B5C2CC267" captionStart="Figures 5, 6" captionStartId="F3" captionText="Figures 5, 6. Habitus of Copelatus 5 C. portior Guignot, 1956 (male, Guadalcanal; TL: 5.2 mm) 6 C. portior (striolate female, Papua; TL: 4.7 mm)." figureDoi="10.3897/zookeys.1023.61478.figures5-6" httpUri="https://binary.pensoft.net/fig/519050" pageId="0" pageNumber="81">6</figureCitation>
). Median lobe in lateral view sickle-shaped; broad and subparallel in basal two thirds, thin and regularly curved in apical third; a distinct hammer-like process present in two thirds on ventral side (Fig.
<figureCitation id="5AF6A92D1A3A821EA0E44F9BB91758C4" captionStart="Figure 19" captionStartId="F11" captionText="Figure 19. Male genitalia of Copelatus portior Guignot, 1956 (Australia) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure19" httpUri="https://binary.pensoft.net/fig/519058" pageId="0" pageNumber="81">19</figureCitation>
). Parameres broad,
<normalizedToken id="C448D0B24D9A716220BE34B1D989C1A2" originalValue="“D”">&quot;D&quot;</normalizedToken>
-shaped; apical lobes moderately long, club-shaped.
</paragraph>
<caption id="6CE78856FE0AC2A220AC6136BEA9E9F7" doi="10.3897/zookeys.1023.61478.figures5-6" httpUri="https://binary.pensoft.net/fig/519050" pageId="0" pageNumber="81" start="Figures 5, 6" startId="F3">
<paragraph id="EC19A1F5C7FDBC8C4F38963657A75BD4" pageId="0" pageNumber="81">
<emphasis id="DFD231CFCCC21A593EB96897D9E73FD3" bold="true" pageId="0" pageNumber="81">Figures 5, 6.</emphasis>
Habitus of
<taxonomicName id="C10D9A28DE4AA5542CC9D65E001023AC" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="379B40D2B45CDC341C430AFFAEF1B1B0" italics="true" pageId="0" pageNumber="81">Copelatus</emphasis>
</taxonomicName>
<emphasis id="00BAAC37A5B5E57DE79717BE41010B89" bold="true" pageId="0" pageNumber="81">5</emphasis>
<taxonomicName id="DBB7E52952F832FD44E3CE06D9A263A3" lsidName="C. portior" pageId="0" pageNumber="81" rank="species" species="portior">
<emphasis id="D7910996819837423B931DF87477812B" italics="true" pageId="0" pageNumber="81">C. portior</emphasis>
</taxonomicName>
Guignot, 1956 (male,
<taxonomicName id="D1CEEEAC8C0E5173D6A839140430F9B7" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
; TL:
<quantity id="DC1B663D04838D1E28987EB12995594D" metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
)
<emphasis id="9BCFCEEEBC1F60B7AE59F98107644A85" bold="true" pageId="0" pageNumber="81">6</emphasis>
<taxonomicName id="9524FF82723EACAFF99832FEE64AB847" lsidName="C. portior" pageId="0" pageNumber="81" rank="species" species="portior">
<emphasis id="F52FDBFB58BC79F39D429F21CE8EDD98" italics="true" pageId="0" pageNumber="81">C. portior</emphasis>
</taxonomicName>
(striolate female, Papua; TL:
<quantity id="F2F99AEC293717FCC8C80F87477AF173" metricMagnitude="-3" metricUnit="m" metricValue="4.7" unit="mm" value="4.7">4.7 mm</quantity>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2F8BD35F3A942FCD19806811346BECAE" pageId="0" pageNumber="81" type="distribution">
<paragraph id="E0F2BF8CDC112C77F6914AEFB3B725B7" pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph id="0024DC982F44CDE9748157FE3AA67F88" pageId="0" pageNumber="81">
The species is originally described from northern
<collectingCountry id="457EAFA767EEF487F5AE64F2A1CCE9D9" name="Papua New Guinea">Papua New Guinea</collectingCountry>
and
<collectingCountry id="B6A50F2E644D05AB66DBD2572C88E982" name="Vanuatu">Vanuatu</collectingCountry>
. It is widely distributed in the Australasian Region: from Lesser Sunda Islands, through New
<collectingCountry id="1909DCC4AF2313DDCEA1034AD7FED04C" name="Guinea-Bissau">Guinea</collectingCountry>
, northern
<collectingCountry id="DD86079ED4D9CA112A49483052F02E1A" name="Australia">Australia</collectingCountry>
, and
<collectingCountry id="4E2E422615B92A63FDB83642DCBCE111" name="Solomon Islands">Solomon Islands</collectingCountry>
to
<collectingCountry id="B24810D952F615E1AD3E574C928E5C2B" name="Vanuatu">Vanuatu</collectingCountry>
. First record from the
<collectingCountry id="EDCFC3A6EF50E43F020894A8F86326E2" name="Solomon Islands">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0A2774AEBEBC356350818D7CA220A5C9" pageId="0" pageNumber="81" type="habitat">
<paragraph id="28CAE01973BB50F1B9F031D22D40A900" pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph id="7ED4231498D3A8BAC8A6E812D2521D4C" pageId="0" pageNumber="81">
Two specimens from Barana were recently collected in a streamlet flowing through secondary forest and gardens near the village; both of them were found in calm water with decaying leaves on the bottom (Fig.
<figureCitation id="5B44E867EF75515308BE6D5E0D22B7E7" captionStart="Figures 26–28" captionStartId="F16" captionText="Figures 26 - 28. Habitats of Copelatus in Guadalcanal 26 temporary forest stream near Barana village 27 shaded pool on the same stream 28 small temporary puddles in a gorge along the road from Barana to Lungga river." figureDoi="10.3897/zookeys.1023.61478.figures26-28" httpUri="https://binary.pensoft.net/fig/519063" pageId="0" pageNumber="81">26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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