treatments-xml/data/75/68/87/756887E70176670BFF03186BFE088A1C.xml

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<mods:title id="A9D624A055818689894B8036D5681CA4">New species and records of the Rhamphomyia subgenus Holoclera (Diptera, Empididae) from the Palaearctic Region</mods:title>
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<mods:namePart id="CFCC4518BE0F59418F2D7441C631AF28">Shamshev, Igor V.</mods:namePart>
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<mods:affiliation id="46378803A5844D700040B3A69CA0269B">All-Russian Institute of Plant Protection, St. Petersburg - Pushkin, Russia</mods:affiliation>
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<mods:namePart id="A9F60335674CD44DC6C3FF1A5DE495EB">Barták, Miroslav</mods:namePart>
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<paragraph id="FD7E36F101766709FF03186BFD178E50" blockId="5.[151,694,582,645]" box="[151,694,582,609]" pageId="5" pageNumber="517">
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<emphasis id="CFB5EAE301766709FF03186BFD178E50" bold="true" box="[151,694,582,609]" pageId="5" pageNumber="517">
<taxonomicName id="3AC14D7201766709FF03186BFDF08E50" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[151,593,582,609]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="517" phylum="Arthropoda" rank="species" species="richterae" status="sp. nov." subGenus="Holoclera">
<emphasis id="CFB5EAE301766709FF03186BFEE88E50" bold="true" box="[151,329,582,609]" italics="true" pageId="5" pageNumber="517">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE301766709FECE186BFE728E50" bold="true" box="[346,467,582,609]" italics="true" pageId="5" pageNumber="517">Holoclera</emphasis>
) 
<emphasis id="CFB5EAE301766709FE70186BFDF08E50" bold="true" box="[484,593,582,609]" italics="true" pageId="5" pageNumber="517">richterae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D486579801766709FDCC186AFD178E50" box="[600,694,583,609]" pageId="5" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B5DB657A01766709FF031846FF4A8EB4" box="[151,235,619,645]" pageId="5" pageNumber="517" type="description">
<paragraph id="FD7E36F101766709FF031846FF4A8EB4" blockId="5.[151,694,582,645]" box="[151,235,619,645]" pageId="5" pageNumber="517">
(
<figureCitation id="65FA2A7401766709FF0B1846FF418EB4" box="[159,224,619,645]" captionStart="FIGURE 1" captionStartId="6.[152,255,1763,1787]" captionTargetBox="[202,1382,309,1739]" captionTargetId="figure-91@6.[196,1392,309,1739]" captionTargetPageId="6" captionText="FIGURE 1. Rhamphomyia (Holoclera) richterae sp. nov., male. A. Habitus, lateral view. B. Hypopygium, lateral view. C. Outline of hypopygium, lateral view (setae omitted). D. Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="5" pageNumber="517">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A01766709FF03189EFB288F6C" pageId="5" pageNumber="517" type="materials_examined">
<paragraph id="FD7E36F101766709FF03189EFB288F6C" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="517">
<emphasis id="CFB5EAE301766709FF03189EFE5B8EFD" bold="true" box="[151,506,690,717]" pageId="5" pageNumber="517">
Type material. 
<typeStatus id="227A885301766709FEC0189FFE5B8EFD" box="[340,506,690,716]" pageId="5" pageNumber="517" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="EBC7FD7801766709FD91189EFDBD8EFC" box="[517,540,691,717]" count="1" pageId="5" pageNumber="517" type="male">♂</specimenCount>
, labelled: [
<emphasis id="CFB5EAE301766709FD34189FFBB48EFC" bold="true" box="[672,1045,690,717]" pageId="5" pageNumber="517">
<collectingCountry id="85D6766101766709FD34189FFCAD8EFD" box="[672,780,690,716]" name="Russia" pageId="5" pageNumber="517">RUSSIA</collectingCountry>
. Krasnodarskiy Terr.
</emphasis>
] Lasarevskoe [
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], 
<taxonomicName id="3AC14D7201766709FF0318FAFF5E8EC0" box="[151,255,727,753]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="517" phylum="Tracheophyta" rank="genus">
<emphasis id="CFB5EAE301766709FF0318FAFF5E8EC0" box="[151,255,727,753]" italics="true" pageId="5" pageNumber="517">Carpinus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="3AC14D7201766709FE9318FAFED08EC0" box="[263,369,727,753]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="517" phylum="Tracheophyta" rank="genus">
<emphasis id="CFB5EAE301766709FE9318FAFED08EC0" box="[263,369,727,753]" italics="true" pageId="5" pageNumber="517">Castanea</emphasis>
</taxonomicName>
forest, 
<date id="897F103101766709FE7A18FAFDC68EC0" box="[494,615,727,753]" pageId="5" pageNumber="517" value="1984-09-11">11.ix.1984</date>
, Richter (
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, 
<collectionCode id="9BD0AE3401766709FCC218FAFC228EC0" box="[854,899,727,753]" pageId="5" pageNumber="517">INS</collectionCode>
_DIP_0000849). 
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<typeStatus id="227A885301766709FBCE18FBFAA78EC1" box="[1114,1286,726,752]" pageId="5" pageNumber="517" type="paratype">PARATYPES</typeStatus>
. 
<collectingCountry id="85D6766101766709FABD18FBFA348EC1" box="[1321,1429,726,752]" name="Russia" pageId="5" pageNumber="517">RUSSIA</collectingCountry>
. Krasnodarskiy Terr.:
</emphasis>
same locality as 
<typeStatus id="227A885301766709FDF818D6FD708F24" box="[620,721,763,789]" pageId="5" pageNumber="517" type="holotype">holotype</typeStatus>
, 
<date id="897F103101766709FD7518D6FCFA8F24" box="[737,859,763,789]" pageId="5" pageNumber="517" value="1982-09-23">23.ix.1982</date>
, Gorodkov (
<specimenCount id="EBC7FD7801766709FC6518D6FB808F24" box="[1009,1057,763,789]" count="2" pageId="5" pageNumber="517" type="female">2 ♀</specimenCount>
, 
<collectionCode id="9BD0AE3401766709FBA018D6FBD38F24" box="[1076,1138,763,789]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="517">ZISP</collectionCode>
). Sochi, mountain Akhun [
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<geoCoordinate id="98F5503601766709FE93190DFEC78F0A" box="[263,358,800,827]" degrees="39" direction="east" minutes="50" orientation="longitude" pageId="5" pageNumber="517" precision="925" value="39.833332">39°50′E</geoCoordinate>
], 24,27. 
<date id="897F103101766709FE561932FDB08F08" box="[450,529,799,825]" pageId="5" pageNumber="517" value="2006-10">x.2006</date>
, N. Vikhrev (
<specimenCount id="EBC7FD7801766709FD381932FD788F08" box="[684,729,799,825]" count="2" pageId="5" pageNumber="517" type="female">2 ♀</specimenCount>
, 
<collectionCode id="9BD0AE3401766709FD7D1932FCE48F08" box="[745,837,799,825]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="517" type="Museum">ZMMU</collectionCode>
). 
<emphasis id="CFB5EAE301766709FCCD1933FC1C8F09" bold="true" box="[857,957,798,824]" pageId="5" pageNumber="517">
<collectingRegion id="3F05F81301766709FCCD1933FC168F09" box="[857,951,798,824]" country="Russia" name="Adygeya" pageId="5" pageNumber="517">Adygea</collectingRegion>
:
</emphasis>
Dakhovskaya env., E slope of Azish-Tau, 
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<emphasis id="CFB5EAE301766709FF03196EFF7C8F6D" box="[151,221,835,860]" italics="true" pageId="5" pageNumber="517">Fagus</emphasis>
</taxonomicName>
forest, 
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<geoCoordinate id="98F5503601766709FE18196EFE708F6C" box="[396,465,835,861]" degrees="40.1" direction="east" orientation="longitude" pageId="5" pageNumber="517" precision="5555" value="40.1">40.1E</geoCoordinate>
, ~ 
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, 
<date id="897F103101766709FDD4196EFD558F6C" box="[576,756,835,861]" pageId="5" pageNumber="517" value="2009-08-26" valueMax="2009-08-29" valueMin="2009-08-26">26–29.viii.2009</date>
, coll. K. Tomkovich (
<specimenCount id="EBC7FD7801766709FC7E196EFBB48F6C" box="[1002,1045,835,861]" count="1" pageId="5" pageNumber="517" type="male">1 ♂</specimenCount>
, 
<collectionCode id="9BD0AE3401766709FBB5196EFBDC8F6C" box="[1057,1149,835,861]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="5" pageNumber="517" type="Museum">ZMMU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A01766709FF53194BFF758FF8" pageId="5" pageNumber="517" type="diagnosis">
<paragraph id="FD7E36F101766709FF53194BFF758FF8" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="517">
<emphasis id="CFB5EAE301766709FF53194BFEE38FB1" bold="true" box="[199,322,870,896]" pageId="5" pageNumber="517">Diagnosis.</emphasis>
A species of the 
<emphasis id="CFB5EAE301766709FD97194AFDB58FB1" box="[515,532,871,896]" italics="true" pageId="5" pageNumber="517">R</emphasis>
. 
<taxonomicName id="3AC14D7201766709FDB7194AFD318FB0" box="[547,656,871,897]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="517" phylum="Arthropoda" rank="species" species="flava" subGenus="Holoclera">
(
<emphasis id="CFB5EAE301766709FDB8194AFDE18FB1" box="[556,576,871,896]" italics="true" pageId="5" pageNumber="517">H</emphasis>
.) 
<emphasis id="CFB5EAE301766709FDCC194AFD318FB0" box="[600,656,871,897]" italics="true" pageId="5" pageNumber="517">flava</emphasis>
</taxonomicName>
group; acrostichal setae absent; thorax and abdomen entirely yellow, male mesoscutum and abdominal tergites with some brownish tinge. Male: halter with brown knob and yellow stem.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A0176670BFF5319FFFB268FB0" lastPageId="7" lastPageNumber="519" pageId="5" pageNumber="517" type="description">
<paragraph id="FD7E36F101766709FF5319FFFD5C8964" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="517">
<emphasis id="CFB5EAE301766709FF5319FFFE3C8FDC" bold="true" box="[199,413,978,1005]" pageId="5" pageNumber="517">Description. Male</emphasis>
(
<figureCitation id="65FA2A7401766709FE3919FEFDA78FDC" box="[429,518,979,1005]" captionStart="FIGURE 1" captionStartId="6.[152,255,1763,1787]" captionTargetBox="[202,1382,309,1739]" captionTargetId="figure-91@6.[196,1392,309,1739]" captionTargetPageId="6" captionText="FIGURE 1. Rhamphomyia (Holoclera) richterae sp. nov., male. A. Habitus, lateral view. B. Hypopygium, lateral view. C. Outline of hypopygium, lateral view (setae omitted). D. Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="5" pageNumber="517">Fig. 1A</figureCitation>
). Body length 
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, wing length 
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(
<typeStatus id="227A885301766709FB9019FEFBCC8FDC" box="[1028,1133,979,1005]" pageId="5" pageNumber="517" type="holotype">holotype</typeStatus>
). Head with greyish, dense pruinescence on ocellar triangle, occiput and postgena; frons above antennae brownish, face and clypeus brownish yellow; black setose (except noted). Eyes holoptic, with upper ommatidia enlarged. Frons represented by very small, subtriangular space just below ocellar triangle and slightly larger space just above antennae. Face moderately broad, almost parallel-sided, bare. Ocellar triangle with 2 long, thin anterior setae and two pairs of minute setulae posteriorly. Occiput with moderately long strong postoculars and almost regular rows of similar setae laterally; postgena with some short, black and longer pale hair-like setae. Antenna with scape and pedicel yellowish brown, postpedicel (except yellowish brown extreme base) and stylus dark brown; scape short, slightly longer than pedicel, both with short setulae; postpedicel elongate-conical, with narrow base, nearly 3X as long as basal width; stylus short, 2.8X shorter than postpedicel. Proboscis short, with labrum only slightly longer than head height (1.3X in 
<typeStatus id="227A885301766709FF031F16FF5F8964" box="[151,254,1339,1365]" pageId="5" pageNumber="517" type="holotype">holotype</typeStatus>
); palpus yellow, with black, scattered setulae.
</paragraph>
<paragraph id="FD7E36F101766709FF531F72FE1C8AA8" blockId="5.[151,1437,690,2013]" pageId="5" pageNumber="517">Thorax almost uniformly yellow, only mesoscutum somewhat darker, with some brownish tinge; faintly pruinose, mesoscutum rather matt; black setose. Proepisternum with 7–10 minute setulae on lower part, 1 similar setula on upper part. Prosternum bare. Antepronotum with 8–9 short, strong setae on each side. Postpronotal lobe with 1 strong, long and 4–5 short, fine setae. Mesonotal setae well-differentiated; acrostichals absent; dorsocentrals uniserial, 9 setae per row, 7 presutural setae subequally moderately long, 2 postsutural setae longer (prescutellar longest); 0 presutural intra-alar, 1 long presutural supra-alar, 3 notopleurals (middle seta longest; additionally, with 2 setulae anteriorly), 1 postsutural supra-alar (with 1–2 setulae anteriorly), 1 long and 1 minute postalars, 4 scutellars (apical setae longer and stronger). Laterotergite with 4–5 long fine setae and numerous minute setulae. Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="FD7E36F10176670AFF531C8EFB998D2C" blockId="5.[151,1437,690,2013]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="518" pageId="5" pageNumber="517">Legs long, slender, podomeres unmodified; extensively yellow and subshiny (including coxae); trochanters with brownish subapical spot, tibiae brownish yellow on extreme apex, basitarsus of all legs brownish on apex, fore and mid tarsomeres 2–5 almost uniformly brownish (tarsomere 2 somewhat paler near base), hind tarsomere 2 brownish at apex and hind tarsomeres 3–5 brownish. Coxae and trochanters with simple setae. Fore femur with simple setulae anteriorly; bare ventrally, with mostly minute anteroventral and posteroventral setae (somewhat longer near apex) and short, strong subapical setae. Fore tibia with 3 moderately long dorsal setae (besides circlet of subapicals). Mid femur covered with mostly uniform minute setulae ventrally; short, strong anteroventral and posteroventral setae on about apical 1/4; subapical circlet of setae of different robustness (1 anterior seta strongest). Mid tibia with 2 moderately long anterodorsal, 3–4 similar posterodorsal, 1 short anteroventral and 2 short posteroventral setae (position somewhat variable even on right and left legs). Hind femur slightly narrower on about basal half, with similar setation to mid femur. Hind tibia slightly evenly thickened on subapical part; with 4 anterodorsal and 4 posterodorsal setae; 1 short seta in posteroapical comb. Basitarsus of all legs with short spine-like setae ventrally; tarsomeres 3–4 with longer setae of subapical circlet than those on tarsomeres 1–2.</paragraph>
<caption id="A9BE66790175670AFF0C1CCEFC708B73" ID-DOI="http://doi.org/10.5281/zenodo.13849162" ID-Zenodo-Dep="13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="6" pageNumber="518" startId="6.[152,255,1763,1787]" targetBox="[202,1382,309,1739]" targetPageId="6" targetType="figure">
<paragraph id="FD7E36F10175670AFF0C1CCEFC708B73" blockId="6.[151,1436,1763,1859]" pageId="6" pageNumber="518">
<emphasis id="CFB5EAE30175670AFF0C1CCEFEBA8ACA" bold="true" box="[152,283,1763,1787]" pageId="6" pageNumber="518">FIGURE 1.</emphasis>
<taxonomicName id="3AC14D720175670AFEB71CCEFD3A8ACA" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[291,667,1763,1787]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="518" phylum="Arthropoda" rank="species" species="richterae" status="sp. nov." subGenus="Holoclera">
<emphasis id="CFB5EAE30175670AFEB71CCEFE148ACA" box="[291,437,1763,1787]" italics="true" pageId="6" pageNumber="518">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE30175670AFE521CCEFD8C8ACA" box="[454,557,1763,1787]" italics="true" pageId="6" pageNumber="518">Holoclera</emphasis>
) 
<emphasis id="CFB5EAE30175670AFDAA1CCEFD3A8ACA" box="[574,667,1763,1787]" italics="true" pageId="6" pageNumber="518">richterae</emphasis>
</taxonomicName>
<emphasis id="CFB5EAE30175670AFD371CCEFD578ACA" bold="true" box="[675,758,1763,1787]" pageId="6" pageNumber="518">
<taxonomicNameLabel id="D48657980175670AFD371CCEFD578ACA" box="[675,758,1763,1787]" pageId="6" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 
<emphasis id="CFB5EAE30175670AFCD11CCEFCFC8ACA" bold="true" box="[837,861,1763,1787]" pageId="6" pageNumber="518">A.</emphasis>
Habitus, lateral view. 
<emphasis id="CFB5EAE30175670AFBDD1CCEFBC08ACA" bold="true" box="[1097,1121,1763,1787]" pageId="6" pageNumber="518">B.</emphasis>
Hypopygium, lateral view. 
<emphasis id="CFB5EAE30175670AFA101CCEFA3D8ACA" bold="true" box="[1412,1436,1763,1787]" pageId="6" pageNumber="518">C.</emphasis>
Outline of hypopygium, lateral view (setae omitted). 
<emphasis id="CFB5EAE30175670AFD221D2AFD6F8B2E" bold="true" box="[694,718,1799,1823]" pageId="6" pageNumber="518">D.</emphasis>
Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="FD7E36F10175670AFF531DB8FAF98BC9" blockId="6.[151,1437,1941,2040]" pageId="6" pageNumber="518">Wing membrane faintly brownish; veins brownish, well sclerotised; CuA+CuP complete but slightly weakened on about middle; cell dm short, with elongate apex. Pterostigma absent. Basal costal seta absent. Anal lobe well-developed; axillary incision acute. Squama yellow, brown fringed. Halter with brown knob and yellow stem.</paragraph>
<paragraph id="FD7E36F10174670BFF531ABAFA318CE4" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">Abdomen extensively yellow, faintly pruinose, black setose; tergites rather brownish yellow dorsally, with long posteromarginal setae. Pregenital segments unmodified; segment 8 with fused tergite and sternite; tergite 8 entire.</paragraph>
<paragraph id="FD7E36F10174670BFF531AF2FB3A8EFC" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">
Hypopygium (
<figureCitation id="65FA2A740174670BFEFB1AF2FE118CC8" box="[367,432,223,249]" captionStart="FIGURE 1" captionStartId="6.[152,255,1763,1787]" captionTargetBox="[202,1382,309,1739]" captionTargetId="figure-91@6.[196,1392,309,1739]" captionTargetPageId="6" captionText="FIGURE 1. Rhamphomyia (Holoclera) richterae sp. nov., male. A. Habitus, lateral view. B. Hypopygium, lateral view. C. Outline of hypopygium, lateral view (setae omitted). D. Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="7" pageNumber="519">Fig. 1</figureCitation>
B-D) small, black setose; cercus and most part of epandrium brownish; basal part of epandrium, hypandrium and visible part of phallus brownish yellow. Epandrial lamellae narrowly connected dorsally (
<figureCitation id="65FA2A740174670BFF0B1B0AFEBC8D70" box="[159,285,295,321]" captionStart="FIGURE 1" captionStartId="6.[152,255,1763,1787]" captionTargetBox="[202,1382,309,1739]" captionTargetId="figure-91@6.[196,1392,309,1739]" captionTargetPageId="6" captionText="FIGURE 1. Rhamphomyia (Holoclera) richterae sp. nov., male. A. Habitus, lateral view. B. Hypopygium, lateral view. C. Outline of hypopygium, lateral view (setae omitted). D. Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="7" pageNumber="519">Fig. 1B, C</figureCitation>
); epandrial lamella subtriangular (lateral view); covered with moderately long to short, fine setae on apical part and along upper margin. Hypandrium separated from epandrium; subtriangular in ventral view, elongate, with rounded apex, well-sclerotised, bearing 2 spinule-like subapical setae; gonocoxal apodeme inconspicuous. Hypoproct weakly sclerotised, bare. Phallus short, almost entirely hidden, slightly extending beyond cerci, gently curved; almost uniformly thick, slightly thicker closer to apex, in lateral view dorsal margin of apex slightly longer than ventral margin; with pair of rod-like, stronger sclerotised spaces on about apical 1/3 forming partly ventral margin of phallotrema; phallotrema large. Ejaculatory apodeme moderately large, without lateral wings, only extending beyond basal curvature of phallus. Cerci separated from epandrium and from each other; cercus broadly deeply concave on upper part posteriorly forming two lobes; upper lobe invisible in lateral view, in dorsal view small (
<figureCitation id="65FA2A740174670BFF711846FE9E8EB4" box="[229,319,619,645]" captionStart="FIGURE 1" captionStartId="6.[152,255,1763,1787]" captionTargetBox="[202,1382,309,1739]" captionTargetId="figure-91@6.[196,1392,309,1739]" captionTargetPageId="6" captionText="FIGURE 1. Rhamphomyia (Holoclera) richterae sp. nov., male. A. Habitus, lateral view. B. Hypopygium, lateral view. C. Outline of hypopygium, lateral view (setae omitted). D. Cerci, dorsal view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849162" httpUri="https://zenodo.org/record/13849162/files/figure.png" pageId="7" pageNumber="519">Fig. 1D</figureCitation>
); lower lobe in lateral view subtriangular and short, apex reaching nearly middle of upper margin of epandrial lamella, produced ventrally forming additional internal subglobular cercal lobe; visible part of cercus covered with simple setulae of different lengths, internal lobe with similar marginal setulae.
</paragraph>
<paragraph id="FD7E36F10174670BFF5318FAFB268FB0" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">
<emphasis id="CFB5EAE30174670BFF5318FAFE848EC0" bold="true" box="[199,293,727,753]" pageId="7" pageNumber="519">Female.</emphasis>
Similar to male, except as follows: Dichoptic; frons below ocellar triangle nearly as broad as anterior ocellus, broadened toward antennae on about lower half, bearing minute marginal setulae; wing hyaline; halter yellow; mesoscutum yellow, with shorter presutural dorsocentral setae; abdomen with shorter setae; abdominal tergites 1–7 with indistinct, brownish yellow, small, subtriangular spots anteriorly; tergite 8, syntergite 9+10 and cerci brownish, rather densely greyish pruinose; cercus long, slender, with minute setulae.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A0174670BFF5319A6FC688FF8" pageId="7" pageNumber="519" type="etymology">
<paragraph id="FD7E36F10174670BFF5319A6FC688FF8" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">
<emphasis id="CFB5EAE30174670BFF5319A6FEEF8F94" bold="true" box="[199,334,907,933]" pageId="7" pageNumber="519">Etymology.</emphasis>
The species epithet, 
<taxonomicName id="3AC14D720174670BFDA219A6FD0B8F94" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[566,682,907,933]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="richterae">
“ 
<emphasis id="CFB5EAE30174670BFDD719A6FD0B8F94" box="[579,682,907,933]" italics="true" pageId="7" pageNumber="519">richterae</emphasis>
</taxonomicName>
”, is a Latin genitive matronym to memory of Russian dipterist Vera Andreevna Richter (1936–2015), who collected part of the 
<typeStatus id="227A88530174670BFCBA1982FCFF8FF8" box="[814,862,943,969]" pageId="7" pageNumber="519">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A0174670BFF5319FEFDDB89AC" pageId="7" pageNumber="519" type="discussion">
<paragraph id="FD7E36F10174670BFF5319FEFDDB89AC" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">
<emphasis id="CFB5EAE30174670BFF5319FEFE9A8FDC" bold="true" box="[199,315,979,1005]" pageId="7" pageNumber="519">Remarks.</emphasis>
The new species belongs to the 
<emphasis id="CFB5EAE30174670BFD2B19FEFD718FDD" box="[703,720,979,1004]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFD7619FEFCF38FDC" box="[738,850,979,1005]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="flava" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFD7F19FEFD5E8FDD" box="[747,767,979,1004]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFC8E19FEFCF38FDC" box="[794,850,979,1005]" italics="true" pageId="7" pageNumber="519">flava</emphasis>
</taxonomicName>
group and is similar to 
<emphasis id="CFB5EAE30174670BFBE619FEFB228FDD" box="[1138,1155,979,1004]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFB0119FEFA3D8FDC" authority="(Fallen)" baseAuthorityName="Fallen" box="[1173,1436,979,1005]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="variabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFB0A19FEFB138FDD" box="[1182,1202,979,1004]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFB5A19FEFA988FDC" box="[1230,1337,979,1005]" italics="true" pageId="7" pageNumber="519">variabilis</emphasis>
(Fallén)
</taxonomicName>
and 
<emphasis id="CFB5EAE30174670BFF5E19DAFF7A8821" box="[202,219,1015,1040]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFF7919DAFD208820" authority="Bartak &amp; Kubik" authorityName="Bartak &amp; Kubik" box="[237,641,1015,1041]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFF6219DAFEAB8821" box="[246,266,1015,1040]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFEB219DAFE198820" box="[294,440,1015,1041]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
Barták &amp; Kubík
</taxonomicName>
, because these three species lack acrostichal setae. The new species can be readily distinguished from these two species by entirely yellow thorax and abdomen. One more species assigned very recently to 
<emphasis id="CFB5EAE30174670BFE561E12FE728869" box="[450,467,1087,1112]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. (
<taxonomicName id="3AC14D720174670BFE7A1E12FDC18868" authorityName="Schiner" authorityYear="1860" box="[494,608,1087,1113]" class="Insecta" family="Empididae" genus="Holoclera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="CFB5EAE30174670BFE7A1E12FDC18868" box="[494,608,1087,1113]" italics="true" pageId="7" pageNumber="519">Holoclera</emphasis>
</taxonomicName>
), 
<emphasis id="CFB5EAE30174670BFDEE1E12FD2A8869" box="[634,651,1087,1112]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. (
<emphasis id="CFB5EAE30174670BFD311E12FD188869" box="[677,697,1087,1112]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFD401E12FCE78868" box="[724,838,1087,1113]" italics="true" pageId="7" pageNumber="519">daugeroni</emphasis>
Barták &amp; Kokan differs from 
<emphasis id="CFB5EAE30174670BFB391E12FB1F8869" box="[1197,1214,1087,1112]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFB441E12FAD28868" box="[1232,1395,1087,1113]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="variabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFB4D1E12FB4C8869" box="[1241,1261,1087,1112]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFA9C1E12FAD28868" box="[1288,1395,1087,1113]" italics="true" pageId="7" pageNumber="519">variabilis</emphasis>
</taxonomicName>
, 
<emphasis id="CFB5EAE30174670BFA101E12FA348869" box="[1412,1429,1087,1112]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFF031E4EFEFF884C" box="[151,350,1123,1149]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFF351E4EFF14884D" box="[161,181,1123,1148]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFF581E4EFEFF884C" box="[204,350,1123,1149]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
</taxonomicName>
and 
<emphasis id="CFB5EAE30174670BFE021E4EFE06884D" box="[406,423,1123,1148]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. (
<emphasis id="CFB5EAE30174670BFE2A1E4EFE73884D" box="[446,466,1123,1148]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFE7E1E4EFD9F884C" box="[490,574,1123,1149]" italics="true" pageId="7" pageNumber="519">richteri</emphasis>
<emphasis id="CFB5EAE30174670BFDD11E4EFD3E884C" bold="true" box="[581,671,1123,1149]" pageId="7" pageNumber="519">
<taxonomicNameLabel id="D48657980174670BFDD11E4EFD3E884C" box="[581,671,1123,1149]" pageId="7" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
primarily by pale setose abdomen (
<bibRefCitation id="99504B000174670BFBA41E4EFAC8884C" author="Daugeron, C. &amp; Lefebvre, V." box="[1072,1385,1122,1149]" pageId="7" pageNumber="519" pagination="605 - 610" refId="ref21546" refString="Daugeron, C. &amp; Lefebvre, V. (2015) Descriptions of two new species of Empidinae Schiner, 1862 (Diptera: Empididae) from the Mercantour National Park, France. Zoosystema, 37 (4), 605 - 610. https: // doi. org / 10.5252 / z 2015 n 4 a 6" type="journal article" year="2015">Daugeron &amp; Lefebvre 2015</bibRefCitation>
—as 
<taxonomicName id="3AC14D720174670BFF031EAAFDFB8890" authorityName="Loew" authorityYear="1861" box="[151,602,1159,1185]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="brevis" subGenus="Lundstroemiella">
<emphasis id="CFB5EAE30174670BFF031EAAFE988890" box="[151,313,1159,1185]" italics="true" pageId="7" pageNumber="519">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE30174670BFED91EAAFDA58890" box="[333,516,1159,1185]" italics="true" pageId="7" pageNumber="519">Lundstroemiella</emphasis>
) 
<emphasis id="CFB5EAE30174670BFD831EAAFDFB8890" box="[535,602,1159,1185]" italics="true" pageId="7" pageNumber="519">brevis</emphasis>
</taxonomicName>
, preoccupied; 
<bibRefCitation id="99504B000174670BFC931EAAFBAC8890" author="Bartak, M. &amp; Kokan, B." box="[775,1037,1159,1185]" pageId="7" pageNumber="519" pagination="425 - 433" refId="ref21144" refString="Bartak, M. &amp; Kokan, B. (2021) New data about Empididae and Hybotidae (Diptera, Empidoidea) from Croatia: new species, new name, new records. Natura Croatica, 30, 425 - 433. https: // doi. org / 10.20302 / NC. 2021.30.28" type="journal article" year="2021">Barták &amp; Kokan 2021</bibRefCitation>
). The new species is more similar to 
<emphasis id="CFB5EAE30174670BFF211E86FF6788F5" box="[181,198,1195,1220]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFF411E86FE3C88F4" box="[213,413,1195,1221]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFF4A1E86FF5388F5" box="[222,242,1195,1220]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFE9F1E86FE3C88F4" box="[267,413,1195,1221]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
</taxonomicName>
sharing the absence of anteroventral setae on mid and hind tibiae and the presence of two very short apical setae on the hypandrium. Besides the colour of the thorax and abdomen, the male of the new species differs from the male of 
<emphasis id="CFB5EAE30174670BFD811EDEFD87893D" box="[533,550,1267,1292]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFDAC1EDEFCA3893C" box="[568,770,1267,1293]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFDD51EDEFDF4893D" box="[577,597,1267,1292]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFDE41EDEFCA3893C" box="[624,770,1267,1293]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
</taxonomicName>
by mostly brown halteres and deeply brownish infuscate wings. The female of the new species has yellow halteres and almost hyaline wings. However, it differs from 
<emphasis id="CFB5EAE30174670BFF031F16FF098965" box="[151,168,1339,1364]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFF2C1F16FE208964" box="[184,385,1339,1365]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFF561F16FF778965" box="[194,214,1339,1364]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFF7B1F16FE208964" box="[239,385,1339,1365]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
</taxonomicName>
primarily by shorter stylus (about 1/3 of postpedicel length versus more than half length of postpedicel). It should be noted that the new species has a shorter seta in the posteroapical comb of hind tibia than 
<emphasis id="CFB5EAE30174670BFF031FAEFF0989AD" box="[151,168,1411,1436]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFF221FAEFEF789AC" box="[182,342,1411,1437]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="variabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFF541FAEFF7589AD" box="[192,212,1411,1436]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFF7F1FAEFEF789AC" box="[235,342,1411,1437]" italics="true" pageId="7" pageNumber="519">variabilis</emphasis>
</taxonomicName>
and 
<emphasis id="CFB5EAE30174670BFE191FAEFE3F89AD" box="[397,414,1411,1436]" italics="true" pageId="7" pageNumber="519">R</emphasis>
. 
<taxonomicName id="3AC14D720174670BFE381FAEFDD289AC" box="[428,627,1411,1437]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="519" phylum="Arthropoda" rank="species" species="subvariabilis" subGenus="Holoclera">
(
<emphasis id="CFB5EAE30174670BFE211FAEFE6889AD" box="[437,457,1411,1436]" italics="true" pageId="7" pageNumber="519">H</emphasis>
.) 
<emphasis id="CFB5EAE30174670BFE751FAEFDD289AC" box="[481,627,1411,1437]" italics="true" pageId="7" pageNumber="519">subvariabilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A0174670BFF531F8BFE088A1C" pageId="7" pageNumber="519" type="distribution">
<paragraph id="FD7E36F10174670BFF531F8BFB6E89F0" blockId="7.[151,1437,151,1581]" box="[199,1231,1446,1473]" pageId="7" pageNumber="519">
<emphasis id="CFB5EAE30174670BFF531F8BFEC089F1" bold="true" box="[199,353,1446,1472]" pageId="7" pageNumber="519">Distribution.</emphasis>
<collectingCountry id="85D676610174670BFEF31F8AFE1489F0" box="[359,437,1447,1473]" name="Russia" pageId="7" pageNumber="519">Russia</collectingCountry>
. Known only from the North Caucasus (
<collectingRegion id="3F05F8130174670BFCEE1F8AFC7489F0" box="[890,981,1447,1473]" country="Russia" name="Adygeya" pageId="7" pageNumber="519">Adygea</collectingRegion>
, Krasnodarskiy Terr.).
</paragraph>
<paragraph id="FD7E36F10174670BFF531FE7FE088A1C" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="519">
<emphasis id="CFB5EAE30174670BFF531FE7FDF089D4" bold="true" box="[199,593,1482,1509]" pageId="7" pageNumber="519">Habitat and seasonal occurrence.</emphasis>
According to label data, the new species appears to inhabit mountain slopes covered with broad-leafed forests (
<taxonomicName id="3AC14D720174670BFD8B1FC2FD268A38" box="[543,647,1519,1545]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="519" phylum="Tracheophyta" rank="genus">
<emphasis id="CFB5EAE30174670BFD8B1FC2FD268A38" box="[543,647,1519,1545]" italics="true" pageId="7" pageNumber="519">Carpinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="3AC14D720174670BFD021FC2FCA18A38" box="[662,768,1519,1545]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="519" phylum="Tracheophyta" rank="genus">
<emphasis id="CFB5EAE30174670BFD021FC2FCA18A38" box="[662,768,1519,1545]" italics="true" pageId="7" pageNumber="519">Castanea</emphasis>
</taxonomicName>
, 
<taxonomicName id="3AC14D720174670BFC9A1FC2FCF58A39" box="[782,852,1519,1544]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="519" phylum="Tracheophyta" rank="genus">
<emphasis id="CFB5EAE30174670BFC9A1FC2FCF58A39" box="[782,852,1519,1544]" italics="true" pageId="7" pageNumber="519">Fagus</emphasis>
</taxonomicName>
), occurring unusually late around the end of August until the end of October.
</paragraph>
</subSubSection>
</treatment>
</document>