418 lines
56 KiB
XML
418 lines
56 KiB
XML
<document id="C81A1DDBC5A0A80991B211AAF68C7EC4" ID-CLB-Dataset="24334" ID-DOI="10.11646/zootaxa.4722.4.5" ID-GBIF-Dataset="740ce08a-799c-4d63-97cd-0b58c2cf5a79" ID-ISSN="1175-5326" ID-Zenodo-Dep="3609601" ID-ZooBank="BFDCA3E9-187B-49A7-9758-B0096CC4A281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579166408655" checkinUser="plazi" docAuthor="Muthukatturaja, Marimuthu, Balasubramanian, Chelliah & Murugan, Alagumalai" docDate="2020" docId="670C87FEFFB51F30FF004ED4C074FAFC" docLanguage="en" docName="zootaxa.4722.4.5.pdf" docOrigin="Zootaxa 4722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clypeocaenis sharadhae Balasubramanian & Muthukatturaja 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="369" masterDocId="9B35FF86FFB11F38FF974A5DC34DFFC8" masterDocTitle="Two new species of Clypeocaenis (Ephemeroptera: Caenidae) from Western Ghats, Southern India" masterLastPageNumber="370" masterPageNumber="361" pageNumber="365" updateTime="1698777561574" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="B5442F8C821D159D3C0EC83E06CE04E8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="494E31C832CEBA5C7BCF0B47C835D4F4">
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<mods:title id="6EA8B85AD43A6E976832BF86BE9C14B1">Two new species of Clypeocaenis (Ephemeroptera: Caenidae) from Western Ghats, Southern India</mods:title>
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</mods:titleInfo>
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<mods:name id="95D0B1961F6C98964C3E6B3B6DA7F09C" type="personal">
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<mods:roleTerm id="81AE2E4AD471CC5FFB9F52027B3359A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E8A32732A98777E6A72402F23CDA0B4">Muthukatturaja, Marimuthu</mods:namePart>
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<mods:roleTerm id="282A716BADC8C68E40CEB40EF7E6EBD8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3D47F49E614D325F9B56CF7BC6C71202">Balasubramanian, Chelliah</mods:namePart>
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</mods:name>
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<mods:name id="1B580A4177844BFF8FFF6097D168AEEF" type="personal">
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<mods:roleTerm id="466672889F0EA140814D32A6C4D7502A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D2949201880526C8F156B5D34A5985E6">Murugan, Alagumalai</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="BE1326D32A90F23FA0A4C184A9931A5C">text</mods:typeOfResource>
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<mods:titleInfo id="1850DD5FA27CEBE6F5834E25A99C374C">
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<mods:title id="0660E7AB303BC70133B84C9394D822F0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="331E88222D69FC132AB2914FBA1C9FE2">
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<mods:date id="A4AC67847A23FEB81B28700251618552">2020</mods:date>
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<mods:detail id="A7BC2E98A26DB8ED538E99BA7D9EC609" type="pubDate">
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<mods:number id="35AFF9681EA3362FE30D9601EE28DF31">2020-01-15</mods:number>
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<mods:number id="625309017F0F55E86C0017B36816F3DA">4722</mods:number>
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<mods:detail id="3CFF015FAC9122A16F78101723E6464B" type="issue">
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<mods:number id="3C749F8FF992C776E6A8B639EAC88F7D">4</mods:number>
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<mods:start id="9195CDC0E5797E932AE63BC18616709E">361</mods:start>
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<mods:classification id="946985D6746CB61B7F1DCD24A3DBBFB3">journal article</mods:classification>
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<mods:identifier id="488483702E650BA906032C354C97951C" type="CLB-Dataset">24334</mods:identifier>
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<mods:identifier id="3A327F079DA421ECFC8B1504B2E9E5E7" type="DOI">10.11646/zootaxa.4722.4.5</mods:identifier>
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<mods:identifier id="4976D93084B3A8DA8B0FB0B7CF494196" type="GBIF-Dataset">740ce08a-799c-4d63-97cd-0b58c2cf5a79</mods:identifier>
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<mods:identifier id="E8EDFB26780B96A6FDB6B16ABC32D3F8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0DFD0E3EA898BD67F1DE8EC5435FF717" type="Zenodo-Dep">3609601</mods:identifier>
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<mods:identifier id="535B8793A85C685743F08CEED72A0A47" type="ZooBank">BFDCA3E9-187B-49A7-9758-B0096CC4A281</mods:identifier>
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<treatment id="670C87FEFFB51F30FF004ED4C074FAFC" ID-DOI="http://doi.org/10.5281/zenodo.5920340" ID-GBIF-Taxon="161755922" ID-Zenodo-Dep="5920340" LSID="urn:lsid:plazi:treatment:670C87FEFFB51F30FF004ED4C074FAFC" httpUri="http://treatment.plazi.org/id/670C87FEFFB51F30FF004ED4C074FAFC" lastPageId="8" lastPageNumber="369" pageId="4" pageNumber="365">
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<subSubSection id="A7BF6563FFB51F3CFF004ED4C748FB6B" box="[151,1029,1161,1188]" pageId="4" pageNumber="365" type="nomenclature">
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<paragraph id="EF1A36E8FFB51F3CFF004ED4C748FB6B" blockId="4.[151,1029,1161,1223]" box="[151,1029,1161,1188]" pageId="4" pageNumber="365">
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<heading id="B4528184FFB51F3CFF004ED4C748FB6B" bold="true" box="[151,1029,1161,1188]" fontSize="11" level="1" pageId="4" pageNumber="365" reason="1">
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<emphasis id="DDD1EAFAFFB51F3CFF004ED4C748FB6B" bold="true" box="[151,1029,1161,1188]" pageId="4" pageNumber="365">
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<taxonomicName id="28A54D6BFFB51F3CFF004ED4C0D7FB6B" authority="Balasubramanian & Muthukatturaja" authorityName="Balasubramanian & Muthukatturaja" authorityYear="2020" box="[151,922,1161,1188]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="sharadhae" status="sp. nov.">
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<emphasis id="DDD1EAFAFFB51F3CFF004ED4C2F0FB6C" bold="true" box="[151,445,1161,1188]" italics="true" pageId="4" pageNumber="365">Clypeocaenis sharadhae</emphasis>
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Balasubramanian & Muthukatturaja
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</taxonomicName>
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,
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<taxonomicNameLabel id="C6E25781FFB51F3CFC304ED4C748FB6B" box="[935,1029,1161,1187]" pageId="4" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A7BF6563FFB51F3CFF004EF0C22AFB0F" box="[151,359,1196,1223]" pageId="4" pageNumber="365" type="description">
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<paragraph id="EF1A36E8FFB51F3CFF004EF0C22AFB0F" blockId="4.[151,1029,1161,1223]" box="[151,359,1196,1223]" pageId="4" pageNumber="365">
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(
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||
<figureCitation id="779E2A6DFFB51F3CFF084EF0C241FB0F" box="[159,268,1197,1223]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1971,1996]" captionTargetBox="[239,1266,1212,1928]" captionTargetId="figure@1.[221,1290,1202,1952]" captionTargetPageId="1" captionText="FIGURES 1–2. Mature nymph, dorsal view. 1–Head of Clypeocaenis gayathri sp. nov., 2–Head of Clypeocaenis sharadhae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3609603" httpUri="https://zenodo.org/record/3609603/files/figure.png" pageId="4" pageNumber="365">Figures 2</figureCitation>
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,
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<figureCitation id="779E2A6DFFB51F3CFE8F4EF1C212FB0F" box="[280,351,1196,1223]" captionStart-0="FIGURES 17–21" captionStart-1="FIGURES 22–26" captionStart-2="FIGURES 27–35" captionStartId-0="5.[151,264,1965,1990]" captionStartId-1="6.[151,264,1983,2008]" captionStartId-2="7.[151,264,1984,2009]" captionTargetBox-0="[178,1401,553,1932]" captionTargetBox-1="[156,1419,553,1945]" captionTargetBox-2="[171,1418,590,1956]" captionTargetId-0="figure@5.[160,1427,517,1946]" captionTargetId-1="figure@6.[151,1436,553,1963]" captionTargetId-2="figure@7.[164,1424,589,1964]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." captionText-1="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." captionText-2="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi-0="http://doi.org/10.5281/zenodo.3609609" figureDoi-1="http://doi.org/10.5281/zenodo.3609611" figureDoi-2="http://doi.org/10.5281/zenodo.3609613" httpUri-0="https://zenodo.org/record/3609609/files/figure.png" httpUri-1="https://zenodo.org/record/3609611/files/figure.png" httpUri-2="https://zenodo.org/record/3609613/files/figure.png" pageId="4" pageNumber="365">17–35</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A7BF6563FFB51F3CFF004EA9C2A9FAB3" pageId="4" pageNumber="365" type="materials_examined">
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<paragraph id="EF1A36E8FFB51F3CFF004EA9C2A9FAB3" blockId="4.[151,1436,1268,2015]" pageId="4" pageNumber="365">
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<emphasis id="DDD1EAFAFFB51F3CFF004EA9C234FAC7" bold="true" box="[151,377,1268,1295]" italics="true" pageId="4" pageNumber="365">Material examined.</emphasis>
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||
<materialsCitation id="5FCD3CB5FFB51F3CFE164EA8C17EFA9F" ID-GBIF-Occurrence="2550485835" collectingDate="2018-04-01" collectorName="Reg. No. & Balasubramanian & Muthukatturaja." country="India" latitude="13.6075" location="Chikkamagalore district" municipality="Tunga river" pageId="4" pageNumber="365" specimenCount="669" stateProvince="Karnataka state" typeStatus="holotype">
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||
<emphasis id="DDD1EAFAFFB51F3CFE164EA8C2A3FAC7" bold="true" box="[385,494,1269,1295]" pageId="4" pageNumber="365">
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||
<typeStatus id="301E884AFFB51F3CFE164EA8C2A3FAC7" box="[385,494,1269,1295]" pageId="4" pageNumber="365" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
(in ethanol): male imago (reared from nymph) (
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||
<collectorName id="4250533EFFB51F3CFB994EA8C720FAC7" box="[1038,1133,1269,1295]" pageId="4" pageNumber="365">Reg.No.</collectorName>
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||
ZSI–SRC/ I/E 464),
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<collectingCountry id="97B27678FFB51F3CFACB4EA8C6D5FAC7" box="[1372,1432,1269,1295]" name="India" pageId="4" pageNumber="365">India</collectingCountry>
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||
,
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<collectingMunicipality id="0F7EAC92FFB51F3CFF004F44C256FAFB" box="[151,283,1305,1331]" pageId="4" pageNumber="365">Tunga river</collectingMunicipality>
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||
,
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||
<location id="EA7A6033FFB51F3CFEB24F44C178FAFB" LSID="urn:lsid:plazi:treatment:670C87FEFFB51F30FF004ED4C074FAFC:EA7A6033FFB51F3CFEB24F44C178FAFB" box="[293,565,1305,1331]" country="India" latitude="13.6075" municipality="Tunga river" name="Chikkamagalore district" pageId="4" pageNumber="365" stateProvince="Karnataka state">Chikkamagalore district</location>
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||
,
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||
<collectingRegion id="2D61F80AFFB51F3CFDA84F44C1A3FAFB" box="[575,750,1305,1331]" country="India" name="Karnataka" pageId="4" pageNumber="365">Karnataka state</collectingRegion>
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||
,
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||
<date id="9B1B1028FFB51F3CFD6F4F44C030FAFB" box="[760,893,1305,1331]" pageId="4" pageNumber="365" value="2018-04-01">
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||
<collectingDate id="8B5FE9C0FFB51F3CFD6F4F44C030FAFB" box="[760,893,1305,1331]" pageId="4" pageNumber="365" value="2018-04-01">
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||
01-IV-
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||
<quantity id="285D9B0DFFB51F3CFCD54F44C030FAFB" box="[834,893,1305,1331]" metricMagnitude="6" metricUnit="m" metricValue="2.018669" pageId="4" pageNumber="365" unit="m" value="2018669.0">2018</quantity>
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</collectingDate>
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</date>
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,
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<specimenCount id="F9A3FD61FFB51F3CFC104F44C099FAFB" box="[903,980,1305,1331]" pageId="4" pageNumber="365" type="generic">669 m.</specimenCount>
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(
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<geoCoordinate id="8A91502FFFB51F3CFC754F45C722FAFB" box="[994,1135,1304,1331]" degrees="13" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="365" precision="15" seconds="27" value="13.6075">13º36′27″ N</geoCoordinate>
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,
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<geoCoordinate id="8A91502FFFB51F3CFBEE4F45C64FFAFB" box="[1145,1282,1304,1331]" pageId="4" pageNumber="365">75º19′89″ E</geoCoordinate>
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), collected by
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||
<collectorName id="4250533EFFB51F3CFF004F60C214FA9F" box="[151,345,1341,1367]" pageId="4" pageNumber="365">Balasubramanian</collectorName>
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&
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<collectorName id="4250533EFFB51F3CFEE84F60C17EFA9F" box="[383,563,1341,1367]" pageId="4" pageNumber="365">Muthukatturaja.</collectorName>
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</materialsCitation>
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<materialsCitation id="5FCD3CB5FFB51F3CFDAD4F60C2ADFAB3" ID-GBIF-Occurrence="2550485838" collectingDate="2018-04-01" collectorName="Reg. No." country="India" latitude="13.6075" location="Chikkamagalore district" municipality="Tunga river" pageId="4" pageNumber="365" specimenCount="1" stateProvince="Karnataka" typeStatus="paratype">
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||
<emphasis id="DDD1EAFAFFB51F3CFDAD4F60C1F6FA9F" bold="true" box="[570,699,1341,1367]" pageId="4" pageNumber="365">
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||
<typeStatus id="301E884AFFB51F3CFDAD4F60C1FBFA9F" box="[570,694,1341,1367]" pageId="4" pageNumber="365" type="paratype">Paratypes</typeStatus>
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||
:
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||
</emphasis>
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11 mature nymphs and one female imago, same data as holotype (
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<collectorName id="4250533EFFB51F3CFF084F3CC3B3FAB3" box="[159,254,1377,1403]" pageId="4" pageNumber="365">Reg.No.</collectorName>
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||
ZSI–SRC/ I/E 465)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A7BF6563FFB51F30FF504FD9C6C5FBFF" lastPageId="8" lastPageNumber="369" pageId="4" pageNumber="365" type="diagnosis">
|
||
<paragraph id="EF1A36E8FFB51F3CFF504FD9C040F9BF" blockId="4.[151,1436,1268,2015]" pageId="4" pageNumber="365">
|
||
<emphasis id="DDD1EAFAFFB51F3CFF504FD9C009FA57" bold="true" box="[199,836,1412,1439]" pageId="4" pageNumber="365">
|
||
Diagnosis.
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<taxonomicName id="28A54D6BFFB51F3CFEDB4FD9C12AFA57" authority="Muthukatturaja & Balasubramanian & Murugan, 2020" authorityName="Muthukatturaja & Balasubramanian & Murugan" authorityYear="2020" box="[332,615,1412,1439]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="sharadhae" status="sp. nov.">
|
||
<emphasis id="DDD1EAFAFFB51F3CFEDB4FD9C12AFA57" bold="true" box="[332,615,1412,1439]" italics="true" pageId="4" pageNumber="365">Clypeocaenis sharadhae</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="C6E25781FFB51F3CFDE54FD8C19EFA57" box="[626,723,1413,1439]" pageId="4" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
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||
: Nymph:
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||
</emphasis>
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||
(i) clypeus strongly protruded with two long setae, absence of setae on pedicel; (ii) femora with ovoid large macula; (iii) foretibia with two rows of long filtering setae on lateral margin and one row on ventral margin arranged vertically, foretarsus with transverse row of long setae, hindtibia with row of lanceolate setae on lateral margin; (iv) gills III–VI approximately triangular shaped, fringed bifid fimbriae, gill III with 12 bifurcated, 2 trifurcated fimbriae and 17 lanceolate setae at margins; (v) gill VI with 8 bifurcated fimbriae, 19 lanceolate setae on ventral margin and lack of trifurcated fimbriae; (vi) small medial projection of tergum II upright and slightly curved backwards.
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</paragraph>
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<paragraph id="EF1A36E8FFB51F3CFF504CDCC397F8CF" blockId="4.[151,1436,1268,2015]" pageId="4" pageNumber="365">
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||
<emphasis id="DDD1EAFAFFB51F3CFF504CDCC228F953" bold="true" box="[199,357,1665,1691]" pageId="4" pageNumber="365">Imago, male:</emphasis>
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(i) 2 cross veins in proximal half of R; (ii) C and Sc brown basally and washed brown distally, remainders translucent; (iii) vein MP forked at near ½ of distance from base to margin, fork symmetrical; (iv) penis lobes completely fused and not produced laterally; (v) absence of short bristles on the posterior margin of penis lobes.
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</paragraph>
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<paragraph id="EF1A36E8FFB51F3CFF504D4CC606F887" blockId="4.[151,1436,1268,2015]" pageId="4" pageNumber="365">
|
||
<emphasis id="DDD1EAFAFFB51F3CFF504D4CC2BCF8E3" bold="true" box="[199,497,1808,1835]" pageId="4" pageNumber="365">
|
||
<emphasis id="DDD1EAFAFFB51F3CFF504D4CC212F8E3" bold="true" box="[199,351,1809,1835]" italics="true" pageId="4" pageNumber="365">Descriptions.</emphasis>
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Male imago
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</emphasis>
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||
(
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<figureCitation id="779E2A6DFFB51F3CFE684D4CC132F8E3" box="[511,639,1809,1835]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="4" pageNumber="365">Figs 17–21</figureCitation>
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) (in alcohol). Body length,
|
||
<quantity id="285D9B0DFFB51F3CFC254D4CC746F8E3" box="[946,1035,1809,1835]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="365" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; cerci lengths,
|
||
<quantity id="285D9B0DFFB51F3CFB274D4DC644F8E3" box="[1200,1289,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="365" unit="mm" value="3.5">3.5 mm</quantity>
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||
; median caudal filament,
|
||
<quantity id="285D9B0DFFB51F3CFEBB4D68C2C8F887" box="[300,389,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="365" unit="mm" value="3.7">3.7 mm</quantity>
|
||
; length of antenna,
|
||
<quantity id="285D9B0DFFB51F3CFDF74D69C18AF887" box="[608,711,1844,1871]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="365" unit="mm" value="0.05">0.05 mm</quantity>
|
||
; length of forewing,
|
||
<quantity id="285D9B0DFFB51F3CFC264D68C747F887" box="[945,1034,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="365" unit="mm" value="3.3">3.3 mm</quantity>
|
||
; width of forewing,
|
||
<quantity id="285D9B0DFFB51F3CFB794D68C60AF887" box="[1262,1351,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="365" unit="mm" value="1.3">1.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB51F3CFF504D04C0A2F873" blockId="4.[151,1436,1268,2015]" pageId="4" pageNumber="365">
|
||
<emphasis id="DDD1EAFAFFB51F3CFF504D04C24AF8BB" bold="true" box="[199,263,1881,1907]" pageId="4" pageNumber="365">Head</emphasis>
|
||
pale brown, length
|
||
<quantity id="285D9B0DFFB51F3CFE7C4D04C108F8BB" box="[491,581,1881,1907]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="365" unit="mm" value="0.1">0.1 mm</quantity>
|
||
, width
|
||
<quantity id="285D9B0DFFB51F3CFD0D4D04C1B9F8BB" box="[666,756,1881,1907]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="365" unit="mm" value="0.4">0.4 mm</quantity>
|
||
; anterior margin and area around antennal base pale brown; scape and pedicel of antennae pale brown and subequal in length, flagellum translucent; dorsal diameter of eye
|
||
<quantity id="285D9B0DFFB51F3CFF004DFDC241F873" box="[151,268,1952,1979]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="365" unit="mm" value="0.075">0.075 mm</quantity>
|
||
; distance between compound eyes
|
||
<quantity id="285D9B0DFFB51F3CFD024DFCC1A3F873" box="[661,750,1953,1979]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="365" unit="mm" value="0.1">0.1 mm</quantity>
|
||
. Eyes and ocelli black.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB51F3DFF504D98C1FCFED7" blockId="4.[151,1436,1268,2015]" lastBlockId="5.[151,1437,152,503]" lastPageId="5" lastPageNumber="366" pageId="4" pageNumber="365">
|
||
<emphasis id="DDD1EAFAFFB51F3CFF504D98C26DF817" bold="true" box="[199,288,1989,2015]" pageId="4" pageNumber="365">Thorax</emphasis>
|
||
uniformly yellowish brown, lateral margins brown.
|
||
<emphasis id="DDD1EAFAFFB51F3CFCE94D98C0F9F817" bold="true" box="[894,948,1989,2015]" italics="true" pageId="4" pageNumber="365">Legs</emphasis>
|
||
translucent; length of segments in foreleg (
|
||
<figureCitation id="779E2A6DFFB41F3DFF374AC4C3BDFF7B" box="[160,240,153,179]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="5" pageNumber="366">Fig. 18</figureCitation>
|
||
) femur/ tibia/ tarsus (mm) viz., 0.4/ 0.6/ 0.52; length ratio of tarsal segments (mm) viz., 0.02/ 0.25/ 0.1/ 0.1/ 0.05.
|
||
<emphasis id="DDD1EAFAFFB41F3DFF424AE0C25FFF1F" bold="true" box="[213,274,189,215]" italics="true" pageId="5" pageNumber="366">Wing</emphasis>
|
||
: Forewings (
|
||
<figureCitation id="779E2A6DFFB41F3DFE324AE0C2B8FF1F" box="[421,501,189,215]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="5" pageNumber="366">Fig. 17</figureCitation>
|
||
) oval, with few cross veins; cross veins arranged singly; two cross veins at proximal half of R; C and Sc brown basally and washed brown distally, remainders translucent; vein MP forked near half distance from base to margin, fork symmetrical.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB41F3DFF504B75C233FE67" blockId="5.[151,1437,152,503]" pageId="5" pageNumber="366">
|
||
<emphasis id="DDD1EAFAFFB41F3DFF504B75C208FE8B" bold="true" box="[199,325,296,323]" pageId="5" pageNumber="366">Abdomen:</emphasis>
|
||
tergum pale yellow to translucent; sterna pale.
|
||
<emphasis id="DDD1EAFAFFB41F3DFCFB4B75C09AFE8B" bold="true" box="[876,983,296,323]" italics="true" pageId="5" pageNumber="366">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="779E2A6DFFB41F3DFC7E4B74C771FE8B" box="[1001,1084,297,323]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="5" pageNumber="366">Fig. 19</figureCitation>
|
||
) pale yellow, forceps
|
||
<quantity id="285D9B0DFFB41F3DFAAA4B75C6D5FE8B" box="[1341,1432,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="366" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, penis
|
||
<quantity id="285D9B0DFFB41F3DFF4F4B10C27DFEAF" box="[216,304,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="366" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; penis lobes completely fused and not produced laterally; absence of short bristles on posterior margin of penis lobes; forceps (
|
||
<figureCitation id="779E2A6DFFB41F3DFE3C4B2CC2B3FE43" box="[427,510,369,395]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="5" pageNumber="366">Fig. 20</figureCitation>
|
||
) formed by one segment with small bristles on surface; cerci and median caudal filament translucent.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB41F3DFF504BE4C709FE3F" blockId="5.[151,1437,152,503]" pageId="5" pageNumber="366">
|
||
<emphasis id="DDD1EAFAFFB41F3DFF504BE4C23CFE1B" bold="true" box="[199,369,440,467]" pageId="5" pageNumber="366">Female imago</emphasis>
|
||
(in alcohol). Body length,
|
||
<quantity id="285D9B0DFFB41F3DFD3B4BE4C04AFE1B" box="[684,775,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="366" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; cerci lengths,
|
||
<quantity id="285D9B0DFFB41F3DFC204BE4C75FFE1B" box="[951,1042,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="366" unit="mm" value="3.8">3.8 mm</quantity>
|
||
; median caudal filament,
|
||
<quantity id="285D9B0DFFB41F3DFAAB4BE4C6D5FE1B" box="[1340,1432,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="366" unit="mm" value="4.2">4.2 mm</quantity>
|
||
; length of antenna,
|
||
<quantity id="285D9B0DFFB41F3DFEFF4B80C28CFE3F" box="[360,449,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="366" unit="mm" value="0.3">0.3 mm</quantity>
|
||
; length of forewing,
|
||
<quantity id="285D9B0DFFB41F3DFD3D4B80C04EFE3F" box="[682,771,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="366" unit="mm" value="3.4">3.4 mm</quantity>
|
||
; width of forewing,
|
||
<quantity id="285D9B0DFFB41F3DFC704B80C70DFE3F" box="[999,1088,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="366" unit="mm" value="1.4">1.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="BBDA6660FFB41F3DFF004DF0C26EF822" ID-DOI="http://doi.org/10.5281/zenodo.3609609" ID-Zenodo-Dep="3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="5" pageNumber="366" startId="5.[151,264,1965,1990]" targetBox="[178,1401,553,1932]" targetPageId="5">
|
||
<paragraph id="EF1A36E8FFB41F3DFF004DF0C26EF822" blockId="5.[151,1436,1965,2026]" pageId="5" pageNumber="366">
|
||
<emphasis id="DDD1EAFAFFB41F3DFF004DF0C21AF80E" bold="true" box="[151,343,1965,1990]" pageId="5" pageNumber="366">FIGURES 17–21.</emphasis>
|
||
Imago,
|
||
<taxonomicName id="28A54D6BFFB41F3DFE384DF3C1E1F80E" authority="Muthukatturaja & Balasubramanian & Murugan, 2020" authorityName="Muthukatturaja & Balasubramanian & Murugan" authorityYear="2020" box="[431,684,1966,1990]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="sharadhae" status="sp. nov.">
|
||
<emphasis id="DDD1EAFAFFB41F3DFE384DF3C1E1F80E" box="[431,684,1966,1990]" italics="true" pageId="5" pageNumber="366">Clypeocaenis sharadhae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDD1EAFAFFB41F3DFD224DF3C045F80E" bold="true" box="[693,776,1966,1990]" pageId="5" pageNumber="366">
|
||
<taxonomicNameLabel id="C6E25781FFB41F3DFD224DF3C045F80E" box="[693,776,1966,1990]" pageId="5" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF1A36E8FFB71F3EFF504AC4C7B0FF33" blockId="6.[151,1437,153,539]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFF504AC4C24AFF7B" bold="true" box="[199,263,153,179]" pageId="6" pageNumber="367">Head</emphasis>
|
||
brownish yellow, length
|
||
<quantity id="285D9B0DFFB71F3EFDBC4AC4C1DEFF7B" box="[555,659,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="6" pageNumber="367" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, width
|
||
<quantity id="285D9B0DFFB71F3EFD7D4AC4C008FF7B" box="[746,837,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="367" unit="mm" value="0.4">0.4 mm</quantity>
|
||
; anterior margin and area around antennal base pale brown to yellow; scape and pedicel of antennae pale brown and subequal in length, flagellum translucent; distance between compound eyes
|
||
<quantity id="285D9B0DFFB71F3EFE264ABCC147FF33" box="[433,522,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="367" unit="mm" value="0.1">0.1 mm</quantity>
|
||
; dorsal diameter of eye
|
||
<quantity id="285D9B0DFFB71F3EFC814ABCC030FF33" box="[790,893,225,251]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="367" unit="mm" value="0.06">0.06 mm</quantity>
|
||
. Compound eyes and ocelli black.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB71F3EFF504B58C257FE43" blockId="6.[151,1437,153,539]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFF504B58C26DFED7" bold="true" box="[199,288,261,287]" pageId="6" pageNumber="367">Thorax</emphasis>
|
||
uniformly yellowish brown, lateral margins brown.
|
||
<emphasis id="DDD1EAFAFFB71F3EFCF64B58C0ECFED7" bold="true" box="[865,929,261,287]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFCF64B58C0DAFED7" bold="true" box="[865,919,261,287]" italics="true" pageId="6" pageNumber="367">Legs</emphasis>
|
||
:
|
||
</emphasis>
|
||
foreleg pale brown to translucent, midleg and hindleg translucent; length of segments in foreleg femur/ tibia/ tarsus (mm) viz., 0.5/ 0.7/ 0.6; length of tarsal segments (mm) viz., 0.03/ 0.3/ 0.11/ 0.1/ 0.06;
|
||
<emphasis id="DDD1EAFAFFB71F3EFD124B10C18FFEAF" bold="true" box="[645,706,333,359]" italics="true" pageId="6" pageNumber="367">Wing</emphasis>
|
||
: C and Sc brown basally and washed brown distally, remainders translucent.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB71F3EFF504BC9C61BFE67" blockId="6.[151,1437,153,539]" box="[199,1366,404,431]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFF504BC9C208FE67" bold="true" box="[199,325,404,431]" pageId="6" pageNumber="367">Abdomen:</emphasis>
|
||
tergum pale yellow to translucent; sterna pale; cerci and median caudal filament translucent.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB71F3EFF504BE4C13DFDD3" blockId="6.[151,1437,153,539]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFF504BE4C236FE1B" bold="true" box="[199,379,441,467]" pageId="6" pageNumber="367">Mature nymph</emphasis>
|
||
(
|
||
<figureCitation id="779E2A6DFFB71F3EFE1E4BE4C29CFE1B" box="[393,465,441,467]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1971,1996]" captionTargetBox="[239,1266,1212,1928]" captionTargetId="figure@1.[221,1290,1202,1952]" captionTargetPageId="1" captionText="FIGURES 1–2. Mature nymph, dorsal view. 1–Head of Clypeocaenis gayathri sp. nov., 2–Head of Clypeocaenis sharadhae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3609603" httpUri="https://zenodo.org/record/3609603/files/figure.png" pageId="6" pageNumber="367">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="779E2A6DFFB71F3EFE4A4BE5C169FE1B" box="[477,548,440,467]" captionStart-0="FIGURES 22–26" captionStart-1="FIGURES 27–35" captionStartId-0="6.[151,264,1983,2008]" captionStartId-1="7.[151,264,1984,2009]" captionTargetBox-0="[156,1419,553,1945]" captionTargetBox-1="[171,1418,590,1956]" captionTargetId-0="figure@6.[151,1436,553,1963]" captionTargetId-1="figure@7.[164,1424,589,1964]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." captionText-1="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi-0="http://doi.org/10.5281/zenodo.3609611" figureDoi-1="http://doi.org/10.5281/zenodo.3609613" httpUri-0="https://zenodo.org/record/3609611/files/figure.png" httpUri-1="https://zenodo.org/record/3609613/files/figure.png" pageId="6" pageNumber="367">22–35</figureCitation>
|
||
) (in 95% ethanol). Body length (without cerci):
|
||
<quantity id="285D9B0DFFB71F3EFBAB4BE5C78EFE1B" box="[1084,1219,440,467]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="6" pageNumber="367" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 mm</quantity>
|
||
in male and female nymphs; cerci lengths
|
||
<quantity id="285D9B0DFFB71F3EFE0F4B81C2BFFE3F" box="[408,498,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="367" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; median caudal filament
|
||
<quantity id="285D9B0DFFB71F3EFC864B80C026FE3F" box="[785,875,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="367" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; width of head
|
||
<quantity id="285D9B0DFFB71F3EFB8A4B80C73AFE3F" box="[1053,1143,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="367" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; antennae length
|
||
<quantity id="285D9B0DFFB71F3EFAA94B80C6D4FE3F" box="[1342,1433,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="367" unit="mm" value="1.2">1.2 mm</quantity>
|
||
. General body coloration brownish yellow.
|
||
</paragraph>
|
||
<caption id="BBDA6660FFB71F3EFF004DE2C175F834" ID-DOI="http://doi.org/10.5281/zenodo.3609611" ID-Zenodo-Dep="3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="6" pageNumber="367" startId="6.[151,264,1983,2008]" targetBox="[156,1419,553,1945]" targetPageId="6">
|
||
<paragraph id="EF1A36E8FFB71F3EFF004DE2C175F834" blockId="6.[151,1436,1983,2044]" pageId="6" pageNumber="367">
|
||
<emphasis id="DDD1EAFAFFB71F3EFF004DE2C21AF810" bold="true" box="[151,343,1983,2008]" pageId="6" pageNumber="367">FIGURES 22–26.</emphasis>
|
||
Mature nymph,
|
||
<taxonomicName id="28A54D6BFFB71F3EFD9F4DE2C049F81F" authority="Muthukatturaja & Balasubramanian & Murugan, 2020" authorityName="Muthukatturaja & Balasubramanian & Murugan" authorityYear="2020" box="[520,772,1983,2007]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="sharadhae" status="sp. nov.">
|
||
<emphasis id="DDD1EAFAFFB71F3EFD9F4DE2C049F81F" box="[520,772,1983,2007]" italics="true" pageId="6" pageNumber="367">Clypeocaenis sharadhae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDD1EAFAFFB71F3EFC9A4DE2C02DF81F" bold="true" box="[781,864,1983,2007]" pageId="6" pageNumber="367">
|
||
<taxonomicNameLabel id="C6E25781FFB71F3EFC9A4DE2C02DF81F" box="[781,864,1983,2007]" pageId="6" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF1A36E8FFB61F3FFF504AC4C2F4FDF7" blockId="7.[151,1437,153,575]" pageId="7" pageNumber="368">
|
||
<emphasis id="DDD1EAFAFFB61F3FFF504AC4C24AFF7B" bold="true" box="[199,263,153,179]" pageId="7" pageNumber="368">Head</emphasis>
|
||
(
|
||
<figureCitation id="779E2A6DFFB61F3FFE8E4AC4C22DFF7B" box="[281,352,153,179]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1971,1996]" captionTargetBox="[239,1266,1212,1928]" captionTargetId="figure@1.[221,1290,1202,1952]" captionTargetPageId="1" captionText="FIGURES 1–2. Mature nymph, dorsal view. 1–Head of Clypeocaenis gayathri sp. nov., 2–Head of Clypeocaenis sharadhae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3609603" httpUri="https://zenodo.org/record/3609603/files/figure.png" pageId="7" pageNumber="368">Fig. 2</figureCitation>
|
||
) brownish yellow, eyes black, ocelli paler. Two long setae on anterior margin of clypeus and strongly protruded (
|
||
<figureCitation id="779E2A6DFFB61F3FFEE14AE0C2F1FF1F" box="[374,444,189,215]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1971,1996]" captionTargetBox="[239,1266,1212,1928]" captionTargetId="figure@1.[221,1290,1202,1952]" captionTargetPageId="1" captionText="FIGURES 1–2. Mature nymph, dorsal view. 1–Head of Clypeocaenis gayathri sp. nov., 2–Head of Clypeocaenis sharadhae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3609603" httpUri="https://zenodo.org/record/3609603/files/figure.png" pageId="7" pageNumber="368">Fig. 2</figureCitation>
|
||
). Antennae yellowish transparent distally, absence of short setae at inner margin of pedicel. Epicranial suture paler. Labrum (
|
||
<figureCitation id="779E2A6DFFB61F3FFDA24ABCC1CBFF33" box="[565,646,225,251]" captionStart="FIGURES 22–26" captionStartId="6.[151,264,1983,2008]" captionTargetBox="[156,1419,553,1945]" captionTargetId="figure@6.[151,1436,553,1963]" captionTargetPageId="6" captionText="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." figureDoi="http://doi.org/10.5281/zenodo.3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="7" pageNumber="368">Fig. 22</figureCitation>
|
||
) length
|
||
<quantity id="285D9B0DFFB61F3FFD754ABCC077FF33" box="[738,826,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="368" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; width
|
||
<quantity id="285D9B0DFFB61F3FFC1C4ABCC0B2FF33" box="[907,1023,225,251]" metricMagnitude="-5" metricUnit="m" metricValue="7.7" pageId="7" pageNumber="368" unit="mm" value="0.077">0.077 mm</quantity>
|
||
, emargination without setae, anterior margin with row of long setae, lateral margin with stout setae, dorsum of labrum with few long setae. Mandibles (
|
||
<figureCitation id="779E2A6DFFB61F3FFF374B74C257FE8C" box="[160,282,297,324]" captionStart="FIGURES 22–26" captionStartId="6.[151,264,1983,2008]" captionTargetBox="[156,1419,553,1945]" captionTargetId="figure@6.[151,1436,553,1963]" captionTargetPageId="6" captionText="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." figureDoi="http://doi.org/10.5281/zenodo.3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="7" pageNumber="368">Fig. 23a, b</figureCitation>
|
||
): rows of 4 and 8 long setae present below base of incisors; basal half of mandible chagrinated. Maxilla (
|
||
<figureCitation id="779E2A6DFFB61F3FFF374B10C252FEA0" box="[160,287,333,360]" captionStart="FIGURES 22–26" captionStartId="6.[151,264,1983,2008]" captionTargetBox="[156,1419,553,1945]" captionTargetId="figure@6.[151,1436,553,1963]" captionTargetPageId="6" captionText="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." figureDoi="http://doi.org/10.5281/zenodo.3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="7" pageNumber="368">Fig. 24a, b</figureCitation>
|
||
): galea with single stout spine and ventral base of lacinia with few bipectinate setae; maxillary palp two-segmented; segment II three times longer than segment I, half of segment II covered with setae. Hypopharynx (
|
||
<figureCitation id="779E2A6DFFB61F3FFF374BC8C3B9FE67" box="[160,244,404,431]" captionStart="FIGURES 22–26" captionStartId="6.[151,264,1983,2008]" captionTargetBox="[156,1419,553,1945]" captionTargetId="figure@6.[151,1436,553,1963]" captionTargetPageId="6" captionText="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." figureDoi="http://doi.org/10.5281/zenodo.3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="7" pageNumber="368">Fig. 25</figureCitation>
|
||
): superlingua covered with stout setae and lingua with long setae at apical margin. Labium (
|
||
<figureCitation id="779E2A6DFFB61F3FFB6B4BC8C61DFE78" box="[1276,1360,405,432]" captionStart="FIGURES 22–26" captionStartId="6.[151,264,1983,2008]" captionTargetBox="[156,1419,553,1945]" captionTargetId="figure@6.[151,1436,553,1963]" captionTargetPageId="6" captionText="FIGURES 22–26. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal views. 22–Labrum, 23a, b–Mandibles, 24a, b– Maxillae, 25–Hypopharynx, 26–Labium." figureDoi="http://doi.org/10.5281/zenodo.3609611" httpUri="https://zenodo.org/record/3609611/files/figure.png" pageId="7" pageNumber="368">Fig. 26</figureCitation>
|
||
): glossae ovate with apical spines, paraglossae covered with thin setae, labial palp three-segmented, segment III longer than segments I and II, 2 transverse rows of stout setae at distal margin of segment III, segment I with eight stout setae on outer margin, basal outer margin of proximal segment and apical margin of middle segment of labial palp with two lanceolate setae.
|
||
</paragraph>
|
||
<caption id="BBDA6660FFB61F3FFF004D9DC0EFF834" ID-DOI="http://doi.org/10.5281/zenodo.3609613" ID-Zenodo-Dep="3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="7" pageNumber="368" startId="7.[151,264,1984,2009]" targetBox="[171,1418,590,1956]" targetPageId="7">
|
||
<paragraph id="EF1A36E8FFB61F3FFF004D9DC0EFF834" blockId="7.[151,1436,1984,2045]" pageId="7" pageNumber="368">
|
||
<emphasis id="DDD1EAFAFFB61F3FFF004D9DC21BF811" bold="true" box="[151,342,1984,2009]" pageId="7" pageNumber="368">FIGURES 27–35.</emphasis>
|
||
Mature nymph,
|
||
<taxonomicName id="28A54D6BFFB61F3FFD924D9CC04CF811" authority="Muthukatturaja & Balasubramanian & Murugan, 2020" authorityName="Muthukatturaja & Balasubramanian & Murugan" authorityYear="2020" box="[517,769,1985,2009]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="sharadhae" status="sp. nov.">
|
||
<emphasis id="DDD1EAFAFFB61F3FFD924D9CC04CF811" box="[517,769,1985,2009]" italics="true" pageId="7" pageNumber="368">Clypeocaenis sharadhae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDD1EAFAFFB61F3FFC9E4D9CC011F811" bold="true" box="[777,860,1985,2009]" pageId="7" pageNumber="368">
|
||
<taxonomicNameLabel id="C6E25781FFB61F3FFC9E4D9CC011F811" box="[777,860,1985,2009]" pageId="7" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF1A36E8FFB91F30FF504AC4C163FD07" blockId="8.[151,1437,153,1332]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504AC4C264FF7B" bold="true" box="[199,297,153,179]" pageId="8" pageNumber="369">Thorax:</emphasis>
|
||
brownish yellow, lateral margins washed yellow.
|
||
<emphasis id="DDD1EAFAFFB91F30FCF64AC4C0DAFF7B" bold="true" box="[865,919,153,179]" italics="true" pageId="8" pageNumber="369">Legs</emphasis>
|
||
evenly covered with scales, pale yellow with washed–brown, femora with ovoid large macula without scales and setae. Foreleg (
|
||
<figureCitation id="779E2A6DFFB91F30FBAA4AE0C7DCFF1F" box="[1085,1169,189,215]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 27</figureCitation>
|
||
): length of femur/ tibia/ tarsus/ claw viz., 0.37/ 0.41/ 0.39/
|
||
<quantity id="285D9B0DFFB91F30FDBF4ABCC1DCFF33" box="[552,657,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="369" unit="mm" value="0.14">0.14 mm</quantity>
|
||
. Coxa with few spines on surface of outer margin and pair of long, thin spines on inner margin. Trochanter with two rows of stout spines on outer margin. Femur with 4 long, stout setae at middle of inner margin and dorsal surface with transverse row of lanceolate setae for 2/3 length, with few monopectinate, stout setae and long setae with feathered cap on outer margins. Tibia with two rows of long, filtering setae on lateral margin and one row on ventral margin. Tarsus with row of monopectinate setae on ventral margin and transverse row of long setae on lateral margin. Claw hooked with 5 denticles. Midleg (
|
||
<figureCitation id="779E2A6DFFB91F30FB154BC8C798FE67" box="[1154,1237,405,431]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 28</figureCitation>
|
||
): length of femur/ tibia/ tarsus/ claw viz., 0.33/ 0.24/ 0.17/
|
||
<quantity id="285D9B0DFFB91F30FDF74BE4C185FE1B" box="[608,712,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="369" unit="mm" value="0.1">0.10 mm</quantity>
|
||
. Coxa and trochanter same as foreleg. Femur with few long and short stout spines at outer margin, inner margin with row of stout spines and thin hair-like setae. Tibia with row of long and short stout spines on ventral margin. Tarsus with few long, hair-like setae. Claw long, slender and slightly curved with 4 denticles. Hindleg (
|
||
<figureCitation id="779E2A6DFFB91F30FD994878C12FFDF7" box="[526,610,549,575]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 29</figureCitation>
|
||
): length of femur/ tibia/ tarsus/ claw viz., 0.44/ 0.24/ 0.23/
|
||
<quantity id="285D9B0DFFB91F30FB7B4878C61CFDF7" box="[1260,1361,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="369" unit="mm" value="0.11">0.11 mm</quantity>
|
||
. Coxa, trochanter similar to those of foreleg. Femur with long spines and hair-like setae on inner and outer margins. Tibia with row of long and short stout spines on inner margin, surface with row of lanceolate setae. Tarsus with few long hair-like seate on surface, inner margin with stout spines, lateral margin with row of lanceolate setae. Claw slender and slightly curved with 6 denticles.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB91F30FF504884C79DFBDB" blockId="8.[151,1437,153,1332]" pageId="8" pageNumber="369">
|
||
A
|
||
<emphasis id="DDD1EAFAFFB91F30FF4C4884C276FD3B" bold="true" box="[219,315,729,755]" pageId="8" pageNumber="369">bdomen</emphasis>
|
||
: posterior margins of terga brown, remainders washed yellow. Tergum II (
|
||
<figureCitation id="779E2A6DFFB91F30FB3D4884C64DFD3B" box="[1194,1280,729,755]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 34</figureCitation>
|
||
) with medial projection upright, slightly curved backwards, lateral margin of tergum with spines and surface with scales, medial projection pale brown with scales and without spines. Lateral margin of tergum VII (
|
||
<figureCitation id="779E2A6DFFB91F30FBDD497CC7D6FCF3" box="[1098,1179,800,827]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 35</figureCitation>
|
||
) with short bipectinate setae; dorsomedial margins of terga VII–IX with crowded scales. Sternum pale yellow. Gills (
|
||
<figureCitation id="779E2A6DFFB91F30FB524918C61CFC97" box="[1221,1361,837,863]" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Figs. 30–32</figureCitation>
|
||
): gill I pointed, two segmented, apex of segment I pulvinate; segment II 2.5 times longer than segment I. Gill covers (
|
||
<figureCitation id="779E2A6DFFB91F30FAF84934C3F9FC6F" captionStart="FIGURES 27–35" captionStartId="7.[151,264,1984,2009]" captionTargetBox="[171,1418,590,1956]" captionTargetId="figure@7.[164,1424,589,1964]" captionTargetPageId="7" captionText="FIGURES 27–35. Mature nymph, Clypeocaenis sharadhae sp. nov., dorsal view. 27–Foreleg, 28–Midleg, 29–Hindleg, 30– Gill I, 31–Gill III, 32–Gill VI, 33–Gill cover, 34–Tergum II, 35–Tergum VII." figureDoi="http://doi.org/10.5281/zenodo.3609613" httpUri="https://zenodo.org/record/3609613/files/figure.png" pageId="8" pageNumber="369">Fig. 33</figureCitation>
|
||
) pale brown, evenly covered with scales, inner margins each with row of spines, outer margins each with small ridges (lanceolate like). Gills III–VI approximately triangular shaped with fringed bifid fimbriae; gill III with 12 bifurcated and 2 trifurcated fimbriae, and 17 lanceolate setae on margins; gill VI with 8 bifurcated fimbriae and 19 lanceolate setae on ventral margin and lack of trifurcated fimbriae. Caudal filaments translucent.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB91F30FF504E40C6C5FBFF" blockId="8.[151,1437,153,1332]" box="[199,1416,1053,1080]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504E40C3B9FBFF" bold="true" box="[199,244,1053,1079]" pageId="8" pageNumber="369">Egg</emphasis>
|
||
(
|
||
<figureCitation id="779E2A6DFFB91F30FE944E40C216FBFF" box="[259,347,1053,1079]" captionStart="FIGURES 17–21" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[178,1401,553,1932]" captionTargetId="figure@5.[160,1427,517,1946]" captionTargetPageId="5" captionText="FIGURES 17–21. Imago, Clypeocaenis sharadhae sp. nov. 17–Forewing, 18–Foreleg, 19–Genitalia, 20–Forceps, 21–SEM image of egg." figureDoi="http://doi.org/10.5281/zenodo.3609609" httpUri="https://zenodo.org/record/3609609/files/figure.png" pageId="8" pageNumber="369">Figs. 21</figureCitation>
|
||
) (dissected from female imago) oval shaped. Chorion with single thread running from polar region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A7BF6563FFB91F30FF504E1DC0F7FAC7" pageId="8" pageNumber="369" type="etymology">
|
||
<paragraph id="EF1A36E8FFB91F30FF504E1DC01CFBB7" blockId="8.[151,1437,153,1332]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504E1DC20FFB93" bold="true" box="[199,322,1088,1115]" italics="true" pageId="8" pageNumber="369">Etymology</emphasis>
|
||
. The species is named after the goddess Sharadhambal, whose temple is in Sringeri (Chikkamagalore district,
|
||
<collectingRegion id="2D61F80AFFB91F30FEB14E38C2D3FB48" box="[294,414,1125,1152]" country="India" name="Karnataka" pageId="8" pageNumber="369">Karnataka</collectingRegion>
|
||
), where the new species was collected.
|
||
</paragraph>
|
||
<paragraph id="EF1A36E8FFB91F30FF504ED5C0F7FAC7" blockId="8.[151,1437,153,1332]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504ED5C268FB6B" bold="true" box="[199,293,1160,1187]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504ED5C253FB6B" bold="true" box="[199,286,1160,1187]" italics="true" pageId="8" pageNumber="369">Habitat</emphasis>
|
||
.
|
||
</emphasis>
|
||
Nymphs of this species were found on the undersides of cobbles and pebbles in medium to fast current streams and associated with
|
||
<taxonomicName id="28A54D6BFFB91F30FE4B4EF0C102FB0F" box="[476,591,1197,1223]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DDD1EAFAFFB91F30FE4B4EF0C102FB0F" box="[476,591,1197,1223]" italics="true" pageId="8" pageNumber="369">Ephemera</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="28A54D6BFFB91F30FD104EF0C06CFB00" box="[647,801,1197,1224]" class="Insecta" family="Ephemeridae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="family">Ephemeridae</taxonomicName>
|
||
),
|
||
<taxonomicName id="28A54D6BFFB91F30FCA14EF0C08CFB0F" authorityName="Eaton" authorityYear="1881" box="[822,961,1197,1223]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DDD1EAFAFFB91F30FCA14EF0C08CFB0F" box="[822,961,1197,1223]" italics="true" pageId="8" pageNumber="369">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="28A54D6BFFB91F30FC6E4EF0C7FBFB00" box="[1017,1206,1197,1224]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="28A54D6BFFB91F30FB5D4EF0C67AFB0F" authorityName="Eaton" authorityYear="1871" box="[1226,1335,1197,1223]" class="Insecta" family="Isonychiidae" genus="Isonychia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DDD1EAFAFFB91F30FB5D4EF0C67AFB0F" box="[1226,1335,1197,1223]" italics="true" pageId="8" pageNumber="369">Isonychia</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="28A54D6BFFB91F30FAF84EF0C245FB24" authorityName="Burks" authorityYear="1953" class="Insecta" family="Isonychiidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="family">Isonychiidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="28A54D6BFFB91F30FE8D4E8CC1BDFB24" authorityName="Balasubramanian, Muthukatturaja & Anbalagan" authorityYear="2019" box="[282,752,1233,1260]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="species" species="tungaiensis">
|
||
<emphasis id="DDD1EAFAFFB91F30FE8D4E8CC17FFB23" box="[282,562,1233,1259]" italics="true" pageId="8" pageNumber="369">Rhoenanthus tungaiensis</emphasis>
|
||
(Potamanthidae)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="28A54D6BFFB91F30FCB34E8FC027FB23" authorityName="Leach" authorityYear="1815" box="[804,874,1234,1259]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DDD1EAFAFFB91F30FCB34E8FC027FB23" box="[804,874,1234,1259]" italics="true" pageId="8" pageNumber="369">Baetis</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="28A54D6BFFB91F30FC0A4E8CC748FB24" authorityName="Leach" authorityYear="1815" box="[925,1029,1233,1260]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="369" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
), possibly feeding on fine particulate organic matter available in the water current and also in the substratum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A7BF6563FFB91F30FF504F45C074FAFC" box="[199,825,1304,1332]" pageId="8" pageNumber="369" type="distribution">
|
||
<paragraph id="EF1A36E8FFB91F30FF504F45C074FAFC" blockId="8.[151,1437,153,1332]" box="[199,825,1304,1332]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504F45C217FAFB" bold="true" box="[199,346,1304,1331]" pageId="8" pageNumber="369">
|
||
<emphasis id="DDD1EAFAFFB91F30FF504F45C21EFAFB" bold="true" box="[199,339,1304,1331]" italics="true" pageId="8" pageNumber="369">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
Central Western Ghats,
|
||
<collectingRegion id="2D61F80AFFB91F30FDFB4F44C192FAFB" box="[620,735,1305,1331]" country="India" name="Karnataka" pageId="8" pageNumber="369">Karnataka</collectingRegion>
|
||
(
|
||
<collectingCountry id="97B27678FFB91F30FD794F44C060FAFC" box="[750,813,1305,1332]" name="India" pageId="8" pageNumber="369">India</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |