treatments-xml/data/4A/01/89/4A018914FFD4FFA8A9AE73534B43FADA.xml

<document ID-DOI="10.11646/phytotaxa.566.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="7103538" checkinTime="1663828622333" checkinUser="plazi" docAuthor="Jiménez, Pedro Díaz, Dötterl, Stefan, Fuchs, Roman, Hentrich, Heiko, Pérez-Farrera, Miguel Ángel, Vega, José Padilla &amp; Aguilar-Rodríguez, Pedro A." docDate="2022" docId="4A018914FFD4FFA8A9AE73534B43FADA" docLanguage="en" docName="phytotaxa.566.1.7.pdf" docOrigin="Phytotaxa 566 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Spathiphyllum wilfridianum Diaz Jim. 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="128" masterDocId="B638F16CFFD1FFAFA926703D4A64FF86" masterDocTitle="Two new species of Spathiphyllum (Araceae) from Tabasco, Mexico with notes on their floral scent" masterLastPageNumber="132" masterPageNumber="121" pageNumber="126" updateTime="1663941463158" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
<mods:title>Two new species of Spathiphyllum (Araceae) from Tabasco, Mexico with notes on their floral scent</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiménez, Pedro Díaz</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dötterl, Stefan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5228-1332</mods:nameIdentifier>
<mods:affiliation>Department of Environment &amp; Biodiversity, Plant Ecology, Paris Lodron University of Salzburg, Hellbrunnerstr. 34, 5020 Salzburg, Austria. &amp; stefan. doetterl @ plus. ac. at; https: // orcid. org / 0000 - 0001 - 5228 - 1332</mods:affiliation>
<mods:nameIdentifier type="email">stefan.doetterl@plus.ac.at</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fuchs, Roman</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9423-5588</mods:nameIdentifier>
<mods:affiliation>Department of Environment &amp; Biodiversity, Plant Ecology, Paris Lodron University of Salzburg, Hellbrunnerstr. 34, 5020 Salzburg, Austria. &amp; roman. fuchs @ plus. ac. at; https: // orcid. org / 0000 - 0001 - 9423 - 5588</mods:affiliation>
<mods:nameIdentifier type="email">roman.fuchs@plus.ac.at</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hentrich, Heiko</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1120-5947</mods:nameIdentifier>
<mods:affiliation>Molecular Cell Biology, Botanical Institute, Karlsruhe Institute of Technology, Fritz-Haber-Weg 4, 76131 Karlsruhe, Germany. &amp; heiko. hentrich @ kit. edu; https: // orcid. org / 0000 - 0002 - 1120 - 5947</mods:affiliation>
<mods:nameIdentifier type="email">heiko.hentrich@kit.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pérez-Farrera, Miguel Ángel</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5329-1505</mods:nameIdentifier>
<mods:affiliation>Herbario Eizi Matuda, Laboratorio de Ecologia, Evolutiva, Instituto de Ciencias Biologicas Universidad de Ciencias y Artes de Chiapas. Libramiento Norte Poniente 1550, Lajas Maciel, Tuxtla Gutiérrez, Chis, 29039, México. &amp; miguel. perez @ unicach. mx; https: // orcid. org / 0000 - 0002 - 5329 - 1505</mods:affiliation>
<mods:nameIdentifier type="email">miguel.perez@unicach.mx</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vega, José Padilla</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8359-1077</mods:nameIdentifier>
<mods:affiliation>Raíces de la Montaña, Carretera Pomoquita-Oxolotán S / N, Col. Pomoquita, C. P. 86895, Tacotalpa, Tabasco, México. &amp; jopave @ gmail. com; https: // orcid. org / 0000 - 0001 - 8359 - 1077</mods:affiliation>
<mods:nameIdentifier type="email">jopave@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aguilar-Rodríguez, Pedro A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9275-4322</mods:nameIdentifier>
<mods:affiliation>Instituto de Investigaciones Forestales, Universidad Veracruzana, Parque Ecologico “ El Haya ”, Carretera Antigua a Coatepec, 91070 Xalapa-Enríquez, Veracruz, México. &amp; pedroaguilarr @ gmail. com; https: // orcid. org / 0000 - 0002 - 9275 - 4322</mods:affiliation>
<mods:nameIdentifier type="email">pedroaguilarr@gmail.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>566</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>121</mods:start>
<mods:end>132</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.566.1.7</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7103538</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:4A018914FFD4FFA8A9AE73534B43FADA" httpUri="http://treatment.plazi.org/id/4A018914FFD4FFA8A9AE73534B43FADA" lastPageId="7" lastPageNumber="128" pageId="5" pageNumber="126">
<subSubSection box="[136,767,878,905]" pageId="5" pageNumber="126" type="nomenclature">
<paragraph blockId="5.[136,767,878,905]" box="[136,767,878,905]" pageId="5" pageNumber="126">
<heading box="[136,767,878,905]" fontSize="11" level="2" pageId="5" pageNumber="126" reason="3">
<taxonomicName authority="Diaz Jim." authorityName="Diaz Jim." authorityYear="2022" box="[136,581,878,905]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="wilfridianum" status="sp. nov.">
<emphasis bold="true" box="[136,464,878,905]" italics="true" pageId="5" pageNumber="126">Spathiphyllum wilfridianum</emphasis>
Díaz Jim.
</taxonomicName>
, 
<emphasis box="[592,677,879,904]" italics="true" pageId="5" pageNumber="126">
<taxonomicNameLabel box="[592,677,879,904]" pageId="5" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[691,759,878,904]" captionStart="FIGURE 4" captionStartId="6.[136,229,1421,1443]" captionTargetBox="[346,1242,191,1385]" captionTargetId="figure-23@6.[340,1247,190,1397]" captionTargetPageId="6" captionText="FIGURE 4. Spathiphyllum wilfridianum. A. Cultivated adult plant. B. Adaxial view of the blade. C. Portion of petioles showing entire sheath margins (arrow). D. Inflorescence showing spathe and spadix in anthesis. E. Pistils. Photos by P. Díaz Jiménez." figureDoi="http://doi.org/10.5281/zenodo.7103546" httpUri="https://zenodo.org/record/7103546/files/figure.png" pageId="5" pageNumber="126">Fig. 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="diagnosis">
<paragraph blockId="5.[136,1452,954,1156]" pageId="5" pageNumber="126">
<taxonomicName box="[136,404,954,975]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[136,404,954,975]" italics="true" pageId="5" pageNumber="126">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
is morphologically similar to 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[691,886,954,975]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[691,886,954,975]" italics="true" pageId="5" pageNumber="126">S. cochlearispathum</emphasis>
</taxonomicName>
, but differs from that species in having petioles longer than the blades and sheathed to 
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from the geniculum (
<emphasis box="[751,777,990,1011]" italics="true" pageId="5" pageNumber="126">vs.</emphasis>
as long as the blade and sheathed near or up to the geniculum), sheath margins entire (
<emphasis box="[341,368,1026,1047]" italics="true" pageId="5" pageNumber="126">vs.</emphasis>
wrinkled), blade attenuate at the base (
<emphasis box="[745,771,1026,1047]" italics="true" pageId="5" pageNumber="126">vs.</emphasis>
rounded), primary veins rising at an angle of 40–55° (
<emphasis box="[1295,1322,1026,1047]" italics="true" pageId="5" pageNumber="126">vs.</emphasis>
60–80°), and up to 7 ovules per ovary (
<emphasis box="[436,463,1062,1083]" italics="true" pageId="5" pageNumber="126">vs.</emphasis>
up to 16).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="materials_examined">
<paragraph blockId="5.[136,1452,954,1156]" pageId="5" pageNumber="126">
<emphasis bold="true" box="[136,194,1098,1119]" pageId="5" pageNumber="126">Type:</emphasis>
— 
<materialsCitation collectingDate="2009-11-16" collectionCode="A" collectorName="Dos Montes &amp; la Naturaleza" country="Mexico" elevation="7" latitude="17.75" location="Centro de Interpretacion" longLatPrecision="1277" longitude="-92.75" pageId="5" pageNumber="126" specimenCount="1" stateProvince="Tabasco">
<collectingCountry box="[219,316,1098,1119]" name="Mexico" pageId="5" pageNumber="126">MEXICO</collectingCountry>
. 
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: Municipio Centro, communal land 
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, 
<location LSID="urn:lsid:plazi:treatment:4A018914FFD4FFA8A9AE73534B43FADA:C7776ED9FFD4FFAAAA5C74774E03FBE6" box="[890,1127,1098,1120]" country="Mexico" latitude="17.75" longLatPrecision="1277" longitude="-92.75" name="Centro de Interpretacion" pageId="5" pageNumber="126" stateProvince="Tabasco">Centro de Interpretación</location>
y Convivencia con 
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“Yumka”, 
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, 
<geoCoordinate box="[386,473,1134,1156]" degrees="92" direction="west" minutes="45" orientation="longitude" pageId="5" pageNumber="126" precision="925" value="-92.75">92°45’W</geoCoordinate>
, 
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<quantity box="[484,521,1134,1155]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="126" unit="m" value="7.0">7 m</quantity>
asl
</elevation>
, 
<date box="[565,752,1134,1156]" pageId="5" pageNumber="126" value="2009-11-16">
<collectingDate box="[565,752,1134,1156]" pageId="5" pageNumber="126" value="2009-11-16">16 November 2009</collectingDate>
</date>
, 
<emphasis box="[762,1003,1134,1155]" italics="true" pageId="5" pageNumber="126">
Pedro Díaz Jiménez &amp; 
<collectionCode box="[985,1003,1134,1155]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="126" type="Herbarium">A</collectionCode>
</emphasis>
</materialsCitation>
<emphasis box="[1002,1006,1134,1155]" italics="true" pageId="5" pageNumber="126">.</emphasis>
<materialsCitation box="[1012,1328,1134,1156]" collectionCode="UJAT" country="Mexico" pageId="5" pageNumber="126" specimenCount="1" stateProvince="Tabasco" typeStatus="holotype">
<emphasis box="[1012,1149,1134,1155]" italics="true" pageId="5" pageNumber="126">Garduza 1092</emphasis>
(
<typeStatus box="[1161,1247,1134,1156]" pageId="5" pageNumber="126">holotype</typeStatus>
<collectionCode box="[1253,1318,1134,1155]" country="Mexico" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14347" lsid="urn:lsid:biocol.org:col:14347" name="Universidad Juárez Autónoma de Tabasco" pageId="5" pageNumber="126" type="Herbarium">UJAT</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="description">
<paragraph blockId="5.[136,1452,1202,2093]" pageId="5" pageNumber="126">
Terrestrial; understory herb, up to 
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tall; internodes short, up to 
<quantity box="[911,966,1202,1229]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="126" unit="cm" value="6.0">6 cm</quantity>
diam. LEAVES 
<quantity box="[1154,1293,1202,1229]" metricMagnitude="0" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.67" metricValueMin="1.22" pageId="5" pageNumber="126" unit="cm" value="144.5" valueMax="167.0" valueMin="122.0">122–167 cm</quantity>
long; petioles up to twice as long as the blades, 
<quantity box="[501,624,1238,1265]" metricMagnitude="-1" metricUnit="m" metricValue="8.7" metricValueMax="10.4" metricValueMin="7.0" pageId="5" pageNumber="126" unit="cm" value="87.0" valueMax="104.0" valueMin="70.0">70–104 cm</quantity>
long, 
<quantity box="[693,784,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="126" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
diam., sheathed near the middle, or above to 
<quantity box="[1281,1391,1238,1265]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" metricValueMax="7.3" metricValueMin="3.3" pageId="5" pageNumber="126" unit="cm" value="53.0" valueMax="73.0" valueMin="33.0">33–73 cm</quantity>
from the geniculum, free portion terete, the sheath and free portion pale green and covered in white dots, sheath margins entire, light green or dark green; geniculum 
<quantity box="[632,757,1310,1337]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.3" metricValueMin="2.5" pageId="5" pageNumber="126" unit="cm" value="3.9" valueMax="5.3" valueMin="2.5">2.5–5.3 cm</quantity>
long, 
<quantity box="[829,943,1310,1337]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="9.5" metricValueMin="7.0" pageId="5" pageNumber="126" unit="mm" value="8.25" valueMax="9.5" valueMin="7.0">7–9.5 mm</quantity>
diam., light green and covered in white dots; blades oblong or oblong-elliptic, widest almost in the middle, 50.0– 
<quantity box="[915,1006,1346,1373]" metricMagnitude="-1" metricUnit="m" metricValue="6.55" pageId="5" pageNumber="126" unit="cm" value="65.5">65.5 cm</quantity>
long, 
<quantity box="[1081,1194,1346,1373]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" metricValueMax="2.9" metricValueMin="1.4" pageId="5" pageNumber="126" unit="cm" value="21.5" valueMax="29.0" valueMin="14.0">14–29 cm</quantity>
wide, approx. 3 times longer than wide, acuminate at apex, attenuate (rarely sub-rounded) at the base, subcoriaceous, dark green to yellowishgreen and semi-glossy above, light green semi-glossy below, drying green dark to dark brown above, greenish-brown to light brown and below; midrib sunken, dark green above, and covered in white dots, light green to whitish below, thicker than broad; primary lateral veins, 24–30 pairs, separated 
<quantity box="[856,965,1490,1517]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="5" pageNumber="126" unit="mm" value="13.5" valueMax="20.0" valueMin="7.0">7–20 mm</quantity>
, arising at a 40–55° angle, sunken and dark green above, whitish below; minor veins dark green below. INFLORESCENCE erect, taller than the leaves; peduncle 
<quantity box="[136,276,1562,1589]" metricMagnitude="0" metricUnit="m" metricValue="1.535" metricValueMax="1.68" metricValueMin="1.39" pageId="5" pageNumber="126" unit="cm" value="153.5" valueMax="168.0" valueMin="139.0">139–168 cm</quantity>
long, 
<quantity box="[348,483,1562,1589]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="5" pageNumber="126" unit="mm" value="7.0" valueMax="8.5" valueMin="5.5">5.5–8.5 mm</quantity>
diam., covered in white dots, green to light yellowish-green; spathe cucullate, oblong or oblong-elliptic, 
<quantity box="[346,458,1598,1625]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" metricValueMax="4.0" metricValueMin="2.4" pageId="5" pageNumber="126" unit="cm" value="32.0" valueMax="40.0" valueMin="24.0">24–40 cm</quantity>
long, 
<quantity box="[530,641,1598,1625]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="5" pageNumber="126" unit="cm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 cm</quantity>
wide, acuminate apex, subcuneate or attenuate, and oblique at the base, almost not decurrent on the peduncle, yellowish-green at anthesis, dark green at post-anthesis; spadix 
<quantity box="[1277,1387,1634,1661]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="5" pageNumber="126" unit="cm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 cm</quantity>
long, 2.0– 
<quantity box="[186,262,1670,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="126" unit="cm" value="2.5">2.5 cm</quantity>
diam., cream-yellowish, emitting a sweet and pleasant scent at anthesis, stipe short, 
<quantity box="[1220,1314,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="5" pageNumber="126" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
long, 
<quantity box="[1386,1452,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="126" unit="mm" value="6.0">6 mm</quantity>
diam., yellowish-green at anthesis, yellowish-green at anthesis; perianth with 6 free tepals, 
<quantity box="[1142,1275,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="5" pageNumber="126" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
long; 6 anthers, 3.5–4.0 mm long, thecae oblong, 1.0– 
<quantity box="[564,652,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="126" unit="mm" value="1.8">1.8 mm</quantity>
; the pistils, sharply emergent, conic, 
<quantity box="[1070,1185,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="7.3" metricValueMin="5.0" pageId="5" pageNumber="126" unit="mm" value="6.15" valueMax="7.3" valueMin="5.0">5–7.3 mm</quantity>
long, style 
<quantity box="[1316,1452,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.1" metricValueMin="1.8" pageId="5" pageNumber="126" unit="mm" value="2.45" valueMax="3.1" valueMin="1.8">1.8–3.1 mm</quantity>
long, 
<quantity box="[198,332,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="126" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
diam. at the base; ovary 3-locular, 1–4 ovules per locule, 4–7 ovules per ovary. INFRUCTESCENCE 
<quantity box="[136,246,1814,1841]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="5" pageNumber="126" unit="cm" value="19.5" valueMax="21.0" valueMin="18.0">18–21 cm</quantity>
long, 
<quantity box="[316,440,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="5" pageNumber="126" unit="cm" value="2.9" valueMax="3.3" valueMin="2.5">2.5–3.3 cm</quantity>
diam.; berries obovoid to oblong, rostrate, 
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long, 3.3–6.0 mm wide, yellowish at maturity; seeds oblong to oblique-ovoid, dark to dark brown when dry and glossy when fresh.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="126" type="distribution">
<paragraph blockId="5.[136,1452,1202,2093]" pageId="5" pageNumber="126">
<emphasis bold="true" box="[189,742,1886,1912]" pageId="5" pageNumber="126">Distribution, habitat and conservation status:</emphasis>
— 
<taxonomicName box="[770,1086,1886,1912]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[770,1086,1886,1912]" italics="true" pageId="5" pageNumber="126">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
is only known from the Centro de Interpretación y Convivencia con la Naturaleza “Yumka”, 
<collectingRegion box="[832,923,1922,1948]" country="Mexico" name="Tabasco" pageId="5" pageNumber="126">Tabasco</collectingRegion>
(
<figureCitation box="[940,1006,1922,1948]" captionStart="FIGURE 1" captionStartId="1.[136,229,1335,1357]" captionTargetBox="[168,1421,439,1298]" captionTargetId="figure-153@1.[151,1436,425,1311]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the collection sites of Spathiphyllum maldonadianum (red dots) and S. wilfridianum (red triangle) in the state of Tabasco. The floral scent of S. maldonadianum was collected in situ in Pomoquita (bottom red dot) and of S. wilfridianum from a cultivated plant in Comalcalco (red star; cultivated individual)." figureDoi="http://doi.org/10.5281/zenodo.7103540" httpUri="https://zenodo.org/record/7103540/files/figure.png" pageId="5" pageNumber="126">Fig. 1</figureCitation>
). Only 
<specimenCount box="[1096,1216,1922,1948]" pageId="5" pageNumber="126" type="adult">three adult</specimenCount>
individuals and 
<specimenCount pageId="5" pageNumber="126" type="juvenile">four juveniles</specimenCount>
were found growing as an understory species in a flooded site in evergreen tropical forest, at 
<quantity box="[1293,1336,1959,1985]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="126" unit="m" value="7.0">7 m</quantity>
asl in this protected natural area. Due to the low number of individuals found, the introduced exotic fauna (such as the deer, 
<taxonomicName baseAuthorityName="Erxleben" baseAuthorityYear="1777" box="[136,239,2031,2057]" class="Mammalia" family="Cervidae" genus="Axis" kingdom="Animalia" order="Artiodactyla" pageId="5" pageNumber="126" phylum="Chordata" rank="species" species="axis">
<emphasis box="[136,239,2031,2057]" italics="true" pageId="5" pageNumber="126">Axis axis</emphasis>
</taxonomicName>
), which often destroys the herbaceous vegetation of the understory, and according to the IUCN Red List Categories and Criteria, the species can be considered as critically endangered [CR B2ab(ii,iii,iv,v); 
<bibRefCitation author="IUCN &amp; Cambridge, UK" box="[1259,1397,2066,2092]" pageId="5" pageNumber="126" refId="ref7937" refString="IUCN (2012) The IUCN Red List categories and criteria: Version 3.1, 2 nd ed. IUCN Species Survival Commission. Gland, Switzerland and Cambridge, UK. Available from: https: // portals. iucn. org / library / sites / library / files / documents / RL- 2001 - 001 - 2 nd-Es. pdf (accessed 22 March 2022)" type="book" year="2012">IUCN 2012</bibRefCitation>
].
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7103546" ID-Zenodo-Dep="7103546" httpUri="https://zenodo.org/record/7103546/files/figure.png" pageId="6" pageNumber="127" startId="6.[136,229,1421,1443]" targetBox="[346,1242,191,1385]" targetPageId="6">
<paragraph blockId="6.[136,1452,1421,1479]" pageId="6" pageNumber="127">
<emphasis bold="true" box="[136,255,1421,1443]" pageId="6" pageNumber="127">FIGURE 4.</emphasis>
<taxonomicName box="[262,532,1421,1442]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[262,532,1421,1442]" italics="true" pageId="6" pageNumber="127">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
. A. Cultivated adult plant. B. Adaxial view of the blade. C. Portion of petioles showing entire sheath margins (arrow). D. Inflorescence showing spathe and spadix in anthesis. E. Pistils. Photos by P. Díaz Jiménez.
</paragraph>
</caption>
<subSubSection pageId="6" pageNumber="127" type="biology_ecology">
<paragraph blockId="6.[136,1452,1526,2093]" pageId="6" pageNumber="127">
<emphasis bold="true" box="[189,519,1526,1552]" pageId="6" pageNumber="127">Phenology and floral scent:</emphasis>
—The specimens at anthesis and with infructescence were collected in November. Flowering in cultivated plants (plants cultivated since 2009 when they were still juveniles and removed with the complete rhizome) has been recorded in March, June, and November. The spadices of 
<taxonomicName box="[1094,1267,1598,1624]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[1094,1267,1598,1624]" italics="true" pageId="6" pageNumber="127">S. wilfridianum</emphasis>
</taxonomicName>
emitted a strong and pleasant floral scent, most intense between 6:00 and 12:00 AM. A total of 25 compounds was detected in the two samples (20 terpenoids, three aromatics and two lipid-derived compounds; Appendix 2). The main compounds (&gt; 5% in any sample) were (
<emphasis box="[375,392,1707,1732]" italics="true" pageId="6" pageNumber="127">E</emphasis>
)- 
<emphasis bold="true" box="[411,425,1705,1732]" italics="true" pageId="6" pageNumber="127">β</emphasis>
-Farnesene, (
<emphasis box="[570,587,1707,1732]" italics="true" pageId="6" pageNumber="127">E</emphasis>
)-Nerolidol, Methyl benzoate, 
<emphasis bold="true" box="[926,941,1706,1732]" italics="true" pageId="6" pageNumber="127">α</emphasis>
-Terpineol, and Eucalyptol (Appendix 2). (
<emphasis box="[1416,1433,1707,1732]" italics="true" pageId="6" pageNumber="127">E</emphasis>
)- Nerolidol is reported in this study for the first time as a main compound in 
<taxonomicName authorityName="Schott" authorityYear="1832" box="[980,1142,1742,1768]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="genus">
<emphasis box="[980,1142,1742,1768]" italics="true" pageId="6" pageNumber="127">Spathiphyllum</emphasis>
</taxonomicName>
(
<bibRefCitation author="Hentrich, H. &amp; Kaiser, R. &amp; Gottsberger, G." box="[1157,1387,1742,1768]" pageId="6" pageNumber="127" pagination="587 - 596" refId="ref7825" refString="Hentrich, H., Kaiser, R. &amp; Gottsberger, G. (2010) Floral biology and reproductive isolation by floral scent in three sympatric aroid species in French Guiana. Plant Biology 12: 587 - 596. https: // doi. org / 10.1111 / j. 1438 - 8677.2009.00256. x" type="journal article" year="2010">
Hentrich 
<emphasis box="[1264,1313,1742,1768]" italics="true" pageId="6" pageNumber="127">et al</emphasis>
. 2010
</bibRefCitation>
; 
<bibRefCitation author="Diaz Jimenez, P. &amp; Hentrich, H. &amp; Aguilar-Rodriguez, P. A. &amp; Kromer, T. &amp; Chartier, M. &amp; MacSwiney-G., M. C. &amp; Gibernau, M." pageId="6" pageNumber="127" pagination="83 - 104" refId="ref7293" refString="Diaz Jimenez, P., Hentrich, H., Aguilar-Rodriguez, P. A., Kromer, T., Chartier, M., MacSwiney-G., M. C. &amp; Gibernau, M. (2019) A review on the pollination of aroids with bisexual flowers. Annals of the Missouri Botanical Garden 104: 83 - 104. https: // doi. org / 10.3417 / 2018219" type="journal article" year="2019">
Díaz Jiménez 
<emphasis box="[235,284,1778,1804]" italics="true" pageId="6" pageNumber="127">et al</emphasis>
. 2019
</bibRefCitation>
; 
<bibRefCitation author="Diaz Jimenez, P. &amp; Hentrich, H. &amp; Dotterl, S. &amp; Kromer, T. &amp; MacSwiney-G., M. C. &amp; Aguilar-Rodriguez, P. A." box="[369,443,1778,1804]" pageId="6" pageNumber="127" pagination="1 - 12" refId="ref7520" refString="Diaz Jimenez, P., Hentrich, H., Dotterl, S., Kromer, T., MacSwiney-G., M. C. &amp; Aguilar-Rodriguez, P. A. (2021 b) Reproductive biology of two Spathiphyllum (Araceae) species in Los Tuxtlas, Veracruz, Mexico. Flora 285: 1 - 12. https: // doi. org / 10.1016 / j. flora. 2021.151958" type="journal article" year="2021">2021b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="127" type="etymology">
<paragraph blockId="6.[136,1452,1526,2093]" pageId="6" pageNumber="127">
<emphasis bold="true" box="[189,314,1814,1840]" pageId="6" pageNumber="127">Eponymy:</emphasis>
— 
<taxonomicName box="[342,662,1814,1840]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[342,662,1814,1840]" italics="true" pageId="6" pageNumber="127">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
is named in honor of the Mexican scientist, Dr. Wilfrido Miguel Contreras Sánchez, a Mexican professor at the Universidad Juárez Autónoma de 
<collectingRegion box="[1047,1138,1850,1876]" country="Mexico" name="Tabasco" pageId="6" pageNumber="127">Tabasco</collectingRegion>
(UJAT). During his time as Faculty Director at the Division of Biological Sciences (DACBiol), he aided and supported the studies of the 
<taxonomicName box="[1360,1452,1886,1912]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
in the state of 
<collectingRegion box="[306,400,1922,1948]" country="Mexico" name="Tabasco" pageId="6" pageNumber="127">Tabasco</collectingRegion>
, helping students to attend national and international conferences. Dr. Wilfrido is currently Secretary for Research, Graduate Studies, and Relations at the UJAT. His main line of research is fish reproduction and sexual inversion.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127" type="materials_examined">
<paragraph blockId="6.[136,1452,1526,2093]" lastBlockId="7.[136,1452,158,1372]" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127">
<materialsCitation collectingDate="2009-11-16" collectionCode="A" collectorName="Dos Montes &amp; la Naturaleza Yumka" country="Mexico" county="Centro de Interpretacion" elevation="7" lastPageId="7" lastPageNumber="128" location="Pedro Diaz Jimenez" municipality="Convivencia" pageId="6" pageNumber="127" specimenCount="2" stateProvince="Tabasco">
<emphasis bold="true" box="[189,707,2029,2056]" pageId="6" pageNumber="127">
<typeStatus box="[189,564,2030,2056]" isOtherTypeStatus="true" pageId="6" pageNumber="127">Additional specimens examined</typeStatus>
(
<typeStatus box="[577,699,2029,2055]" pageId="6" pageNumber="127">paratypes</typeStatus>
)
</emphasis>
:— 
<collectingCountry box="[740,856,2030,2056]" name="Mexico" pageId="6" pageNumber="127">MEXICO</collectingCountry>
. 
<collectingRegion box="[865,961,2030,2056]" country="Mexico" name="Tabasco" pageId="6" pageNumber="127">Tabasco</collectingRegion>
: 
<emphasis bold="true" box="[969,1122,2030,2056]" pageId="6" pageNumber="127">Mun. Centro</emphasis>
, ejido 
<collectorName box="[1196,1334,2030,2056]" pageId="6" pageNumber="127">Dos Montes</collectorName>
, 
<collectingCounty pageId="6" pageNumber="127">Centro de Interpretación</collectingCounty>
y 
<collectingMunicipality box="[320,462,2066,2092]" pageId="6" pageNumber="127">Convivencia</collectingMunicipality>
con 
<collectorName box="[516,751,2066,2092]" pageId="6" pageNumber="127">la Naturaleza Yumka</collectorName>
, 17°45’, 92°45’, 
<quantity box="[943,987,2066,2092]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="127" unit="m" value="7.0">
<elevation box="[943,987,2066,2092]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="127" unit="m" value="7.0">7 m</elevation>
</quantity>
, 
<date box="[999,1217,2066,2092]" pageId="6" pageNumber="127" value="2009-11-16">
<collectingDate box="[999,1217,2066,2092]" pageId="6" pageNumber="127" value="2009-11-16">16 November 2009</collectingDate>
</date>
, 
<emphasis italics="true" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127">
<location LSID="urn:lsid:plazi:treatment:4A018914FFD4FFA8A9AE73534B43FADA:C7776ED9FFD7FFA9ADEA782F4FCFF7AA" box="[1228,1451,2066,2092]" country="Mexico" county="Centro de Interpretacion" municipality="Convivencia" name="Pedro Diaz Jimenez" pageId="6" pageNumber="127" stateProvince="Tabasco">Pedro Díaz Jiménez</location>
&amp; 
<collectionCode box="[164,184,159,185]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="128" type="Herbarium">A</collectionCode>
</emphasis>
</materialsCitation>
<emphasis box="[184,353,158,185]" italics="true" pageId="7" pageNumber="128">. Garduza 1093</emphasis>
(MO!). 
<materialsCitation box="[446,690,158,185]" collectingDate="2009-11-16" collectionCode="R" collectorName="Dos Montes &amp; la Naturaleza Yumka" country="Mexico" county="Centro de Interpretacion" elevation="7" isTailEnumeration="true" location="Pedro Diaz Jimenez" municipality="Convivencia" pageId="7" pageNumber="128" specimenCount="1" stateProvince="Mun. Comalcalco">
<emphasis bold="true" box="[446,657,158,184]" pageId="7" pageNumber="128">
<collectingRegion box="[446,657,158,184]" pageId="7" pageNumber="128">Mun. Comalcalco</collectingRegion>
</emphasis>
, 
<collectionCode box="[670,690,158,184]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="128">R</collectionCode>
</materialsCitation>
/ 
<materialsCitation collectingDate="2021-03-09" collectionCode="UJAT" collectorName="Independencia &amp; Seccion" country="Mexico" elevation="8" latitude="18.283333" longLatPrecision="1275" longitude="-93.15" pageId="7" pageNumber="128" specimenCount="1" stateProvince="Mun. Comalcalco">
a. 
<collectorName box="[723,886,158,184]" pageId="7" pageNumber="128">Independencia</collectorName>
2da. 
<collectorName box="[946,1035,158,184]" pageId="7" pageNumber="128">Sección</collectorName>
(cultivated), 
<geoCoordinate box="[1183,1280,158,184]" degrees="18" direction="north" minutes="17" orientation="latitude" pageId="7" pageNumber="128" precision="925" value="18.283333">18º17’N</geoCoordinate>
, 
<geoCoordinate box="[1291,1391,158,184]" degrees="93" direction="west" minutes="09" orientation="longitude" pageId="7" pageNumber="128" precision="925" value="-93.15">93º09’W</geoCoordinate>
, 
<quantity box="[1402,1447,158,185]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="128" unit="m" value="8.0">
<elevation box="[1402,1447,158,185]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="128" unit="m" value="8.0">8 m</elevation>
</quantity>
, 
<date box="[136,284,194,221]" pageId="7" pageNumber="128" value="2021-03-09">
<collectingDate box="[136,284,194,221]" pageId="7" pageNumber="128" value="2021-03-09">09 Mar 2021</collectingDate>
</date>
, 
<emphasis box="[295,582,194,221]" italics="true" pageId="7" pageNumber="128">Pedro Díaz Jiménez 1496</emphasis>
(
<collectionCode box="[597,674,194,220]" country="Mexico" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14347" lsid="urn:lsid:biocol.org:col:14347" name="Universidad Juárez Autónoma de Tabasco" pageId="7" pageNumber="128" type="Herbarium">UJAT</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="128" type="discussion">
<paragraph blockId="7.[136,1452,158,1372]" pageId="7" pageNumber="128">
<emphasis bold="true" box="[189,265,230,256]" pageId="7" pageNumber="128">Notes:</emphasis>
— 
<taxonomicName box="[294,615,230,256]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[294,615,230,256]" italics="true" pageId="7" pageNumber="128">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
had been mistakenly identified as 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[1049,1286,230,257]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[1049,1286,230,257]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
. However, 
<taxonomicName class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis italics="true" pageId="7" pageNumber="128">S. wilfridianum</emphasis>
</taxonomicName>
has a petiole almost twice as long as the blade and is sheathed almost to the middle, the margin of the sheath is entire and not wrinkled, its blades are narrower with an attenuated base (rarely sub-rounded), the angle of the primary veins is less than 60°, and the number of ovules per ovary is less than that recorded in 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[1221,1452,338,365]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[1221,1452,338,365]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
(
<bibRefCitation author="Bunting, G. S." box="[144,303,374,400]" pageId="7" pageNumber="128" pagination="1 - 53" refId="ref6812" refString="Bunting, G. S. (1960) A revision of Spathiphyllum (Araceae). Memoirs of the New York Botanical Garden 10: 1 - 53." type="journal article" year="1960">Bunting 1960</bibRefCitation>
, 
<bibRefCitation author="Diaz Jimenez, P. &amp; Perez-Farrera, M. A. &amp; Hentrich, H. &amp; Gomez-Dominguez, H. &amp; Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A." box="[316,618,374,401]" pageId="7" pageNumber="128" refId="ref7447" refString="Diaz Jimenez, P., Perez-Farrera, M. A., Hentrich, H., Gomez-Dominguez, H., Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A. (2021 a) Mistaken identity: a new Spathiphyllum (Araceae) from Veracruz, Mexico. Phytotaxa 522: 056 - 062. https: // doi. org / 10.11646 / phytotaxa. 522.1.6" type="journal volume" year="2021">
Díaz Jiménez 
<emphasis box="[479,530,374,400]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. 2021a
</bibRefCitation>
). 
<taxonomicName box="[639,954,374,400]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[639,954,374,400]" italics="true" pageId="7" pageNumber="128">Spathiphyllum wilfridianum</emphasis>
</taxonomicName>
is a rare species, only known from its 
<typeStatus box="[1403,1451,375,401]" pageId="7" pageNumber="128">type</typeStatus>
locality, growing below 
<quantity box="[406,462,410,437]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="128" unit="m" value="10.0">10 m</quantity>
asl in tropical forests, while 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[782,1010,410,437]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[782,1010,410,437]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
has been collected mostly above 
<quantity box="[1381,1452,410,437]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="128" unit="m" value="800.0">800 m</quantity>
asl in coffee crops and montane cloud forests (
<bibRefCitation author="Diaz Jimenez, P. &amp; Perez-Farrera, M. A. &amp; Hentrich, H. &amp; Gomez-Dominguez, H. &amp; Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A." box="[662,960,446,473]" pageId="7" pageNumber="128" refId="ref7447" refString="Diaz Jimenez, P., Perez-Farrera, M. A., Hentrich, H., Gomez-Dominguez, H., Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A. (2021 a) Mistaken identity: a new Spathiphyllum (Araceae) from Veracruz, Mexico. Phytotaxa 522: 056 - 062. https: // doi. org / 10.11646 / phytotaxa. 522.1.6" type="journal volume" year="2021">
Díaz Jiménez 
<emphasis box="[823,873,446,472]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. 2021a
</bibRefCitation>
). The new species represents the fourteenth species for 
<collectingCountry box="[266,352,482,508]" name="Mexico" pageId="7" pageNumber="128">Mexico</collectingCountry>
and the third species for 
<collectingRegion box="[643,734,482,508]" country="Mexico" name="Tabasco" pageId="7" pageNumber="128">Tabasco</collectingRegion>
(
<bibRefCitation author="Diaz Jimenez, P. &amp; Perez-Farrera, M. A. &amp; Hentrich, H. &amp; Gomez-Dominguez, H. &amp; Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A." box="[750,1050,482,509]" pageId="7" pageNumber="128" refId="ref7447" refString="Diaz Jimenez, P., Perez-Farrera, M. A., Hentrich, H., Gomez-Dominguez, H., Ameca-Juarez, E. O. &amp; Aguilar-Rodriguez, P. A. (2021 a) Mistaken identity: a new Spathiphyllum (Araceae) from Veracruz, Mexico. Phytotaxa 522: 056 - 062. https: // doi. org / 10.11646 / phytotaxa. 522.1.6" type="journal volume" year="2021">
Díaz Jiménez 
<emphasis box="[913,964,482,508]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. 2021a
</bibRefCitation>
, Croat 
<emphasis box="[1133,1184,482,508]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. unpubl.). On the other hand, unlike 
<taxonomicName box="[282,493,518,545]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="maldonadianum">
<emphasis box="[282,493,518,545]" italics="true" pageId="7" pageNumber="128">S. maldonadianum</emphasis>
</taxonomicName>
, 
<taxonomicName box="[507,681,518,545]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="wilfridianum">
<emphasis box="[507,681,518,545]" italics="true" pageId="7" pageNumber="128">S. wilfridianum</emphasis>
</taxonomicName>
has entire sheath margins, shortest geniculum, narrower blades with attenuated base, fewer pairs of primary veins per side, shortest pistil, and fewer ovules per ovary.
</paragraph>
<paragraph blockId="7.[136,1452,158,1372]" pageId="7" pageNumber="128">
We strongly recommend that all specimens identified under 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[884,1116,590,617]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[884,1116,590,617]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
in 
<collectingCountry box="[1157,1243,590,616]" name="Mexico" pageId="7" pageNumber="128">Mexico</collectingCountry>
require a detailed review, being revealed as a complex of several species morphologically similar, but with distinct floral scent composition. Initially, 
<bibRefCitation author="Liebmann, F. M." box="[235,427,662,688]" pageId="7" pageNumber="128" pagination="1 - 120" refId="ref8006" refString="Liebmann, F. M. (1849) Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn (1 - 2): 1 - 120." type="book chapter" year="1849">Liebmann (1849)</bibRefCitation>
published 
<taxonomicName authority="Liebmann (1849: 24)" authorityName="Liebmann" authorityPageNumber="24" authorityYear="1849" box="[545,1163,662,688]" class="Liliopsida" family="Araceae" genus="Hydnostachyon" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[545,925,662,688]" italics="true" pageId="7" pageNumber="128">Hydnostachyon cochlearispathum</emphasis>
<bibRefCitation author="Liebmann, F. M." box="[930,1163,662,688]" pageId="7" pageNumber="128" pagination="1 - 120" refId="ref8006" refString="Liebmann, F. M. (1849) Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KjObenhavn (1 - 2): 1 - 120." type="book chapter" year="1849">Liebmann (1849: 24)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Liebm. (1849: 24)" authorityName="Liebm." authorityPageNumber="24" authorityYear="1849" class="Liliopsida" family="Araceae" genus="Hydnostachyon" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="longirostre">
<emphasis box="[1212,1366,662,688]" italics="true" pageId="7" pageNumber="128">H. longirostre</emphasis>
Liebm. (1849: 24)
</taxonomicName>
, and 
<bibRefCitation author="Schott, H. W." box="[319,475,698,725]" pageId="7" pageNumber="128" refId="ref8556" refString="Schott, H. W. (1853) Aroideae 1. Gerold, Vienna, 2. https: // doi. org / 10.5962 / bhl. title. 118627" type="book" year="1853">Schott (1853)</bibRefCitation>
, transferred them to 
<taxonomicName authorityName="Schott" authorityYear="1832" box="[713,875,698,724]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="genus">
<emphasis box="[713,875,698,724]" italics="true" pageId="7" pageNumber="128">Spathiphyllum</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Engler, A." box="[890,1047,698,724]" pageId="7" pageNumber="128" pagination="1 - 681" refId="ref7597" refString="Engler, A. (1879) Araceae. In: De Candolle, A. &amp; De Candolle, C. (eds.) Monographiae Phanoerogamorum 2: 1 - 681." type="journal article" year="1879">Engler (1879)</bibRefCitation>
published 
<taxonomicName authorityName="Liebmann" authorityYear="1849" box="[1173,1403,698,725]" class="Liliopsida" family="Araceae" genus="Hydnostachyon" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[1173,1403,698,725]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
and 
<taxonomicName authority="(Liebm.) Engler (1879: 221)" authorityName="Engler" authorityPageNumber="221" authorityYear="1879" baseAuthorityName="Liebm." box="[136,869,734,761]" class="Liliopsida" family="Araceae" genus="Hydnostachyon" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="variety" species="cochlearispathum" variety="longirostre">
<emphasis box="[136,365,734,761]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
var. 
<emphasis box="[419,541,734,760]" italics="true" pageId="7" pageNumber="128">longirostre</emphasis>
(Liebm.) 
<bibRefCitation author="Engler, A." box="[653,869,734,760]" pageId="7" pageNumber="128" pagination="1 - 681" refId="ref7597" refString="Engler, A. (1879) Araceae. In: De Candolle, A. &amp; De Candolle, C. (eds.) Monographiae Phanoerogamorum 2: 1 - 681." type="journal article" year="1879">Engler (1879: 221)</bibRefCitation>
</taxonomicName>
. 
<taxonomicName box="[879,1425,734,761]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="variety" species="cochlearispathum" variety="longirostre">
<emphasis box="[879,1249,734,760]" italics="true" pageId="7" pageNumber="128">Spathiphyllum cochlearispathum</emphasis>
var. 
<emphasis box="[1303,1425,734,760]" italics="true" pageId="7" pageNumber="128">longirostre</emphasis>
</taxonomicName>
, a taxon that is currently a synonym of 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[545,774,770,797]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[545,774,770,797]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
, has similar characteristics to the latter species as referred to by 
<bibRefCitation author="Bunting, G. S." box="[172,348,806,833]" pageId="7" pageNumber="128" pagination="291 - 382" refId="ref6842" refString="Bunting, G. S. (1965) Commentary on Mexican Araceae. Gentes Herbarum 9: 291 - 382." type="journal article" year="1965">Bunting (1965)</bibRefCitation>
. However, 
<taxonomicName authorityName="Lundell" authorityYear="1941" box="[476,594,806,833]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="lacustre">
<emphasis box="[476,594,806,833]" italics="true" pageId="7" pageNumber="128">S. lacustre</emphasis>
</taxonomicName>
, also considered a synonym of 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[953,1183,806,833]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[953,1183,806,833]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Bunting, G. S." box="[1226,1401,806,833]" pageId="7" pageNumber="128" pagination="291 - 382" refId="ref6842" refString="Bunting, G. S. (1965) Commentary on Mexican Araceae. Gentes Herbarum 9: 291 - 382." type="journal article" year="1965">Bunting (1965)</bibRefCitation>
, has characteristics that separate it from 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[539,768,842,869]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[539,768,842,869]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
, therefore, it is considered a different species (Díaz Jiménez 
<emphasis box="[136,185,878,904]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. unpubl.).
</paragraph>
<paragraph blockId="7.[136,1452,158,1372]" pageId="7" pageNumber="128">
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[189,557,914,940]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[189,557,914,940]" italics="true" pageId="7" pageNumber="128">Spathiphyllum cochlearispathum</emphasis>
</taxonomicName>
was formerly considered a species with a wide morphological variation, growing from sea level to 
<quantity box="[339,426,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="128" unit="m" value="1600.0">1600 m</quantity>
in different 
<typeStatus box="[571,630,951,977]" pageId="7" pageNumber="128">types</typeStatus>
of vegetation in the south of 
<collectingCountry box="[974,1062,950,976]" name="Mexico" pageId="7" pageNumber="128">Mexico</collectingCountry>
, in the states of 
<collectingRegion box="[1254,1347,950,976]" country="Mexico" name="Chiapas" pageId="7" pageNumber="128">Chiapas</collectingRegion>
, 
<collectingRegion box="[1361,1448,950,976]" country="Mexico" name="Oaxaca" pageId="7" pageNumber="128">Oaxaca</collectingRegion>
, 
<collectingRegion box="[136,230,986,1012]" country="Mexico" name="Tabasco" pageId="7" pageNumber="128">Tabasco</collectingRegion>
, 
<collectingRegion box="[242,345,987,1013]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="7" pageNumber="128">Veracruz</collectingRegion>
, and 
<collectingRegion box="[404,494,986,1012]" country="Mexico" name="Yucatan" pageId="7" pageNumber="128">Yucatán</collectingRegion>
(
<bibRefCitation author="Croat, T. B. &amp; Acebey, A." box="[509,763,986,1013]" pageId="7" pageNumber="128" refId="ref7055" refString="Croat, T. B. &amp; Acebey, A. (2015) Araceae. In: Flora de Veracruz. Fasciculo 164. Instituto de Ecologia A, C., Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa, 211 pp." type="book" year="2015">Croat &amp; Acebey 2015</bibRefCitation>
), and the only species reported for 
<collectingRegion box="[1162,1253,986,1012]" country="Mexico" name="Tabasco" pageId="7" pageNumber="128">Tabasco</collectingRegion>
(
<bibRefCitation author="Diaz Jimenez, P. &amp; Guadarrama-Olivera, M. A. &amp; Croat, T. B." pageId="7" pageNumber="128" pagination="131 - 142" refId="ref7239" refString="Diaz Jimenez, P., Guadarrama-Olivera, M. A. &amp; Croat, T. B. (2015) Diversidad floristica de Araceae en el estado de Tabasco, Mexico. Botanical Sciences 93: 131 - 142. https: // doi. org / 10.17129 / botsci. 238" type="journal article" year="2015">
Díaz Jiménez 
<emphasis italics="true" pageId="7" pageNumber="128">et al</emphasis>
. 2015
</bibRefCitation>
), but our detailed review of herbarium specimens and the collection of specimens from different populations, revealed that none of the studied specimens corresponds to 
<taxonomicName baseAuthorityName="Bunting" baseAuthorityYear="1960" box="[815,1046,1058,1085]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cochlearispathum">
<emphasis box="[815,1046,1058,1085]" italics="true" pageId="7" pageNumber="128">S. cochlearispathum</emphasis>
</taxonomicName>
. Therefore, its presence in the state of 
<collectingRegion box="[168,259,1094,1120]" country="Mexico" name="Tabasco" pageId="7" pageNumber="128">Tabasco</collectingRegion>
is currently doubtful (Díaz Jiménez 
<emphasis box="[683,734,1094,1120]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. unpubl.). So far, the use of floral scent composition has been scarcely used to distinguish botanical species but is becoming more used in recent years to separate species (
<bibRefCitation author="Peakall, R. &amp; Whitehead, M. R." pageId="7" pageNumber="128" pagination="341 - 355" refId="ref8201" refString="Peakall, R. &amp; Whitehead, M. R. (2014) Floral odour chemistry defines species boundaries and underpins strong reproductive isolation in sexually deceptive orchids. Annals of Botany 113: 341 - 355. https: // doi. org / 10.1093 / aob / mct 199" type="journal article" year="2014">Peakall &amp; Whitehead 2014</bibRefCitation>
, 
<bibRefCitation author="Chen, G. &amp; Gong, W. - G. &amp; Ge, J. &amp; Schinnerl, J. &amp; Wang, B. &amp; Sun, W. - B." box="[361,548,1166,1192]" pageId="7" pageNumber="128" pagination="825 - 839" refId="ref6894" refString="Chen, G., Gong, W. - G., Ge, J., Schinnerl, J., Wang, B. &amp; Sun, W. - B. (2017) Variation in floral characters, particularly floral scent, in sapromyophilous Stemona species. Journal of Integrative Plant Biology 59: 825 - 839. https: // doi. org / 10.1111 / jipb. 12580" type="journal article" year="2017">
Chen 
<emphasis box="[427,483,1166,1192]" italics="true" pageId="7" pageNumber="128">et al.</emphasis>
2017
</bibRefCitation>
, 
<bibRefCitation author="Gervasi, D. D. L. &amp; Selosse, M-A. &amp; Sauve, M. &amp; Francke, W. &amp; Vereecken, N. J. &amp; Cozzolino, S. &amp; Schiestl, F. P." box="[559,774,1166,1192]" pageId="7" pageNumber="128" pagination="6023 - 6034" refId="ref7662" refString="Gervasi, D. D. L., Selosse, M-A., Sauve, M., Francke, W., Vereecken, N. J., Cozzolino, S. &amp; Schiestl, F. P. (2017) Floral scent and species divergence in a pair of sexually deceptive orchids. Ecology and Evolution 7: 6023 - 6034. https: // doi. org / 10.1002 / ece 3.3147" type="journal article" year="2017">
Gervasi 
<emphasis box="[653,708,1166,1192]" italics="true" pageId="7" pageNumber="128">et al.</emphasis>
2017
</bibRefCitation>
, 
<bibRefCitation author="Powers, J. M. &amp; Seco, R. &amp; Faiola, C. L. &amp; Sakai, A. K. &amp; Weller, S. G. &amp; Campbell, D. R. &amp; Guenther, A." box="[784,995,1166,1193]" pageId="7" pageNumber="128" pagination="1 - 16" refId="ref8414" refString="Powers, J. M., Seco, R., Faiola, C. L., Sakai, A. K., Weller, S. G., Campbell, D. R. &amp; Guenther, A. (2020) Floral scent composition and finescale timing in two moth-pollinated hawaiian Schiedea (Caryophyllaceae). Frontiers in Plant Science 11: 1 - 16. https: // doi. org / 10.3389 / fpls. 2020.01116" type="journal article" year="2020">
Powers 
<emphasis box="[873,929,1166,1192]" italics="true" pageId="7" pageNumber="128">et al.</emphasis>
2020
</bibRefCitation>
), even in sympatric and morphologically similar species (
<bibRefCitation author="Okamoto, T. &amp; Su, Z. - H." box="[319,562,1202,1228]" pageId="7" pageNumber="128" pagination="1 - 12" refId="ref8136" refString="Okamoto, T. &amp; Su, Z. - H. (2021) Chemical analysis of floral scents in sympatric Ficus species: highlighting different compositions of floral scents in morphologically and phylogenetically close species. Plant Systematics and Evolution 307: 1 - 12. https: // doi. org / 10.1007 / s 00606 - 021 - 01767 - y" type="journal article" year="2021">Okamoto &amp; Su 2021</bibRefCitation>
). These new species of 
<taxonomicName authorityName="Schott" authorityYear="1832" box="[833,995,1202,1228]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="genus">
<emphasis box="[833,995,1202,1228]" italics="true" pageId="7" pageNumber="128">Spathiphyllum</emphasis>
</taxonomicName>
have different odors, and whether these differences have any impact on their pollinator´s spectrum is still to be determined (see for example 
<bibRefCitation author="Diaz Jimenez, P. &amp; Hentrich, H. &amp; Aguilar-Rodriguez, P. A. &amp; Kromer, T. &amp; Chartier, M. &amp; MacSwiney-G., M. C. &amp; Gibernau, M." pageId="7" pageNumber="128" pagination="83 - 104" refId="ref7293" refString="Diaz Jimenez, P., Hentrich, H., Aguilar-Rodriguez, P. A., Kromer, T., Chartier, M., MacSwiney-G., M. C. &amp; Gibernau, M. (2019) A review on the pollination of aroids with bisexual flowers. Annals of the Missouri Botanical Garden 104: 83 - 104. https: // doi. org / 10.3417 / 2018219" type="journal article" year="2019">
Díaz Jiménez 
<emphasis box="[136,187,1274,1300]" italics="true" pageId="7" pageNumber="128">et al</emphasis>
. 2019
</bibRefCitation>
). In this sense, at least with scented aroids, we consider it important to, when possible, reinforce species description with ecological data such as the volatile composition of floral scents and pollinators, especially for closely related aroids.
</paragraph>
</subSubSection>
</treatment>
</document>