502 lines
49 KiB
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502 lines
49 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1009.54303" ID-GBIF-Dataset="4f67a727-4d9e-47e7-9c0a-6cf024d77da5" ID-PMC="PMC7801368" ID-Pensoft-Pub="1313-2970-1009-81" ID-Pensoft-UUID="24DEEA8C6B435153B6A5794E603BB79A" ID-PubMed="33505196" ID-ZooBank="F0C4C58B5F484CDBA443E258D732AC5B" ModsDocID="1313-2970-1009-81" checkinTime="1609809790016" checkinUser="pensoft" docAuthor="Klotz, Werner, Rintelen, Thomas von, Wowor, Daisy, Lukhaup, Chris & Rintelen, Kristina von" docDate="2021" docId="2A9F9EBD609958BCB9A4CB1614FA4B24" docLanguage="en" docName="ZooKeys 1009: 81-122" docOrigin="ZooKeys 1009" docPubDate="2021-01-04" docSource="http://dx.doi.org/10.3897/zookeys.1009.54303" docTitle="Caridina marlenae Klotz, Wowor & K. von Rintelen 2021, sp. nov." docType="treatment" docUuid="801EC24A-93F5-48EF-9394-AF0930D50E36" docUuidSource="ZooBank" docVersion="4" id="24DEEA8C6B435153B6A5794E603BB79A" lastPageNumber="81" masterDocId="24DEEA8C6B435153B6A5794E603BB79A" masterDocTitle="Lake Poso's shrimp fauna revisited: the description of five new species of the genus Caridina (Crustacea, Decapoda, Atyidae) more than doubles the number of endemic lacustrine species" masterLastPageNumber="122" masterPageNumber="81" pageNumber="81" updateTime="1668149687226" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Lake Poso's shrimp fauna revisited: the description of five new species of the genus Caridina (Crustacea, Decapoda, Atyidae) more than doubles the number of endemic lacustrine species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klotz, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1104-5613</mods:nameIdentifier>
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<mods:affiliation>Wiesenweg 1, A- 6063 Rum, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">wklotz@aon.at</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rintelen, Thomas von</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6253-3078</mods:nameIdentifier>
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<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, D- 10115 Berlin, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wowor, Daisy</mods:namePart>
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<mods:affiliation>Division of Zoology, Research Center for Biology, Indonesian Institute of Sciences (LIPI), Jalan Raya Jakarta Bogor Km 46, Cibinong 16911, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lukhaup, Chris</mods:namePart>
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<mods:affiliation>Waldstrasse 5 a, D- 66999 Hinterweidenthal, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rintelen, Kristina von</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4167-3570</mods:nameIdentifier>
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<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, D- 10115 Berlin, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-01-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1009</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>81</mods:start>
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<mods:end>122</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1009.54303</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1009.54303</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1009-81</mods:identifier>
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<mods:identifier type="ZooBank">F0C4C58B5F484CDBA443E258D732AC5B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">24DEEA8C6B435153B6A5794E603BB79A</mods:identifier>
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<treatment ID-GBIF-Taxon="176432961" LSID="urn:lsid:zoobank.org:act:801EC24A-93F5-48EF-9394-AF0930D50E36" httpUri="http://treatment.plazi.org/id/2A9F9EBD609958BCB9A4CB1614FA4B24" lastPageNumber="81" pageId="0" pageNumber="81">
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<subSubSection pageId="0" pageNumber="81" type="nomenclature">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName LSID="http://zoobank.org/801EC24A-93F5-48EF-9394-AF0930D50E36" authority="Klotz, Wowor & K. von Rintelen" authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae" status="sp. nov.">Caridina marlenae Klotz, Wowor & K. von Rintelen</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living specimens of Caridina spp in Lake Poso 2. A, B Caridina mayamareenae sp. nov. C C. lilianae sp. nov. D Two snail species Celetaia persculpta and Tylomelania sp. on soft substrate. Empty shells of these species are shelter for C. mayamareenae sp. nov. E, F C. poso sp. nov. G C. marlenae sp. nov. H C. fusca sp. nov. Not to scale. All photographs: W. Klotz." figureDoi="10.3897/zookeys.1009.54303.figure3" httpUri="https://binary.pensoft.net/fig/493860" pageId="0" pageNumber="81">Figures 3G</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="materials_examined">
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<paragraph pageId="0" pageNumber="81">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype</emphasis>
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: ♀ cl. 2.8 mm (MZB Cru 5039), Indonesia, Central Sulawesi, Lake Poso, E shore, S of Tentena, dive at small cape, 15 m,
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<geoCoordinate degrees="1" direction="south" minutes="46.394" orientation="latitude" precision="1" value="-1.7732334">1°46.394'S</geoCoordinate>
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,
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<geoCoordinate degrees="120" direction="east" minutes="38.327" orientation="longitude" precision="1" value="120.63879">120°38.327'E</geoCoordinate>
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, T. von Rintelen and W. Klotz leg., 12 May 2017.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Paratypes</emphasis>
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: 4 ♀♀ cl. 1.4-2.1 mm, 1 ♂ cl. 1.6 mm (MZB Cru 5040), 1 ♀ cl. 2.3 mm, 3 ♂♂ cl. 2.0-2.2 mm (ZMB 30199), same data as holotype; 2 ov. ♀♀ cl. 2.7 and 2.8 mm, 2 ♂♂ cl. 2.5 and 2.8 mm (MZB Cru 5041), 2 ♀♀ cl. 2.6 and 3.1 mm, 1 ♂ cl. 2.4 mm, 2 sequenced specimens without anterior pleopods cl. 2.2 and 2.9 mm (ZMB 29519), Lake Poso, E shore, S of Tentena, dive at small cape, in 15 m depth,
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<geoCoordinate degrees="1" direction="south" minutes="46.394" orientation="latitude" precision="1" value="-1.7732334">1°46.394'S</geoCoordinate>
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,
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<geoCoordinate degrees="120" direction="east" minutes="38.327" orientation="longitude" precision="1" value="120.63879">120°38.327'E</geoCoordinate>
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, M. Glaubrecht and T. von Rintelen leg., 16 May 2007; 2 ♀♀ cl. 2.1 and 2.5 mm, 1 ♂ cl. 2.0 mm, 2 juv. specimens (MZB Cru 5092), 2 ♀♀ cl. 1.9 and 2.2 mm, 1 ♂ cl. 1.8 mm, 1 juv. specimen (ZMB 31661), Lake Poso, E shore, S of Tentena, dive at small cape, in 15 m depth,
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<geoCoordinate degrees="1" direction="south" minutes="46.394" orientation="latitude" precision="1" value="-1.7732334">1°46.394'S</geoCoordinate>
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,
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<geoCoordinate degrees="120" direction="east" minutes="38.327" orientation="longitude" precision="1" value="120.63879">120°38.327'E</geoCoordinate>
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, T. von Rintelen leg., 14 Jul 2019.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="materials_examined">
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<paragraph pageId="0" pageNumber="81">Comparative material examined.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName authorityName="Schenkel" authorityYear="1902" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina sarasinorum" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="sarasinorum">
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<emphasis italics="true" pageId="0" pageNumber="81">Caridina sarasinorum</emphasis>
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</taxonomicName>
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Schenkel, 1902, 1 ov. ♀ cl. 3.0 mm, 1 ♂ cl. 2.6 mm (ZMB 29288), Lake Poso, E shore,
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<geoCoordinate degrees="2" direction="south" minutes="0.825" orientation="latitude" precision="1" value="-2.01375">2°0.825'S</geoCoordinate>
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,
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<geoCoordinate degrees="120" direction="east" minutes="42.007" orientation="longitude" precision="1" value="120.70012">120°42.007'E</geoCoordinate>
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, K. Zitzler leg., 16 Aug. 2004; 2 ov. ♀♀ cl. 3.2 and 3.6 mm, 1 ♀ cl. 3.mm, 5 ♂♂ cl. 2.1-2.6 mm, 2 juv. specimens cl. 1.7 mm, 1 damaged specimen cl. 2.0 mm (ZMB 30224), Lake Poso, E shore, beach in front of Dolidi Ndano Cottages,
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<geoCoordinate degrees="1" direction="south" minutes="48.14" orientation="latitude" precision="9" value="-1.8023334">1°48.14'S</geoCoordinate>
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,
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<geoCoordinate degrees="120" direction="east" minutes="38.043" orientation="longitude" precision="1" value="120.63405">120°38.043'E</geoCoordinate>
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, W. Klotz leg., 12 May 2017.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="description">
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<paragraph pageId="0" pageNumber="81">Description.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" pageId="0" pageNumber="81">Cephalothorax and cephalic appendages.</emphasis>
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Postorbital carapace length 1.44-3.07 mm (n = 19). Rostrum (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9A, B</figureCitation>
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) long and slender, curved upwards or sigmoid, reaching slightly beyond end of scaphocerite or in small specimen, to end of the antennular peduncle, distal 0.16-0.40 (median 0.38, n = 11) of dorsal margin unarmed, ventral margin armed throughout, dorsal teeth more widely spaced distally, 0.95-1.54 (median 1.20, n = 14) times as long as carapace, rostral formula 3-6 (4-6) + 10-20 / 6-18. Antennal spine slightly separated from orbital margin. Pterygostomial angle broadly rounded. Eyes well developed with globular cornea. Antennular peduncle (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9A, B, G</figureCitation>
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), 0.89-1.04 (median 0.96, n = 5) times as long as carapace, first segment 1.56-1.79 (median 1.71, n = 4) times as long as second segment, second segment 2.40-2.80 (median 2.58, n = 4) times longer than third segment. Tooth on distolateral margin of first segment of antennular peduncle acute. Stylocerite reaching to 0.78-0.83 (median 0.80, n = 4) of first segment of antennular peduncle. Scaphocerite (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9H</figureCitation>
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) 4.10-4.25 (median 4.20, n = 3) times as long as wide.
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</paragraph>
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<caption doi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81" start="Figure 9" startId="F9">
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" pageId="0" pageNumber="81">Figure 9.</emphasis>
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<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
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<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
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</taxonomicName>
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sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199
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<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
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cephalothorax and cephalic appendages
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<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
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rostrum
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<emphasis bold="true" pageId="0" pageNumber="81">C</emphasis>
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preanal carina
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<emphasis bold="true" pageId="0" pageNumber="81">D</emphasis>
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uropodal diaeresis
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<emphasis bold="true" pageId="0" pageNumber="81">E, F</emphasis>
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telson
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<emphasis bold="true" pageId="0" pageNumber="81">G</emphasis>
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antennular peduncle
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<emphasis bold="true" pageId="0" pageNumber="81">H</emphasis>
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scaphocerite
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<emphasis bold="true" pageId="0" pageNumber="81">I</emphasis>
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mandible
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<emphasis bold="true" pageId="0" pageNumber="81">K</emphasis>
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maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199
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<emphasis bold="true" pageId="0" pageNumber="81">J</emphasis>
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maxillula. Scale bars: 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="81">A-E</emphasis>
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); 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="81">F-K</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" pageId="0" pageNumber="81">
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<emphasis italics="true" pageId="0" pageNumber="81">Abdominal somites, telson and uropods</emphasis>
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.
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</emphasis>
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Sixth abdominal somite 0.55-0.64 (median 0.62, n = 5) times carapace length, 1.93-2.23 (median 2.15, n = 5) times as long as fifth somite, 1.00-1.13 (median 1.12, n = 4) times as long as telson. Distal margin of telson (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9E, F</figureCitation>
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) convex or subtriangular without a median projection, with 3-5 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; distal end with 7-10 long spiniform setae, lateral pair slightly longer than others, innermost pair very tiny. Preanal carina (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9C</figureCitation>
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) with a hook-like spine. Uropodal diaeresis (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9D</figureCitation>
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) with 11-14 stout movable spiniform setae, outermost ones shorter than lateral angle.
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</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" pageId="0" pageNumber="81">
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<emphasis italics="true" pageId="0" pageNumber="81">Mouthparts and branchiae</emphasis>
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.
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</emphasis>
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Incisor process of mandible (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9I</figureCitation>
|
||
) ending in irregular teeth, molar process truncated. Lower lacinia of maxillula (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9J</figureCitation>
|
||
) broadly rounded, upper lacinia elongate, with numerous distinct teeth on inner margin, palp slender with few pappose setae and one conical spiniform seta near tip. Upper endites of maxilla (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Caridina marlenae sp. nov. Morphology 1. Paratype ♂, cl. 2.2 mm, ZMB 30199 A cephalothorax and cephalic appendages B rostrum C preanal carina D uropodal diaeresis E, F telson G antennular peduncle H scaphocerite I mandible K maxilla; paratype ♀, cl. 2.3 mm, ZMB 30199 J maxillula. Scale bars: 1 mm (A-E); 0.5 mm (F-K)." figureDoi="10.3897/zookeys.1009.54303.figure9" httpUri="https://binary.pensoft.net/fig/493866" pageId="0" pageNumber="81">9K</figureCitation>
|
||
) subdivided, palp slender, scaphognathite tapering posteriorly, fringed with long, curved setae at posterior margin. Palp of first maxilliped (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10A, B</figureCitation>
|
||
) ending in triangular shape. Podobranch on second maxilliped (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10C</figureCitation>
|
||
) reduced to a lamina. Third maxilliped (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10D</figureCitation>
|
||
) with one well developed and one small arthrobranch, ultimate segment of maxilliped shorter than penultimate segment. First pereiopod with an arthrobranch. Pleurobranchs present on all pereiopods. Epipod slightly reduced (without distal hook) on third maxilliped, absent from all pereiopods.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">
|
||
<taxonomicName genus="Pereiopods" lsidName="Pereiopods" pageId="0" pageNumber="81" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Pereiopods</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Chelae of first and second pereiopod (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10E, F</figureCitation>
|
||
) well developed; chela of first pereiopod 3.00-3.83 (median 3.20, n = 5) times as long as wide, 1.00-1.05 (median 1.03, n = 5) times length of carpus; tips of fingers (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10E</figureCitation>
|
||
) rounded, without hooks, with tufts of hairs near tip; dactylus 1.88-2.50 (median 2.00, n = 5) times as long as palm; carpus slender, hardly excavated distally, 3.85-4.77 (median 4.00, n = 5) times as long as wide, 1.25-1.33 (median 1.29, n = 5) times length of merus. Merus 3.80-4.25 (median 4.00, n = 5) times as long as wide, 0.89-0.95 (median 0.92, n = 4) times as long as ischium. Chela of second pereiopod 3.69-4.38 (median 3.71, n = 5) times as long as wide, 0.68-0.79 (median 0.71, n = 6) times length of carpus; tips of fingers rounded, without hooks, with tufts of hairs near tip; dactylus 1.80-2.25 (median 2.00, n = 5) times as long as palm; carpus 6.78-9.80 (median 7.60, n = 5) times as long as wide, 1.27-1.58 (median 1.47, n = 4) times as long as merus; merus 5.33-6.50 (median 6.00, n = 4) times as long as wide, 0.82-0.93 (median 0.86, n = 4) times as long as ischium. Third pereiopod (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10G, H</figureCitation>
|
||
) not sexually dimorphic, dactylus 4.40-5.33 (median 4.50, n = 5) times as long as wide (terminal claw and spiniform setae on flexor margin included), terminating in one large claw with four or five short spiniform setae on flexor margin; propodus slender, 13.67-16.25 (median 14.57, n = 5) times as long as wide, 3.56-4.06 (median 3.78, n = 5) times as long as dactylus; carpus bearing one strong and 6 small short spiniform setae on posterior margin of outer surface, 7.50-8.36 (median 8.00, n = 5) times as long as wide, 0.63-0.75 (median 0.68, n = 5) times as long as propodus; merus slender, 10.36-12.62 (median 11.60, n = 5) times as long as wide, 1.68-1.93 (median 1.78, n = 5) times as long as carpus, bearing 3-5 strong spiniform setae on posterior margin of outer surface. Ischium with one spiniform seta. Fifth pereiopod (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10I, J</figureCitation>
|
||
) slender, dactylus 3.50-4.86 (median 4.13, n = 4) times as long as wide (terminal claw and serrate setae on flexor margin included), terminating in one large claw with 24-31 serrate setae on flexor margin; propodus slender, 13.43-16.00 (median 14.00, n = 4) times as long as wide, 3.29-3.50 (median 3.33, n = 4) times length of dactylus, carpus bearing one strong and 5-7 small spiniform setae on posterior margin of outer surface, 7.57-9.00 (median 7.88, n = 4) times as long as wide, 0.55-0.64 (median 0.57, n = 4) times as long as propodus, 0.63-0.70 (median 0.66, n = 4) times as long as merus; merus slender, 9.20-10.67 (median 10.55, n = 4) times as long as wide, 1.44-1.53 (median 1.51, n = 4) times length of carpus, bearing three or four strong spiniform setae on posterior margin of outer surface. Ischium without a strong spiniform seta.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">Figure 10.</emphasis>
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199
|
||
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
|
||
first maxilliped
|
||
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
|
||
palp of first maxilliped
|
||
<emphasis bold="true" pageId="0" pageNumber="81">C</emphasis>
|
||
second maxilliped
|
||
<emphasis bold="true" pageId="0" pageNumber="81">D</emphasis>
|
||
third maxilliped
|
||
<emphasis bold="true" pageId="0" pageNumber="81">E</emphasis>
|
||
first pereiopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">F</emphasis>
|
||
second pereiopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">I</emphasis>
|
||
fifth pereiopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">K</emphasis>
|
||
endopod of male first pleopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">L</emphasis>
|
||
appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199
|
||
<emphasis bold="true" pageId="0" pageNumber="81">G</emphasis>
|
||
third pereiopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">H</emphasis>
|
||
dactylus of third pereiopod
|
||
<emphasis bold="true" pageId="0" pageNumber="81">J</emphasis>
|
||
dactylus of fifth pereiopod. Scale bars: 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="81">A, C, H, J-L</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
|
||
); 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="81">D-G, I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Pleopods</emphasis>
|
||
.
|
||
</emphasis>
|
||
Endopod of male first pleopod (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10K</figureCitation>
|
||
) subtriangular, without an appendix interna, two pappose setae on outer, ~ eight on inner margin, 1.91-2.57 (median 2.00, n = 3) times as long as proximal width, 0.29-0.41 (median 0.29, n = 3) times as long as exopod. Appendix masculina on male second pleopod (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Caridina marlenae sp. nov. Morphology 2. Paratype ♂, cl. 2.2 mm, ZMB 30199 A first maxilliped B palp of first maxilliped C second maxilliped D third maxilliped E first pereiopod F second pereiopod I fifth pereiopod K endopod of male first pleopod L appendix masculina on male second pleopod; paratype ♀, cl. 2.3 mm, ZMB 30199 G third pereiopod H dactylus of third pereiopod J dactylus of fifth pereiopod. Scale bars: 0.5 mm (A, C, H, J-L); 0.1 mm (B); 1 mm (D-G, I)." figureDoi="10.3897/zookeys.1009.54303.figure10" httpUri="https://binary.pensoft.net/fig/493867" pageId="0" pageNumber="81">10L</figureCitation>
|
||
) rod-shaped, 6.00-8.40 (median 6.00, n = 3) times as long as wide, with long spiniform setae on inner and distal margin, few pappose setae on basal part, appendix interna reaching to ~ 0.72-0.80 (median 0.80, n = 3) of appendix masculina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="colouration">
|
||
<paragraph pageId="0" pageNumber="81">Colouration.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
Body colouration bright reddish with large white dots (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living specimens of Caridina spp in Lake Poso 2. A, B Caridina mayamareenae sp. nov. C C. lilianae sp. nov. D Two snail species Celetaia persculpta and Tylomelania sp. on soft substrate. Empty shells of these species are shelter for C. mayamareenae sp. nov. E, F C. poso sp. nov. G C. marlenae sp. nov. H C. fusca sp. nov. Not to scale. All photographs: W. Klotz." figureDoi="10.3897/zookeys.1009.54303.figure3" httpUri="https://binary.pensoft.net/fig/493860" pageId="0" pageNumber="81">3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="reproductive biology and larval development">
|
||
<paragraph pageId="0" pageNumber="81">Reproductive biology and larval development.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
Ovigerous females with few eggs (9, n = 1). Size of eggs 0.81-0.83
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
0.48-0.0.51 mm (n = 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="etymology">
|
||
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
Named after the second and last
|
||
<normalizedToken originalValue="authors’">authors'</normalizedToken>
|
||
second daughter who is very interested in field work and helped to observe and document this species while visiting the lake in 2019.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="distribution">
|
||
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is endemic to Lake Poso. Specimens were found only at one locality in a bay south of the town of Tentena at the east shore of the lake.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="ecology">
|
||
<paragraph pageId="0" pageNumber="81">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is found under rocks in deep water (more than 5 m).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="remarks">
|
||
<paragraph pageId="0" pageNumber="81">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
With its long rostrum, approximately anterior 0.4 unarmed,
|
||
<taxonomicName lsidName="C. marlenae" pageId="0" pageNumber="81" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
, all endemic to Lake Poso. In the field, body colouration alone is sufficient to differ
|
||
<taxonomicName lsidName="C. marlenae" pageId="0" pageNumber="81" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
but it might be confused with reddish specimens of
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is showing large bright white dots on reddish colouration of the entire body. In
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, the body is coloured dark brown with faint light transversal bands on first, third, fifth and sixth abdominal segments (W. Klotz, pers. observation on the comparative material listed above). In
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
the colouration of the body is mostly transparent with some brownish or whitish blotches.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
In preserved condition,
|
||
<taxonomicName lsidName="C. marlenae" pageId="0" pageNumber="81" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
by the more reduced epipods (slightly reduced on the third maxilliped, absent from all pereiopods vs. well developed (with distal hooks) on the third maxilliped and first pereiopod in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
and the slender chelipeds and pereiopods (in detail: chela of first pereiopod 3.00-3.83 times as long as wide vs. 1.74-2.1 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, 1.00-1.05 times as long as carpus vs. 1.35-1.48 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, dactylus 1.88-2.50 times as long as palm vs. 0.83-1.05 times as long in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, carpus 3.85-4.77 times as long as wide vs. 1.75-2.22 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, merus 3.80-4.25 times as long as wide vs. 1.78-2.63 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. Chela of second pereiopod 3.69-4.38 times as long as wide vs. 2.19-2.64 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, dactylus 1.80-2.25 times as long as palm vs. 1.05-1.33 times as long in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, carpus 6.78-9.80 times as long as wide vs. 4.56-5.05 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, merus 5.33-6.50 times as long as wide vs. 3.60-4.29 times in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. Dactylus of third pereiopod 4.40-5.33 times as long as wide vs. 4.0-4.20 in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, propodus 13.67-16.25 times as long as wide vs. 10.00-11.11 in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
, 3.56-4.06 times as long as dactylus vs. 2.86-3.13 times as long as dactylus in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. Propodus of fifth pereiopod 13.43-16.00 times as long as wide vs. 10.00-12.00 in
|
||
<taxonomicName lsidName="C. sarasinorum" pageId="0" pageNumber="81" rank="species" species="sarasinorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
by the more reduced epipods (slightly reduced on the third maxilliped, absent from all pereiopods vs. well developed (with distal hooks) on the third maxilliped and first and second pereiopod in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
and the slender chelipeds and armature of the dactyli of pereiopods (in detail: chela of first pereiopod 3.00-3.83 times as long as wide vs. 1.90-3.2 times in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
, dactylus 1.88-2.50 times as long as palm vs. 1.0-1.4 times as long in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
. Carpus of first cheliped 3.85-4.77 times as long as wide vs. 2.1-3.2 times as long as wide in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
. Dactylus of second pereiopod 1.80-2.25 times as long as palm vs. 1.2-1.4 times as long in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
, carpus 6.78-9.80 times as long as wide vs. 4.5-6.5 times in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
. Dactylus of third pereiopod with four or five spiniform setae on flexor margin vs. with 6-8 spiniform setae in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
. Dactylus of fifth pereiopod with 24-31 serrate setae on flexor margin vs. with 57-64 in
|
||
<taxonomicName lsidName="C. schenkeli" pageId="0" pageNumber="81" rank="species" species="schenkeli">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. schenkeli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<taxonomicName authorityName="Klotz, Wowor & K. von Rintelen" authorityYear="2021" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caridina marlenae" order="Decapoda" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marlenae">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Caridina marlenae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can also be distinguished from
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
by the type of chelae built for scraping vs. for filter-feeding in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
(brushes of setae short on tips of fingers of chelipeds vs. setae long, chela of first cheliped 3.00-3.83 times as long as wide vs. 4.6-6.5 times as long as wide in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
, dactylus 1.88-2.50 times as long as palm vs. 3.6-4.6 times as long in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
, carpus 3.85-4.77 times as long as wide vs. 4.8-8.1 times in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
; chela of second pereiopod 3.69-4.38 times as long as wide vs. 4.8-6.4 times in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
, dactylus 1.80-2.25 times as long as palm vs. 3.4-3.9 times as long in
|
||
<taxonomicName lsidName="C. longidigita" pageId="0" pageNumber="81" rank="species" species="longidigita">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. longidigita</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |