207 lines
33 KiB
XML
207 lines
33 KiB
XML
<document id="4766113F0A9B1E1804137FB75E0E25EE" ID-CLB-Dataset="29126" ID-DOI="10.11646/zootaxa.4497.4.3" ID-GBIF-Dataset="f09380a8-4d31-4e79-9b72-74ee26438553" ID-ISSN="1175-5326" ID-Zenodo-Dep="1456311" ID-ZooBank="7DFD16E7-D628-48DA-AAD6-4AB1E4D679E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539238820092" checkinUser="plazi" docAuthor="Reid, C. A. M. & Beatson, M." docDate="2018" docId="2A2D13766B68FF99FF1AFD21064A5D3E" docLanguage="en" docName="zootaxa.4497.4.3.pdf" docOrigin="Zootaxa 4497 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheiloxena monga Reid & Beatson 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="527" masterDocId="D6146B0E6B71FF83FF8DFFF604615833" masterDocTitle="Revision of the Australian leaf beetle genus Cheiloxena Baly, 1860 (Coleoptera: Chrysomelidae: Spilopyrinae)" masterLastPageNumber="534" masterPageNumber="501" pageNumber="526" updateTime="1698574941947" updateUser="ExternalLinkService">
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<mods:mods id="457C35AA42A7675044E2D9DC2A92E13E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E85EC269936950F6B6F741D1DD1E3464">
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<mods:title id="2E9A900569B901FEA2D2A857C445EBDB">Revision of the Australian leaf beetle genus Cheiloxena Baly, 1860 (Coleoptera: Chrysomelidae: Spilopyrinae)</mods:title>
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<mods:name id="1C48578F9033C6FDDCDE71ECAE155B10" type="personal">
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<mods:roleTerm id="49A77B69B8BC9F293D0578C607BBCE40">Author</mods:roleTerm>
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<mods:namePart id="B5E8D486CB8C06D486A20708BBD70AEA">Reid, C. A. M.</mods:namePart>
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<mods:roleTerm id="10AB30F2A83E37A33B675013A371B852">Author</mods:roleTerm>
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<mods:namePart id="57E70EA7E0D633380FB111548A1A800E">Beatson, M.</mods:namePart>
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<mods:typeOfResource id="E30AED72D96BBD480DFBD3DEA83486F9">text</mods:typeOfResource>
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<mods:title id="3008EB34956DF76FBDDC4F1883398409">Zootaxa</mods:title>
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<mods:part id="9714E816F1B57665B4F6CA0B9234622E">
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<mods:date id="0C70ADD1CE517796DBE8936504F2779C">2018</mods:date>
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<mods:number id="FDAEAFDF238E118DB30DF48269D390DE">2018-10-11</mods:number>
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<mods:number id="46557AD1F36E9D7336953A1C29CF307B">4497</mods:number>
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<mods:detail id="35ECAF117C79B8438C2DAE83FCE20EC2" type="issue">
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<mods:classification id="9260FCB907365FAA879BCD0D3DA90CE2">journal article</mods:classification>
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<mods:identifier id="BABE4200D7190F6FC618C00367CD4B86" type="DOI">10.11646/zootaxa.4497.4.3</mods:identifier>
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<treatment id="2A2D13766B68FF99FF1AFD21064A5D3E" ID-DOI="http://doi.org/10.5281/zenodo.5958942" ID-GBIF-Taxon="148673168" ID-Zenodo-Dep="5958942" LSID="urn:lsid:plazi:treatment:2A2D13766B68FF99FF1AFD21064A5D3E" httpUri="http://treatment.plazi.org/id/2A2D13766B68FF99FF1AFD21064A5D3E" lastPageId="26" lastPageNumber="527" pageId="25" pageNumber="526">
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<subSubSection id="EA9EF1EB6B68FF9AFF1AFD21058A5B20" pageId="25" pageNumber="526" type="nomenclature">
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<paragraph id="A23BA2606B68FF9AFF1AFD2105805AC1" blockId="25.[151,491,727,787]" box="[151,481,727,754]" pageId="25" pageNumber="526">
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<heading id="F973150C6B68FF9AFF1AFD2105805AC1" bold="true" box="[151,481,727,754]" fontSize="11" level="1" pageId="25" pageNumber="526" reason="1">
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<emphasis id="90F07E726B68FF9AFF1AFD2105805AC1" bold="true" box="[151,481,727,754]" pageId="25" pageNumber="526">
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<taxonomicName id="6584D9E36B68FF9AFF1AFD21051D5AC2" authority="Reid & Beatson, 2018" authorityName="Reid & Beatson" authorityYear="2018" box="[151,380,727,753]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="monga" status="sp. nov.">
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<emphasis id="90F07E726B68FF9AFF1AFD21051D5AC2" bold="true" box="[151,380,727,753]" italics="true" pageId="25" pageNumber="526">Cheiloxena monga</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8BC3C3096B68FF9AFE0EFD2E05805AC1" box="[387,481,728,754]" pageId="25" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A23BA2606B68FF9AFF1AFD0C058A5B20" blockId="25.[151,491,727,787]" box="[151,491,762,787]" pageId="25" pageNumber="526">
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(
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<figureCitation id="3ABFBEE56B68FF9AFF12FD0C04865B20" box="[159,231,762,787]" captionStart="FIGURES 7–10" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 7–10. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid & Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="25" pageNumber="526">Figs 9</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFF7EFD0D05705B20" box="[243,273,763,787]" captionStart="FIGURES 13–21" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 13–21. Lateral of Cheiloxena species: 13, C. aitori Reid & Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid & Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid & Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="25" pageNumber="526">18</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFE90FD0D055A5B20" box="[285,315,763,787]" captionStart="FIGURES 22–30" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 22–30. Anterior view, Cheiloxena species: 22, C. aitori Reid & Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid & Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid & Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="25" pageNumber="526">27</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFECAFD0D05045B20" box="[327,357,763,787]" captionStart="FIGURES 31–34" captionStartId="15.[151,264,1652,1675]" captionTargetBox="[217,1368,257,1517]" captionTargetId="figure@15.[150,1436,141,1632]" captionTargetPageId="15" captionText="FIGURES 31–34. Dorsal of pronotum, Cheiloxena species: 31, C. aitori Reid & Beatson, male (holotype); 32, C. blackburni Reid; 33, C. frenchae Blackburn; 34, C. monga Reid & Beatson." httpUri="https://zenodo.org/record/1456325/files/figure.png" pageId="25" pageNumber="526">34</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFEFCFD0D05EE5B20" box="[369,399,763,787]" captionStart="FIGURES 43–51" captionStartId="18.[151,264,1789,1812]" captionTargetId="figure@18.[152,1435,193,1768]" captionTargetPageId="18" captionText="FIGURES 43–51. Cheiloxena species, male genitalia lateral and dorsal: 43, C. aitori Reid & Beatson; 44, C. blackburni Reid; 45, C. conani Reid & Beatson; 46, C. frenchae Blackburn; 47, C. insignis Blackburn; 48, C. monga Reid & Beatson; 49, C. tuberosa Reid; 50, C. westwoodii Baly (northern form, Dorrigo); 51, C. westwoodii (southern form, Victoria). All images to scale, scale bar = 2mm." httpUri="https://zenodo.org/record/1457873/files/figure.png" pageId="25" pageNumber="526">48</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFE16FD0C05DB5B20" box="[411,442,762,787]" captionStart="FIGURES 52–61" captionStartId="19.[151,264,1911,1934]" captionTargetBox="[224,1367,255,1760]" captionTargetId="figure@19.[152,1435,131,1947]" captionTargetPageId="19" captionText="FIGURES 52–61. Cheiloxena species, apex of penis: 52, C. aitori Reid & Beatson; 53, C. blackburni Reid; 54, C. conani Reid & Beatson; 55, C. frenchae Blackburn; 56, C. insignis Blackburn (Launching Place, Victoria); 57, C. insignis Blackburn (Lake Mountain, Victoria); 58, C. monga Reid & Beatson; 59, C. tuberosa Reid; 60, C. westwoodii Baly (northern form, Dorrigo); 61, C. westwoodii (southern form, Victoria). All images to same scale." httpUri="https://zenodo.org/record/1456331/files/figure.png" pageId="25" pageNumber="526">58</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFE4BFD0D05825B20" box="[454,483,763,787]" captionStart="FIGURE 77" captionStartId="31.[151,250,1971,1993]" captionTargetId="figure@31.[151,1435,193,1950]" captionTargetPageId="31" captionText="FIGURE 77. Eastern Australia showing distribution of Cheiloxena species." httpUri="https://zenodo.org/record/1457875/files/figure.png" pageId="25" pageNumber="526">77</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="EA9EF1EB6B68FF9AFF1AFCB401235BAD" pageId="25" pageNumber="526" type="reference_group">
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<paragraph id="A23BA2606B68FF9AFF1AFCB4074C5B6B" blockId="25.[151,813,834,856]" box="[151,813,834,856]" pageId="25" pageNumber="526">
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<taxonomicName id="6584D9E36B68FF9AFF1AFCB405095B6B" authorityName="Blackburn. Not" authorityYear="1893" box="[151,360,834,856]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="frenchae">
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<emphasis id="90F07E726B68FF9AFF1AFCB405095B6B" box="[151,360,834,856]" italics="true" pageId="25" pageNumber="526">Cheiloxena frenchae</emphasis>
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</taxonomicName>
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male, sensu
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<bibRefCitation id="C615DF916B68FF9AFE66FCB406375B6B" box="[491,598,834,856]" pageId="25" pageNumber="526" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">Reid 1992</bibRefCitation>
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, nec
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<bibRefCitation id="C615DF916B68FF9AFD07FCB407485B6B" box="[650,809,834,856]" pageId="25" pageNumber="526" refId="ref19416" refString="Blackburn, T. (1893) Further notes on Australian Coleoptera, with descriptions of new genera and species. XIII. Transactions of the Royal Society of South Australia, 17, 130 - 140." type="journal article">Blackburn1893</bibRefCitation>
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.
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</paragraph>
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<paragraph id="A23BA2606B68FF9AFF1AFC7301235BAD" blockId="25.[151,1437,901,2006]" box="[151,1346,901,926]" pageId="25" pageNumber="526">
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<emphasis id="90F07E726B68FF9AFF1AFC7305E15BAD" bold="true" box="[151,384,901,926]" pageId="25" pageNumber="526">Mateiral examined.</emphasis>
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<typeStatus id="7D3F1CC26B68FF9AFE0AFC7005A35BAD" box="[391,450,902,926]" pageId="25" pageNumber="526">Type</typeStatus>
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.
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<typeStatus id="7D3F1CC26B68FF9AFE41FC7306585BAD" box="[460,569,901,926]" pageId="25" pageNumber="526" type="holotype">Holotype</typeStatus>
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: Ƌ*/ 5k SSW Monga NSW
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<emphasis id="90F07E726B68FF9AFCF9FC7007985BAE" box="[884,1017,901,925]" italics="true" pageId="25" pageNumber="526">
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Euc.
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<taxonomicName id="6584D9E36B68FF9AFC23FC7307985BAE" ID-CoL="3BQG3" authorityName="H.Deane & Maiden" authorityYear="1899" box="[942,1017,901,925]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="25" pageNumber="526" phylum="Tracheophyta" rank="species" species="rubida">rubida</taxonomicName>
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</emphasis>
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<date id="D63A84A06B68FF9AFB8DFC7300165BAD" box="[1024,1143,901,926]" pageId="25" pageNumber="526" value="1985-11-12">12.xi.1985</date>
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C. Reid/ (ANIC).
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</paragraph>
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</subSubSection>
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<subSubSection id="EA9EF1EB6B68FF9AFF4AFC5F07FF5C3A" pageId="25" pageNumber="526" type="diagnosis">
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<paragraph id="A23BA2606B68FF9AFF4AFC5F07FF5C3A" blockId="25.[151,1437,901,2006]" pageId="25" pageNumber="526">
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<emphasis id="90F07E726B68FF9AFF4AFC5F05205BF1" bold="true" box="[199,321,937,962]" pageId="25" pageNumber="526">Diagnosis.</emphasis>
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<taxonomicName id="6584D9E36B68FF9AFEC5FC5F067B5BF2" box="[328,538,937,961]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="monga">
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<emphasis id="90F07E726B68FF9AFEC5FC5F067B5BF2" box="[328,538,937,961]" italics="true" pageId="25" pageNumber="526">Cheiloxena monga</emphasis>
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</taxonomicName>
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is distinguished by: upper and lower surfaces scaled; anterior margin of clypeus truncate; pronotum with almost parallel paired ridges; hypomeral lobe rugose, without scales; apical half of elytra conspicuously tuberculate, with largest tubercles arranged in a curve.
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</paragraph>
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</subSubSection>
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<subSubSection id="EA9EF1EB6B68FF99FF4AFBE306C05BDE" lastPageId="26" lastPageNumber="527" pageId="25" pageNumber="526" type="description">
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<paragraph id="A23BA2606B68FF9AFF4AFBE306E45E31" blockId="25.[151,1437,901,2006]" pageId="25" pageNumber="526">
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<emphasis id="90F07E726B68FF9AFF4AFBE305305C1D" bold="true" box="[199,337,1045,1070]" pageId="25" pageNumber="526">Description</emphasis>
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(female unknown). Length
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<quantity id="657C0F856B68FF9AFD19FBE006865C1E" box="[660,743,1046,1070]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="526" unit="mm" value="11.0">11 mm</quantity>
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; body moderately convex in profile, length about
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<date id="D63A84A06B68FF9AFAA4FBE001245C1D" box="[1321,1349,1046,1070]" pageId="25" pageNumber="526">3x</date>
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height; colour black, but elytra mostly dark brown with black tubercles, and maxillary and labial palpi reddish-brown; almost entirely clothed with adpressed creamy-white scale-like setae (length 3–
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<date id="D63A84A06B68FF9AFB9EFBAB004E5C45" box="[1043,1071,1117,1142]" pageId="25" pageNumber="526">5x</date>
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width), not distinctly variegated in colour, setae not thinner on depressed areas of pronotum and elytra, thinner on anterior of clypeus, apices of mandibles, prosternal and mesoventral processes, legs, apex of abdominal ventrite V, and antennomeres 1–7, scalelike setae absent from antennomeres 8–11 (minute simple setae only); scale-like setae distributed as follows: almost evenly on head but clypeal margins anteriorly glabrous; almost evenly on pronotum but slightly sparser on depressed areas and absent from apices of ridges; scutellum clothed with thin setae; elytra with irregular mosaic of dense patches and more sparsely setose areas, setae of larger tubercles not differentiated except slightly more elongate; ventral surfaces with dense white adpressed setae except thin and sparse on apical half of ventrite V, setae of appendages thin but dense (except antennomeres 8–11); surface sculpture: head including clypeal margins, pronotum and metaventrite densely punctured and microreticulate, dull, elytra and rest of thoracic venter, including pronotal hypomera, densely microsculptured and dull; apices of tubercles and ridges, and abdominal ventrites shiny and not or shallowly microsculptured.
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</paragraph>
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<paragraph id="A23BA2606B68FF9AFF4AF9FB050C5F5A" blockId="25.[151,1437,901,2006]" pageId="25" pageNumber="526">
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Head (
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<figureCitation id="3ABFBEE56B68FF9AFE9CF9FB053B5E15" box="[273,346,1549,1574]" captionStart="FIGURES 7–10" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 7–10. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid & Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="25" pageNumber="526">Figs 9</figureCitation>
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,
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<figureCitation id="3ABFBEE56B68FF9AFEEAF9F805E45E15" box="[359,389,1550,1574]" captionStart="FIGURES 13–21" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 13–21. Lateral of Cheiloxena species: 13, C. aitori Reid & Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid & Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid & Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="25" pageNumber="526">18</figureCitation>
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,
|
||
<figureCitation id="3ABFBEE56B68FF9AFE1CF9F805D05E15" box="[401,433,1550,1574]" captionStart="FIGURES 22–30" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 22–30. Anterior view, Cheiloxena species: 22, C. aitori Reid & Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid & Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid & Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="25" pageNumber="526">27</figureCitation>
|
||
): slightly narrower than pronotum; densely and slightly confluently punctured, punctures small but larger than ommatidia, intervals dull and microsculptured, including narrow impunctate frontoclypeal sutures, apical margin clypeus, and margin of antennal cavities; convex between eyes, shallowly concave between antennae; eyes laterally prominent, separated by about 3 eye widths; gena at shortest point about 0.3x eye length; genal lobe
|
||
<date id="D63A84A06B68FF9AFE97F96805575E85" box="[282,310,1694,1718]" pageId="25" pageNumber="526">2x</date>
|
||
shortest length of gena; antennae
|
||
<date id="D63A84A06B68FF9AFD40F96806885E85" box="[717,745,1694,1718]" pageId="25" pageNumber="526">3x</date>
|
||
socket diameters apart; antennae about 0.75x body length; antennomeres 1–6 dull with shining apices, apical half of 7, and whole of 8–11 matt; antennomeres 1 and 3–11 elongate, 2 slightly transverse and about 0.5x length of 1, 0.25x length of 3, relative lengths of antennomeres, with 3 longest and 7 next, and middle segments unequal in length: 2, <1, <8, <4=9=10, <6, <5, <11, <7, <3; clypeus with 2 pairs of long subapical setae; clypeal anterior margin truncate; apical maxillary palpomere broadly ovoid, with truncate apex.
|
||
</paragraph>
|
||
<paragraph id="A23BA2606B68FF99FF4AF88305B85B0B" blockId="25.[151,1437,901,2006]" lastBlockId="26.[151,1437,151,1293]" lastPageId="26" lastPageNumber="527" pageId="25" pageNumber="526">
|
||
Thorax (
|
||
<figureCitation id="3ABFBEE56B68FF9AFEA5F88305125FBD" box="[296,371,1909,1934]" captionStart="FIGURES 7–10" captionStartId="7.[151,264,1975,1998]" captionTargetBox="[194,1366,311,1853]" captionTargetId="figure@7.[152,1435,139,1955]" captionTargetPageId="7" captionText="FIGURES 7–10. Habitus of Cheiloxena species: 7, C. insignis Baly (Launching Place, Victoria); 8, C. insignis (Lake Mountain, Victoria); 9, C. monga Reid & Beatson (holotype); 10, C. tuberosa Reid; Not to scale." httpUri="https://zenodo.org/record/1456317/files/figure.png" pageId="25" pageNumber="526">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="3ABFBEE56B68FF9AFE0CF88005FE5FBD" box="[385,415,1910,1934]" captionStart="FIGURES 13–21" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[230,1395,232,1777]" captionTargetId="figure@13.[152,1435,141,1956]" captionTargetPageId="13" captionText="FIGURES 13–21. Lateral of Cheiloxena species: 13, C. aitori Reid & Beatson, male (holotype); 14, C. blackburni Reid; 15, C. conani Reid & Beatson; 16, C. frenchae Blackburn, (holotype); 17, C. insignis Blackburn; 18, C. monga Reid & Beatson (holotype); 19, C. tuberosa Reid; 20, C. westwoodii Baly (northern form, Dorrigo); 21, C. westwoodii Baly (southern form, Victoria). Not to scale." httpUri="https://zenodo.org/record/1456321/files/figure.png" pageId="25" pageNumber="526">18</figureCitation>
|
||
,
|
||
<figureCitation id="3ABFBEE56B68FF9AFE20F88005AD5FBD" box="[429,460,1910,1934]" captionStart="FIGURES 22–30" captionStartId="14.[151,264,1700,1723]" captionTargetBox="[212,1381,282,1587]" captionTargetId="figure@14.[150,1436,193,1679]" captionTargetPageId="14" captionText="FIGURES 22–30. Anterior view, Cheiloxena species: 22, C. aitori Reid & Beatson, male (holotype); 23, C. blackburni Reid; 24, C. conani Reid & Beatson; 25, C. frenchae Blackburn; 26, C. insignis Baly; 27, C. monga Reid & Beatson; 28, C. tuberosa Reid; 29, C. westwoodii Baly (northern form, Dorrigo); 30, C. westwoodii Baly (southern form, Victoria)." httpUri="https://zenodo.org/record/1456323/files/figure.png" pageId="25" pageNumber="526">27</figureCitation>
|
||
,
|
||
<figureCitation id="3ABFBEE56B68FF9AFE54F88005985FBD" box="[473,505,1910,1934]" captionStart="FIGURES 31–34" captionStartId="15.[151,264,1652,1675]" captionTargetBox="[217,1368,257,1517]" captionTargetId="figure@15.[150,1436,141,1632]" captionTargetPageId="15" captionText="FIGURES 31–34. Dorsal of pronotum, Cheiloxena species: 31, C. aitori Reid & Beatson, male (holotype); 32, C. blackburni Reid; 33, C. frenchae Blackburn; 34, C. monga Reid & Beatson." httpUri="https://zenodo.org/record/1456325/files/figure.png" pageId="25" pageNumber="526">34</figureCitation>
|
||
): sides bulging slightly at middle, no clear distinction between dorsal and ventral (hypomeral) areas of pronotum; pronotal punctures dense, diameters 1.5–
|
||
<date id="D63A84A06B68FF9AFC5BF86F07935F81" box="[982,1010,1945,1970]" pageId="25" pageNumber="526">2x</date>
|
||
larger than on head; interspaces dull, densely microsculptured; pronotum slightly wider than long (measured at midline), length 0.95x width, greatest width at middle, sides weakly sinuate; anterior angles prominent but apices blunt, 90°; anterior margin produced, concave at middle; sides without tubercles; basal margin truncate; pronotal disc with two prominent subparallel keels from middle to anterior margin, almost symmetrically convex in lateral view, sharper in anterior half; remainder of disc unevenly surfaced, concave apicolaterally and either side of small postmedian swelling; hypomeron and prosternum densely punctured and pubescent, as pronotum, except hypomeral lobe rugose and glabrous; scutellum closely punctured and setose, scales more elongate than on pronotum; elytra irregularly scaled, forming network of pale flecks; elytron conspicuously tuberculate, with variably sized shiny-tipped tubercles arranged in approximately 4 clusters close to suture, from base to apex, as follows: first (basal) with 1 large blunt tubercle at apex of ridge from base of elytra; second (median) with 1 smaller conical or elongate tubercle; third (subapical) with 3 conical tubercles arranged in an equilateral triangle, largest lateral and smallest basal; fourth (apical) with 3 conspicuous but smaller well-separated tubercles arranged in an equilateral triangle; small tubercles scattered at base and sides of elytron; elytral disc shallowly arcuately depressed in basal half, posterior to humerus, and concave at sides of middle; elytral punctures large, similar diameter to largest on pronotum, but much deeper, and slightly sparser, separated by 1–1.5x diameters; interspaces dull, densely microsculptured; discal elytral punctures with two small shining tubercles laterally on rim; elytral apex narrowly truncate; epipleuron finely and shallowly punctured, with scale-like pubescence; mesoventrite punctured as prosternum; metepisternum and metaventrite dull and microsculptured, densely punctured and scaled, except midline impunctate and shining; metatibiae with shallow irregular lateral longitudinal grooves, thin and elongated; bases of tarsomeres 1–3 depressed and not expanded.
|
||
</paragraph>
|
||
<paragraph id="A23BA2606B6BFF99FF4AFCB206C05BDE" blockId="26.[151,1437,151,1293]" pageId="26" pageNumber="527">
|
||
Abdomen (
|
||
<figureCitation id="3ABFBEE56B6BFF99FEC8FCB205FC5B6F" box="[325,413,836,861]" captionStart="FIGURES 43–51" captionStartId="18.[151,264,1789,1812]" captionTargetId="figure@18.[152,1435,193,1768]" captionTargetPageId="18" captionText="FIGURES 43–51. Cheiloxena species, male genitalia lateral and dorsal: 43, C. aitori Reid & Beatson; 44, C. blackburni Reid; 45, C. conani Reid & Beatson; 46, C. frenchae Blackburn; 47, C. insignis Blackburn; 48, C. monga Reid & Beatson; 49, C. tuberosa Reid; 50, C. westwoodii Baly (northern form, Dorrigo); 51, C. westwoodii (southern form, Victoria). All images to scale, scale bar = 2mm." httpUri="https://zenodo.org/record/1457873/files/figure.png" pageId="26" pageNumber="527">Figs 48</figureCitation>
|
||
,
|
||
<figureCitation id="3ABFBEE56B6BFF99FE26FCB205AA5B6E" box="[427,459,836,861]" captionStart="FIGURES 52–61" captionStartId="19.[151,264,1911,1934]" captionTargetBox="[224,1367,255,1760]" captionTargetId="figure@19.[152,1435,131,1947]" captionTargetPageId="19" captionText="FIGURES 52–61. Cheiloxena species, apex of penis: 52, C. aitori Reid & Beatson; 53, C. blackburni Reid; 54, C. conani Reid & Beatson; 55, C. frenchae Blackburn; 56, C. insignis Blackburn (Launching Place, Victoria); 57, C. insignis Blackburn (Lake Mountain, Victoria); 58, C. monga Reid & Beatson; 59, C. tuberosa Reid; 60, C. westwoodii Baly (northern form, Dorrigo); 61, C. westwoodii (southern form, Victoria). All images to same scale." httpUri="https://zenodo.org/record/1456331/files/figure.png" pageId="26" pageNumber="527">58</figureCitation>
|
||
): ventrites I–V shinier than thoracic ventrites, strongly but less densely punctured and more shallowly microreticulate; ventrite pubescence recumbent, scale-like, but much narrower on ventrite V, only erect setae present on apical half of ventrite V; apex of ventrite V feebly convex; apex of penis contracted to blunt mucronate tip in dorsal view, apex thick and short in lateral view, strongly curved; tegminal keel shallowly sinuate in lateral view, with acute simple proximal tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA9EF1EB6B6BFF99FF4AFC01064A5D3E" pageId="26" pageNumber="527" type="etymology">
|
||
<paragraph id="A23BA2606B6BFF99FF4AFC0107D55C23" blockId="26.[151,1437,151,1293]" box="[199,948,1015,1040]" pageId="26" pageNumber="527">
|
||
<emphasis id="90F07E726B6BFF99FF4AFC01052F5C23" bold="true" box="[199,334,1015,1040]" pageId="26" pageNumber="527">Etymology.</emphasis>
|
||
Named from the
|
||
<typeStatus id="7D3F1CC26B6BFF99FD9EFC0E06225C23" box="[531,579,1016,1040]" pageId="26" pageNumber="527">type</typeStatus>
|
||
locality, as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="A23BA2606B6BFF99FF4AFBEA003E5C4E" blockId="26.[151,1437,151,1293]" pageId="26" pageNumber="527">
|
||
<emphasis id="90F07E726B6BFF99FF4AFBEA05705C06" bold="true" box="[199,273,1052,1077]" pageId="26" pageNumber="527">Notes.</emphasis>
|
||
<taxonomicName id="6584D9E36B6BFF99FE93FBEA05995C07" box="[286,504,1052,1076]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="monga">
|
||
<emphasis id="90F07E726B6BFF99FE93FBEA05995C07" box="[286,504,1052,1076]" italics="true" pageId="26" pageNumber="527">Cheiloxena monga</emphasis>
|
||
</taxonomicName>
|
||
is only known from the
|
||
<typeStatus id="7D3F1CC26B6BFF99FCBEFBEA07F95C06" box="[819,920,1052,1077]" pageId="26" pageNumber="527" type="holotype">holotype</typeStatus>
|
||
, collected at the margin of cool temperate rainforest and tall eucalypt forest in what is now Monga National Park. It was beaten off a sapling of
|
||
<taxonomicName id="6584D9E36B6BFF99FA92FBB604835C4F" authorityName="H.Deane & Maiden" authorityYear="1899" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="26" pageNumber="527" phylum="Tracheophyta" rank="species" species="rubida">
|
||
<emphasis id="90F07E726B6BFF99FA92FBB604835C4F" italics="true" pageId="26" pageNumber="527">Eucalyptus rubida</emphasis>
|
||
</taxonomicName>
|
||
, now a synonym of
|
||
<taxonomicName id="6584D9E36B6BFF99FE4BFB9307755C4F" authorityName="Maiden" authorityYear="1920" box="[454,788,1124,1148]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="26" pageNumber="527" phylum="Tracheophyta" rank="species" species="dalrympleana">
|
||
<emphasis id="90F07E726B6BFF99FE4BFB9306E05C4F" box="[454,641,1124,1148]" italics="true" pageId="26" pageNumber="527">E. dalrympleana</emphasis>
|
||
(Myrtaceae)
|
||
</taxonomicName>
|
||
, with no evidence of feeding.
|
||
</paragraph>
|
||
<paragraph id="A23BA2606B6BFF99FF4AFB71064A5D3E" blockId="26.[151,1437,151,1293]" pageId="26" pageNumber="527">
|
||
This specimen was previously incorporated in a description of
|
||
<taxonomicName id="6584D9E36B6BFF99FC34FB7E00215C93" authorityName="Blackburn. Not" authorityYear="1893" box="[953,1088,1160,1184]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="frenchae">
|
||
<emphasis id="90F07E726B6BFF99FC34FB7E00215C93" box="[953,1088,1160,1184]" italics="true" pageId="26" pageNumber="527">C. frenchae</emphasis>
|
||
</taxonomicName>
|
||
, when the only other known specimen of that species was the
|
||
<typeStatus id="7D3F1CC26B6BFF99FD98FB5A06165CF6" box="[533,631,1196,1221]" pageId="26" pageNumber="527" type="holotype">holotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="C615DF916B6BFF99FD0AFB5A07625CF7" box="[647,771,1196,1221]" pageId="26" pageNumber="527" refId="ref20270" refString="Reid, C. A. M. (1992) Revision of the genus Cheiloxena Baly (Coleoptera: Chrysomelidae: Eumolpinae). Memoirs of the Museum of Fictoria, 53 (1), 101 - 114. https: // doi. org / 10.24199 / j. mmv. 1992.53.04" type="journal article">Reid 1992</bibRefCitation>
|
||
). Based on the small amount of additional material of this complex (
|
||
<taxonomicName id="6584D9E36B6BFF99FE80FB2605135CD4" authorityName="Reid & Beatson" authorityYear="2018" box="[269,370,1232,1255]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="aitori">
|
||
<emphasis id="90F07E726B6BFF99FE80FB2605135CD4" box="[269,370,1232,1255]" italics="true" pageId="26" pageNumber="527">C. aitori</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6584D9E36B6BFF99FE0BFB26066C5CDB" authorityName="Blackburn. Not" authorityYear="1893" box="[390,525,1232,1256]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="frenchae">
|
||
<emphasis id="90F07E726B6BFF99FE0BFB26066C5CDB" box="[390,525,1232,1256]" italics="true" pageId="26" pageNumber="527">C. frenchae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6584D9E36B6BFF99FDC2FB2606A35CDB" box="[591,706,1232,1256]" class="Insecta" family="Chrysomelidae" genus="Cheiloxena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="monga">
|
||
<emphasis id="90F07E726B6BFF99FDC2FB2606A35CDB" box="[591,706,1232,1256]" italics="true" pageId="26" pageNumber="527">C. monga</emphasis>
|
||
</taxonomicName>
|
||
) made available since 1992 we now believe that the Monga specimen represents a valid species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |