562 lines
68 KiB
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562 lines
68 KiB
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<mods:title id="34E233AD8FFC00AAEEA822370D1A4868">Two new species of the genus Sitana Cuvier, 1829 (Reptilia: Agamidae) from Sri Lanka, including a taxonomic revision of the Indian Sitana species</mods:title>
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<mods:namePart id="1F0331DF5E9A9F19D583239B9210CAE0">Thasun Amarasinghe, A. A.</mods:namePart>
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<mods:namePart id="FE27480137FFE62E2E1A1D1DD6BE6524">Ineich, Ivan</mods:namePart>
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<mods:namePart id="691C91B698387CF1F4D416BFE1A98B30">Suranjan Karunarathna, D. M. S.</mods:namePart>
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<mods:namePart id="44A1A56CCFAAB2BA36CD85A239E81D2B">Botejue, W. Madhava S.</mods:namePart>
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<mods:title id="A67FC73C766009C900C516CA0E2BF8D4">Zootaxa</mods:title>
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<treatment id="DA632C027E50E50E46A6FE65FEE8E802" ID-DOI="http://doi.org/10.5281/zenodo.6113349" ID-GBIF-Taxon="127671455" ID-Zenodo-Dep="6113349" LSID="urn:lsid:plazi:treatment:DA632C027E50E50E46A6FE65FEE8E802" httpUri="http://treatment.plazi.org/id/DA632C027E50E50E46A6FE65FEE8E802" lastPageId="14" lastPageNumber="81" pageId="10" pageNumber="77">
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<paragraph id="52759D147E50E50A46A6FE65FC17E8EF" blockId="10.[151,918,437,496]" box="[151,918,437,463]" pageId="10" pageNumber="77">
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<heading id="093D2A787E50E50A46A6FE65FC17E8EF" bold="true" box="[151,918,437,463]" fontSize="11" level="1" pageId="10" pageNumber="77" reason="1">
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<emphasis id="60BE41067E50E50A46A6FE65FC17E8EF" bold="true" box="[151,918,437,463]" pageId="10" pageNumber="77">
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<taxonomicName id="95CAE6977E50E50A46A6FE65FEC8E8EF" box="[151,329,437,463]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
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<emphasis id="60BE41067E50E50A46A6FE65FEC8E8EF" bold="true" box="[151,329,437,463]" italics="true" pageId="10" pageNumber="77">Sitana devakai</emphasis>
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<taxonomicNameLabel id="7B8DFC7D7E50E50A4761FE65FE2CE8EF" box="[336,429,437,463]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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Amarasinghe, Ineich & Karunarathna
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="52759D147E50E50A46A6FE07FE1BE8D0" blockId="10.[151,918,437,496]" box="[151,410,471,496]" pageId="10" pageNumber="77">
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(
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<figureCitation id="CAF181917E50E50A46AEFE07FF6CE8D0" box="[159,237,471,496]" captionStart="FIGURE 3" captionStartId="7.[151,250,1337,1359]" captionTargetBox="[311,1275,301,1312]" captionTargetId="figure@7.[311,1275,301,1316]" captionTargetPageId="7" captionText="FIGURE 3. Sitana bahiri sp. nov. (A) lateral body scales of holotype male (WHT 1434 A); (B) pectoral scales and (C) abdominal scales of paratype female (WHT 0195 A); Sitana devakai sp. nov. (D) lateral body scales of holotype male (WHT 0111 E); (E) pectoral scales and (F) abdominal scales of paratype female (WHT 0111 C) (illustrated by A. A. T. Amarasinghe)." httpUri="https://zenodo.org/record/287774/files/figure.png" pageId="10" pageNumber="77">Figs. 3</figureCitation>
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,
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<figureCitation id="CAF181917E50E50A46CBFE08FE8AE8D0" box="[250,267,472,496]" captionStart="FIGURE 4" captionStartId="11.[151,250,1528,1550]" captionTargetBox="[311,1275,179,1502]" captionTargetId="figure@11.[311,1275,179,1503]" captionTargetPageId="11" captionText="FIGURE 4. Holotype male WHT 0111 E of Sitana devakai sp. nov., (A) dorsal head view; (B) lateral head view; (C) ventral head view; (D) ventral thigh and tail base (photo: M. C. De Silva); (E) Sitana devakai sp. nov., live breeding male with expanded throat-fan (not collected) at Wilpaththu National Park, Sri Lanka (photo: V. Weeratunge)." httpUri="https://zenodo.org/record/287775/files/figure.png" pageId="10" pageNumber="77">4</figureCitation>
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;
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<tableCitation id="1F48A8AF7E50E50A4726FE07FE10E8D0" box="[279,401,471,496]" captionStart-0="TABLE 1. A" captionStart-1="TABLE 2" captionStart-2="TABLE 3. A" captionStartId-0="8.[151,239,152,175]" captionStartId-1="8.[151,239,1396,1419]" captionStartId-2="9.[151,239,152,175]" captionTargetBox-0="[159,1414,234,1331]" captionTargetBox-1="[159,1424,1478,1655]" captionTargetBox-2="[159,1400,234,1362]" captionTargetPageId-0="8" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="TABLE 1. A comparison of morphometric and meristic characters of male specimens of Sitana ponticeriana, S. deccanensis, S. bahiri sp. nov., and S. devakai sp. nov. based on examined materials." captionText-1="TABLE 2. External morphological character variation in Sitana ponticeriana, S. deccanensis, S. bahiri sp. nov., and S. devakai sp. nov. based on examined male specimens." captionText-2="TABLE 3. A comparison of morphometric and meristic characters between the Sri Lanka species: S. bahiri sp. nov. and S. devakai sp. nov. based on examined specimens; NA, not applicable." httpUri-0="http://table.plazi.org/id/06B5CD9C7E52E50846A6FF48FBA5E9EF" httpUri-1="http://table.plazi.org/id/06B5CD9C7E52E50846A6FAA4FD51EC8B" httpUri-2="http://table.plazi.org/id/06B5CD9C7E53E50946A6FF48FC09E9EF" pageId="10" pageNumber="77" tableUuid-0="06B5CD9C7E52E50846A6FF48FBA5E9EF" tableUuid-1="06B5CD9C7E52E50846A6FAA4FD51EC8B" tableUuid-2="06B5CD9C7E53E50946A6FF48FC09E9EF">Tables 1–3</tableCitation>
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)
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</paragraph>
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<paragraph id="52759D147E50E50A46A6FDF1FD29EB16" blockId="10.[151,1436,544,719]" box="[151,680,544,566]" pageId="10" pageNumber="77">
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<taxonomicName id="95CAE6977E50E50A46A6FDF1FEE7EB16" authority="Ponticereana" authorityName="Ponticereana" box="[151,358,545,566]" class="Reptilia" family="Agamidae" genus="Litana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="genus">
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<emphasis id="60BE41067E50E50A46A6FDF1FEE7EB16" box="[151,358,545,566]" italics="true" pageId="10" pageNumber="77">Litana Ponticereana</emphasis>
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</taxonomicName>
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[sic]—
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<bibRefCitation id="365BE0E57E50E50A479EFDF0FDC6EB16" author="Kelaart" box="[431,583,544,566]" pageId="10" pageNumber="77" refString="Kelaart, E. F. (1854) Catalogue of reptiles collected in Ceylon. Annals and Magazine of Natural History, 2 (XIII), 137 - 140. http: // dx. doi. org / 10.1080 / 03745485709495092" type="journal article" year="1854">Kelaart (1854)</bibRefCitation>
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[
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<emphasis id="60BE41067E50E50A4467FDF1FD1BEB16" box="[598,666,545,566]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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].
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</paragraph>
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<emphasis id="60BE41067E50E50A46A6FDEFFEE2EB75" box="[151,355,575,597]" italics="true" pageId="10" pageNumber="77">Sitana ponticeriana</emphasis>
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</taxonomicName>
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—
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[
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<emphasis id="60BE41067E50E50A442BFD90FDDFEB75" box="[538,606,576,597]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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]; Taylor (1957) [
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];
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[
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<emphasis id="60BE41067E50E50A4204FD90FBF8EB75" box="[1077,1145,576,597]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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];
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<bibRefCitation id="365BE0E57E50E50A42A2FDEEFEEFEB53" author="Manamendra-Arachchi" pageId="10" pageNumber="77" refString="Manamendra-Arachchi, K. & Liyanage, S. (1994) Conservation and distributions of the agamid lizards of Sri Lanka with illustrations of the extant species. Journal of South Asian Natural History, 1, 77 - 96." type="journal article" year="1994">Manamendra-Arachchi & Liyanage (1994)</bibRefCitation>
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[
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<emphasis id="60BE41067E50E50A474CFD8EFE41EB53" box="[381,448,606,627]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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];
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<bibRefCitation id="365BE0E57E50E50A47E7FD8DFDEFEB53" author="Erdelen" box="[470,622,605,627]" pageId="10" pageNumber="77" refString="Erdelen, W. (1998) The genera Otocryptis and Sitana: Geographic distribution, microhabitat use and morphometric relations. In: Biology and Conservation of the Amphibians, Reptiles and their habitats in South Asia. Proceedings of the International Conference on the Biology and Conservation of the Amphibians and Reptiles of South Asia, Sri Lanka, pp. 232 - 246." type="proceedings paper" year="1998">Erdelen (1998)</bibRefCitation>
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[
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<emphasis id="60BE41067E50E50A444CFD8EFD41EB53" box="[637,704,606,627]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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]; Das & de Silva (2005) [
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<emphasis id="60BE41067E50E50A45F8FD8EFB8DEB53" box="[969,1036,606,627]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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]; de
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<bibRefCitation id="365BE0E57E50E50A4271FD8DFB3FEB53" author="Silva" box="[1088,1214,605,627]" pageId="10" pageNumber="77" refString="de Silva, A. (2006) Current status of the Reptiles of Sri Lanka. In: Bambaradeniya, C. N. B. (Ed.). Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation, IUCN Sri Lanka, Colombo, 134 - 163 pp." type="book chapter" year="2006">Silva (2006)</bibRefCitation>
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[
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];
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<bibRefCitation id="365BE0E57E50E50A4317FD8DFE21EBB2" author="Somaweera" pageId="10" pageNumber="77" refString="Somaweera, R. & Somaweera, N. (2009) Lizards of Sri Lanka: A colour guide with field keys. Edition Chimaira, Frankfurt / Main, Germany, 303 pp." type="book" year="2009">Somaweera & Somaweera (2009)</bibRefCitation>
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[
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<emphasis id="60BE41067E50E50A479EFDADFE72EBB2" box="[431,499,637,658]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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];
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<bibRefCitation id="365BE0E57E50E50A4438FDACFD2DEBB2" author="Manthey" box="[521,684,636,658]" pageId="10" pageNumber="77" refString="Manthey, U. (2010) Agamid Lizards of Southern Asia, Draconinae 2 & Leiolepidinae. Edition Chimaira & A. C. S. Glaser, Germany, Frankfurt am Main, 168 pp." type="book" year="2010">Manthey (2010)</bibRefCitation>
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[
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<emphasis id="60BE41067E50E50A448AFDADFD7EEBB2" box="[699,767,637,658]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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].
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</paragraph>
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<paragraph id="52759D147E50E50A46A6FD4BFCB3EB90" blockId="10.[151,1436,544,719]" box="[151,818,666,688]" pageId="10" pageNumber="77">
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<taxonomicName id="95CAE6977E50E50A46A6FD4BFE67EB90" box="[151,486,667,688]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="subSpecies" species="ponticeriana" subSpecies="ponticeriana">
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<emphasis id="60BE41067E50E50A46A6FD4BFE67EB90" box="[151,486,667,688]" italics="true" pageId="10" pageNumber="77">Sitana ponticeriana ponticeriana</emphasis>
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</taxonomicName>
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||
—
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||
<bibRefCitation id="365BE0E57E50E50A4430FD4AFD50EB90" author="Deraniyagala" box="[513,721,666,688]" pageId="10" pageNumber="77" refString="Deraniyagala, P. E. P. (1953) A Colored Atlas of some vertebrates from Ceylon, Tetrapod Reptilia. Vol. 2. National Museums of Sri Lanka, Colombo, 101 pp." type="book" year="1953">Deraniyagala (1953)</bibRefCitation>
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[
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||
<emphasis id="60BE41067E50E50A44D1FD4BFCA5EB90" box="[736,804,667,688]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
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].
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</paragraph>
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<paragraph id="52759D147E50E50A46A6FD6AFCA7EBEF" blockId="10.[151,1436,544,719]" box="[151,806,697,719]" pageId="10" pageNumber="77">
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<taxonomicName id="95CAE6977E50E50A46A6FD6AFEE6EBEF" box="[151,359,698,719]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="ponticertiana">
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<emphasis id="60BE41067E50E50A46A6FD6AFEE6EBEF" box="[151,359,698,719]" italics="true" pageId="10" pageNumber="77">Sitana ponticertiana</emphasis>
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</taxonomicName>
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[sic]—
|
||
<bibRefCitation id="365BE0E57E50E50A4781FD69FD47EBEF" author="Bahir" box="[432,710,697,719]" pageId="10" pageNumber="77" refString="Bahir, M. M. & Silva, A. (2005) Otocryptis nigristigma, a new species of Agamid Lizard from Sri Lanka. In: Yeo, C. J., Ng P. K. L. & Pethiyagoda, R. (Ed.), Contributions to biodiversity exploration and research in Sri Lanka. The Raffles Bulletin of Zoology, 12 (Supplement), 393 - 406." type="journal article" year="2005">Bahir & Surasinghe (2005)</bibRefCitation>
|
||
[
|
||
<emphasis id="60BE41067E50E50A44E5FD6AFC99EBEF" box="[724,792,698,719]" italics="true" pageId="10" pageNumber="77">partim</emphasis>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E50E50A46A6FD2CFCBBED15" pageId="10" pageNumber="77" type="materials_examined">
|
||
<paragraph id="52759D147E50E50A46A6FD2CFC3DEA18" blockId="10.[151,1437,764,2013]" pageId="10" pageNumber="77">
|
||
<emphasis id="60BE41067E50E50A46A6FD2CFE8AEA35" bold="true" box="[151,267,764,789]" pageId="10" pageNumber="77">
|
||
<typeStatus id="8D7123B67E50E50A46A6FD2CFE87EA35" box="[151,262,764,789]" pageId="10" pageNumber="77" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male,
|
||
<collectionCode id="34DB05D17E50E50A476CFD2DFE1CEA34" box="[349,413,765,788]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
0111E, SVL
|
||
<quantity id="953230F17E50E50A440BFD2CFD23EA34" box="[570,674,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="10" pageNumber="77" unit="mm" value="45.6">45.6 mm</quantity>
|
||
, collected at Nagagamuwa-Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A4260FD2CFB41EA35" box="[1105,1216,764,789]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
(
|
||
<geoCoordinate id="37FEFBD37E50E50A42E3FD2CFAA2EA35" box="[1234,1315,764,789]" direction="north" orientation="latitude" pageId="10" pageNumber="77" precision="925" value="8.166667">8°10′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="37FEFBD37E50E50A4303FD2CFA11EA35" box="[1330,1424,764,789]" direction="east" orientation="longitude" pageId="10" pageNumber="77" precision="925" value="79.833336">79°50′E</geoCoordinate>
|
||
), alt.
|
||
<quantity id="953230F17E50E50A46F0FCF0FF6FEA17" box="[193,238,800,824]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="77" unit="m" value="2.0">2 m</quantity>
|
||
, by K. Manamendra-Arachchi & D. Gabadage,
|
||
<date id="2674BBD47E50E50A4536FCF0FC39EA18" box="[775,952,799,824]" pageId="10" pageNumber="77" value="1993-10-09">9 October 1993</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="52759D147E50E50A46F6FC94FCBBED15" blockId="10.[151,1437,764,2013]" pageId="10" pageNumber="77">
|
||
<emphasis id="60BE41067E50E50A46F6FC94FEC7EA7D" bold="true" box="[199,326,836,861]" pageId="10" pageNumber="77">
|
||
<typeStatus id="8D7123B67E50E50A46F6FC94FEC0EA7D" box="[199,321,836,861]" pageId="10" pageNumber="77" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Males,
|
||
<collectionCode id="34DB05D17E50E50A4793FC95FE63EA7C" box="[418,482,837,860]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
0174A–B, respective SVL
|
||
<quantity id="953230F17E50E50A452CFC94FC03EA7C" box="[797,898,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="4.11" pageId="10" pageNumber="77" unit="mm" value="41.1">41.1 mm</quantity>
|
||
and 40.0 mm, collected at Palavi-Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A434AFC94FF60EAA0" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Male,
|
||
<collectionCode id="34DB05D17E50E50A4702FCB8FEF2EA5F" box="[307,371,872,895]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
1417, SVL
|
||
<quantity id="953230F17E50E50A47CDFCB8FDE3EA5F" box="[508,610,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="4.39" pageId="10" pageNumber="77" unit="mm" value="43.9">43.9 mm</quantity>
|
||
, collected at Angamuwewa-Rajanganaya,
|
||
<collectingCountry id="2ADDDD847E50E50A420FFCB7FB31EAA0" box="[1086,1200,871,896]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Male,
|
||
<collectionCode id="34DB05D17E50E50A4333FCB8FAC3EA5F" box="[1282,1346,872,895]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
1424A, SVL
|
||
<quantity id="953230F17E50E50A46E2FC5CFEB8EA84" box="[211,313,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="3.9899999999999998" pageId="10" pageNumber="77" unit="mm" value="39.9">39.9 mm</quantity>
|
||
, collected at Mundel-Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A4491FC5CFC93EA85" box="[672,786,908,933]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Female,
|
||
<collectionCode id="34DB05D17E50E50A454FFC5CFC41EA84" box="[894,960,908,932]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
0174C, SVL
|
||
<quantity id="953230F17E50E50A4262FC5CFB38EA84" box="[1107,1209,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="4.27" pageId="10" pageNumber="77" unit="mm" value="42.7">42.7 mm</quantity>
|
||
, collected at Palavi- Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A4738FC7FFEFBEAE8" box="[265,378,943,968]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Females,
|
||
<collectionCode id="34DB05D17E50E50A47C0FC60FDB0EAE7" box="[497,561,944,967]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
0111B–C, respective SVL 42.0 mm and
|
||
<quantity id="953230F17E50E50A45CEFC60FBE4EAE7" box="[1023,1125,944,968]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="10" pageNumber="77" unit="mm" value="44.9">44.9 mm</quantity>
|
||
, collected at Nagagamuwa- Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A473BFC04FEFDEACD" box="[266,380,980,1005]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Sub adult male,
|
||
<collectionCode id="34DB05D17E50E50A4473FC05FD03EACC" box="[578,642,981,1004]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
1424B, SVL
|
||
<quantity id="953230F17E50E50A4510FC04FC09EACC" box="[801,904,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="3.86" pageId="10" pageNumber="77" unit="mm" value="38.6">38.6 mm</quantity>
|
||
, collected at Mundel-Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A42C3FC04FAE4EACD" box="[1266,1381,980,1005]" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
; Sub adult females,
|
||
<collectionCode id="34DB05D17E50E50A4770FC28FE00ED2F" box="[321,385,1016,1039]" pageId="10" pageNumber="77">WHT</collectionCode>
|
||
0111A,D, respective SVL
|
||
<quantity id="953230F17E50E50A448DFC27FCA2ED2F" box="[700,803,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.76" pageId="10" pageNumber="77" unit="mm" value="37.6">37.6 mm</quantity>
|
||
and
|
||
<quantity id="953230F17E50E50A456EFC28FC49ED2F" box="[863,968,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="10" pageNumber="77" unit="mm" value="38.1">38.1 mm</quantity>
|
||
, collected at Nagagamuwa-Puttalam,
|
||
<collectingCountry id="2ADDDD847E50E50A434DFC27FF60ED15" name="Sri Lanka" pageId="10" pageNumber="77">Sri Lanka</collectingCountry>
|
||
. All those specimens were collected by D. Gabadage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E50E50A46F6FBEFFC7CEFB8" pageId="10" pageNumber="77" type="diagnosis">
|
||
<paragraph id="52759D147E50E50A46F6FBEFFC7CEFB8" blockId="10.[151,1437,764,2013]" pageId="10" pageNumber="77">
|
||
<emphasis id="60BE41067E50E50A46F6FBEFFD8AED78" bold="true" box="[199,523,1087,1112]" pageId="10" pageNumber="77">Diagnosis for adult males.</emphasis>
|
||
<taxonomicName id="95CAE6977E50E50A4426FB90FD43ED78" box="[535,706,1087,1112]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E50E50A4426FB90FD43ED78" box="[535,706,1087,1112]" italics="true" pageId="10" pageNumber="77">Sitana devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E50E50A44FFFBEFFCADED78" box="[718,812,1087,1112]" pageId="10" pageNumber="77" rank="species">
|
||
<emphasis id="60BE41067E50E50A44FFFBEFFCADED78" bold="true" box="[718,812,1087,1112]" pageId="10" pageNumber="77">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from each congener by having the following characters:
|
||
<emphasis id="60BE41067E50E50A4729FBB3FE5EED5D" bold="true" box="[280,479,1123,1149]" pageId="10" pageNumber="77">
|
||
<taxonomicName id="95CAE6977E50E50A4729FBB3FEFDED5C" box="[280,380,1123,1148]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="bahiri" status="sp. nov.">
|
||
<emphasis id="60BE41067E50E50A4729FBB3FEFDED5C" bold="true" box="[280,380,1123,1148]" italics="true" pageId="10" pageNumber="77">S. bahiri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E50E50A47B5FBB4FE5EED5D" box="[388,479,1124,1149]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: longer throat-fan ADW 29.9–32.5% of AG (vs. shorter, ADW 28.3–29.2% of AG), midbody scale rows 57–59 (vs. 49–56), ventral scale number 100–108 (vs. 87–89), abdominal scales pointed (vs. bluntly pointed), equal and regular lateral scales with intermediate enlarged scales (vs. unequal and irregular lateral scales with intermediate enlarged scales), breeding-males with bright red colour patch (vs. bleached orange colour patch);
|
||
<emphasis id="60BE41067E50E50A46DDFB24FE1CEC2D" bold="true" box="[236,413,1267,1293]" pageId="10" pageNumber="77">
|
||
S.
|
||
<taxonomicName id="95CAE6977E50E50A4738FB23FE15EC2C" box="[265,404,1267,1292]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="deccanensis">
|
||
<emphasis id="60BE41067E50E50A4738FB23FE15EC2C" bold="true" box="[265,404,1267,1292]" italics="true" pageId="10" pageNumber="77">deccanensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
smaller SVL 40.0–
|
||
<quantity id="953230F17E50E50A4449FB24FD5DEC2C" box="[632,732,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="10" pageNumber="77" unit="mm" value="45.6">45.6 mm</quantity>
|
||
(vs. larger SVL above 60.0 mm), snout longer ES 57.2–63.2% of HW (vs. shorter, ES 53.1–53.4% of HW), throat-fan shorter ADW 29.9–32.5% of AG (vs. longer, ADW 50.3–60.2% of AG), femur shorter FEL 72.2–78.4% of TBL (vs. longer, FEL 79.7–82.9% of TBL), foot longer FOL 154.7–180.2 of HL (vs. shorter, FOL 131.9–144.6% of HL), supralabials 8–9 (vs. 12), midbody scale rows 57–59 (vs. 66–70), ventral scale number 100–108 (vs. 82–92), subdigital lamellae on toe
|
||
<date id="2674BBD47E50E50A42F0FA55FAAEECBD" box="[1217,1327,1412,1437]" pageId="10" pageNumber="77" value="1915-03-16">III 15–16</date>
|
||
(vs. 13), enlarged scales above the tympanum present (vs. absent), lateral scales with intermediate enlarged scales (vs. without enlarged scales), breeding-males with bright red colour patch restricted to mid throat-fan (vs. bright red patch extending to vent), single pale stripe running from the snout up to the shoulder (vs. 3–4 bright stripes from snout and behind the eye up to the shoulders);
|
||
<emphasis id="60BE41067E50E50A44AEF9C3FCD9EF0D" bold="true" box="[671,856,1555,1581]" pageId="10" pageNumber="77">
|
||
<taxonomicName id="95CAE6977E50E50A44AEF9C3FCCEEF0C" box="[671,847,1555,1580]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="ponticeriana">
|
||
<emphasis id="60BE41067E50E50A44AEF9C3FCCEEF0C" bold="true" box="[671,847,1555,1580]" italics="true" pageId="10" pageNumber="77">S. ponticeriana</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
femur shorter FEL 72.2–78.4% of TBL (vs. longer, FEL 80.0–84.8% of TBL), ventral scale number 100–108 (vs. 65–69), abdominal scales pointed (vs. rounded);
|
||
<emphasis id="60BE41067E50E50A46A6F98BFE53EF55" bold="true" box="[151,466,1627,1653]" pageId="10" pageNumber="77">
|
||
<taxonomicName id="95CAE6977E50E50A46A6F98BFEDBEF54" box="[151,346,1627,1652]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="sivalensis">
|
||
<emphasis id="60BE41067E50E50A46A6F98BFEDBEF54" bold="true" box="[151,346,1627,1652]" italics="true" pageId="10" pageNumber="77">Sitana sivalensis</emphasis>
|
||
</taxonomicName>
|
||
complex:
|
||
</emphasis>
|
||
throat-fan extending up to the chest (vs. not extending to the level of axilla), foot longer with FOL 154.7–180.2 of HL (vs. shorter, FOL 120.0–144.3% of HL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E50E50D46F6F974FD7AEA87" lastPageId="13" lastPageNumber="80" pageId="10" pageNumber="77" type="description">
|
||
<paragraph id="52759D147E50E50C46F6F974FE29E9DB" blockId="10.[151,1437,764,2013]" lastBlockId="12.[151,1437,151,2015]" lastPageId="12" lastPageNumber="79" pageId="10" pageNumber="77">
|
||
<emphasis id="60BE41067E50E50A46F6F974FED8EF9D" bold="true" box="[199,345,1700,1725]" pageId="10" pageNumber="77">Description.</emphasis>
|
||
(Based on
|
||
<typeStatus id="8D7123B67E50E50A47EFF974FDC2EF9D" box="[478,579,1700,1725]" pageId="10" pageNumber="77" type="holotype">holotype</typeStatus>
|
||
, WHT 0111E). An adult male,
|
||
<quantity id="953230F17E50E50A4581F974FB97EF9C" box="[944,1046,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="4.56" pageId="10" pageNumber="77" unit="mm" value="45.6">45.6 mm</quantity>
|
||
SVL; head moderately large (HL 27.9% of SVL), elongate (HW 60.0% of HL), narrow (HW 16.7% of SVL), distinct from neck; snout short (ES 58.9% of HW); snout length somewhat greater than eye diameter (ED 98.0% of ES); interorbital distance broad (IO 11.7% of HL); eye large (ED 34.7% of HL); pupil rounded; ear-opening shallow, its greatest diameter located dorsoventrally, ear-opening with enlarged, unkeeled scales, tympanum smaller than orbit (
|
||
<collectingCountry id="2ADDDD847E50E50A42A1F8E4FB4AEE6C" box="[1168,1227,1844,1868]" name="Chad" pageId="10" pageNumber="77">TYD</collectingCountry>
|
||
60.6% of ED); no spines above the tympanum but an enlarged keeled scale above the tympanum separated from the tympanum by two smaller scale rows; diameter of eyes greater than eye to ear distance (ED 168.7% of TYE); forehead concave; scales on snout keeled, smaller in size than those of the occipital region; scales on interorbital and supercilium area keeled; no nuchal or dorsal crest; rostral scale width greater than height, ventro-posteriorly in contact with the first supralabial, in contact posteriorly with the prenasal and three postrostral scales unequal in size; two supranasals, on each side of the nostrils, three postnasals approximately equal in size and one prenasal; nostril rounded and located in the middle of an undivided nasal plate; three postnasals bordering the nasal, the middle one largest;
|
||
<emphasis id="60BE41067E51E50B4374F96BFF74EFD8" italics="true" pageId="11" pageNumber="78">canthus rostralis</emphasis>
|
||
and superciliary edges sharp; parietal plate lozenge shaped (anterior sides shorter than posterior sides) including a grey-blue coloured “pineal eye”, parietal plate larger than adjacent plates. Mental pentagonal, width and length of which approximately equal, posteriolaterally in contact with two enlarged postmentals separated by a smaller scale preventing contact between them; each postmental pair bordered posteriorly by three (left side) and four (right side) smooth scales including the medial scale, but exclusive of infralabials. Throat-fan small, posterior scales do not extend to mid venter (DWL 60.3% of SVL, ADW 32.1% of AG), margin of the anterior part of the throat-fan covered with keeled, overlapping, bluntly pointed scales, posterior part (from the middle of the throat to the midventer) with long, elongate lanceolate bluntly pointed scales; throat scales elongate, keeled and overlapping; three scale rows separating orbit from supralabials; supralabials 8 (6th in mid orbit position); infralabials 9, decreasing in size towards gape; scales on the throat-fan and adjacent ventral scales keeled and mucronate, overlapping.
|
||
</paragraph>
|
||
<caption id="06B5CD9C7E51E50B46A6FA28FB07EF6B" httpUri="https://zenodo.org/record/287775/files/figure.png" pageId="11" pageNumber="78" targetBox="[311,1275,179,1502]" targetPageId="11">
|
||
<paragraph id="52759D147E51E50B46A6FA28FB07EF6B" blockId="11.[151,1436,1527,1611]" pageId="11" pageNumber="78">
|
||
<emphasis id="60BE41067E51E50B46A6FA28FE95EF2D" bold="true" box="[151,276,1528,1550]" pageId="11" pageNumber="78">FIGURE 4.</emphasis>
|
||
Holotype male WHT 0111E of
|
||
<taxonomicName id="95CAE6977E51E50B446EFA29FD77EF2E" box="[607,758,1528,1550]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E51E50B446EFA29FD77EF2E" box="[607,758,1528,1550]" italics="true" pageId="11" pageNumber="78">Sitana devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E51E50B44CCFA28FCCEEF2D" box="[765,847,1528,1549]" pageId="11" pageNumber="78" rank="species">
|
||
<emphasis id="60BE41067E51E50B44CCFA28FCCEEF2D" bold="true" box="[765,847,1528,1549]" pageId="11" pageNumber="78">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, (
|
||
<emphasis id="60BE41067E51E50B4554FA27FC01EF2E" bold="true" box="[869,896,1527,1550]" pageId="11" pageNumber="78">A)</emphasis>
|
||
dorsal head view; (
|
||
<emphasis id="60BE41067E51E50B427DFA27FBE7EF2E" bold="true" box="[1100,1126,1527,1550]" pageId="11" pageNumber="78">B)</emphasis>
|
||
lateral head view; (
|
||
<emphasis id="60BE41067E51E50B4302FA27FACFEF2E" bold="true" box="[1331,1358,1527,1550]" pageId="11" pageNumber="78">C)</emphasis>
|
||
ventral head view; (
|
||
<emphasis id="60BE41067E51E50B472DF9C6FEB7EF0D" bold="true" box="[284,310,1558,1581]" pageId="11" pageNumber="78">D)</emphasis>
|
||
ventral thigh and tail base (photo: M.C. De Silva); (
|
||
<emphasis id="60BE41067E51E50B4543F9C7FC02EF0C" bold="true" box="[882,899,1559,1580]" pageId="11" pageNumber="78">E</emphasis>
|
||
)
|
||
<taxonomicName id="95CAE6977E51E50B45A7F9C8FBB1EF0D" box="[918,1072,1559,1581]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E51E50B45A7F9C8FBB1EF0D" box="[918,1072,1559,1581]" italics="true" pageId="11" pageNumber="78">Sitana devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E51E50B420BF9C7FB0EEF0C" box="[1082,1167,1559,1580]" pageId="11" pageNumber="78" rank="species">
|
||
<emphasis id="60BE41067E51E50B420BF9C7FB0EEF0C" bold="true" box="[1082,1167,1559,1580]" pageId="11" pageNumber="78">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, live breeding male with expanded throat-fan (not collected) at Wilpaththu National Park, Sri Lanka (photo: V. Weeratunge).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="52759D147E56E50C46F6FED6FC1FE8D7" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">Body slender (AG 50.0% of SVL); mid dorsal scales equal, keeled, not elongate with pointed dorsal scales at midbody, keels straight, forming longitudinal continuous parallel ridges; scales on dorsum at midbody larger in size than those of venter at the same level; lateral body scales unequal, keeled, rounded, smaller than dorsals and with randomly distributed enlarged, rounded, keeled scales; upper dorsolateral scales directed upwards and backwards, dorsoventral ones directed downwards and backwards; 59 scale rows around midbody; pectoral scales enlarged, elongate and keeled; abdominal scales not enlarged, short, mucronate and keeled; ventral keels forming regular and parallel continuous ridges; no preanal or femoral pores. Ventrals 105.</paragraph>
|
||
<paragraph id="52759D147E56E50C46F6FDD3FD32EA37" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">Upper arm somewhat longer than lower arm (UAL 18.9% of SVL; LAL 18.1% of SVL); femur shorter than tibia (FEL 26.5% of SVL; TBL 36.7% of SVL; FEL 72.2% of TBL). Dorsal scales on fore and hind limbs enlarged, elongate, keeled, overlapping, and mucronate; ventral scales on upper and lower arm enlarged, slightly keeled, overlapping, and blunt ended; scales on dorsal and ventral surface of thigh and shank enlarged, keeled, overlapping and blunt ended; keels on dorsal and ventral arm and foot forming a series of continuous parallel ridges. Digits elongate, slender, all bearing slightly recurved claws; claws are sharp and elongate; subdigital lamellae entire and tri-mucronate, 16 subdigital lamellae on toe III and 23 on toe IV; inter-digital webbing absent; relative length of fingers 3> 4> 5> 2> 1, and toes 4> 3> 2> 1.</paragraph>
|
||
<paragraph id="52759D147E56E50C46F6FCF2FC21EAA3" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">
|
||
Tail complete (
|
||
<quantity id="953230F17E56E50C4743FCF2FE69EA1B" box="[370,488,802,827]" metricMagnitude="-1" metricUnit="m" metricValue="1.258" pageId="12" pageNumber="79" unit="mm" value="125.8">125.8 mm</quantity>
|
||
) and base swollen, ventral scales on tail base rounded and keeled; dorsal scales on tail keeled, elongate, overlapping, directed backwards, keels forming continuous parallel ridges; tail with subcaudals on median row not enlarged, slightly keeled, overlapping.
|
||
</paragraph>
|
||
<paragraph id="52759D147E56E50C46F6FC5EFE54ED33" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">
|
||
<emphasis id="60BE41067E56E50C46F6FC5EFDC9EA87" bold="true" box="[199,584,910,935]" pageId="12" pageNumber="79">Colour in preserved specimen.</emphasis>
|
||
Dorsum light brown, darker brown patch on the neck, five rhomboid markings on the back. Light coloured blotches on lateral sides of the body, white coloured incomplete cross bars on the tail beginning just behind the hind limbs, a persian blue stripe runs from mental and continues up to mid dewlap point along mid ventral line.
|
||
</paragraph>
|
||
<paragraph id="52759D147E56E50C46F6FBCEFE49EC5B" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">
|
||
<emphasis id="60BE41067E56E50C46F6FBCEFEECED17" bold="true" box="[199,365,1054,1079]" pageId="12" pageNumber="79">Colour in life.</emphasis>
|
||
Based on the specimen in
|
||
<figureCitation id="CAF181917E56E50C44ABFBCEFD5FED17" box="[666,734,1054,1079]" captionStart="FIGURE 4" captionStartId="11.[151,250,1528,1550]" captionTargetBox="[311,1275,179,1502]" captionTargetId="figure@11.[311,1275,179,1503]" captionTargetPageId="11" captionText="FIGURE 4. Holotype male WHT 0111 E of Sitana devakai sp. nov., (A) dorsal head view; (B) lateral head view; (C) ventral head view; (D) ventral thigh and tail base (photo: M. C. De Silva); (E) Sitana devakai sp. nov., live breeding male with expanded throat-fan (not collected) at Wilpaththu National Park, Sri Lanka (photo: V. Weeratunge)." httpUri="https://zenodo.org/record/287775/files/figure.png" pageId="12" pageNumber="79">Fig. 4</figureCitation>
|
||
. Upper head and back dorsally cream to yellowish brown with five black-edged rhomboidal spots, the back is bordered on each side with a dark brownish band, a light vertebral line along the middle of the back separating them. Upper and lower lip a light buff cream with dark markings. A buff light cream streak from the nasal across the tympanum to the forelimb; pupil black. Iris black with a golden outer rim; reddish-brown patch on anterior part of interorbital. Anterior part of the throat-fan is persian blue, black and cream. A bright red coloured patch at the middle of the throat-fan on the cream coloured background during the breeding season. Upper arm, lower arm, thigh and tibia all with three indistinct buff cream bands. Vague, light patches on flank and lateral body. Tail with 20 light cross-bands. Three buff cross-bands on fingers and three or four on toes. Ventral scales buff cream. See the following corresponding figures in
|
||
<bibRefCitation id="365BE0E57E56E50C42A9FAEEFA19EC77" author="Manthey" box="[1176,1432,1342,1367]" pageId="12" pageNumber="79" refString="Manthey, U. (2010) Agamid Lizards of Southern Asia, Draconinae 2 & Leiolepidinae. Edition Chimaira & A. C. S. Glaser, Germany, Frankfurt am Main, 168 pp." type="book" year="2010">Manthey (2010: 154)</bibRefCitation>
|
||
: RA04168-4 (Wiplattu NP).
|
||
</paragraph>
|
||
<paragraph id="52759D147E56E50C46F6FA56FB42EF2B" blockId="12.[151,1437,151,2015]" pageId="12" pageNumber="79">
|
||
<emphasis id="60BE41067E56E50C46F6FA56FD9FECBF" bold="true" box="[199,542,1414,1439]" pageId="12" pageNumber="79">
|
||
Variation in male
|
||
<typeStatus id="8D7123B67E56E50C4791FA56FD98ECBF" box="[416,537,1414,1439]" pageId="12" pageNumber="79" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Nine supralabials in
|
||
<typeStatus id="8D7123B67E56E50C4521FA58FCFAEC80" box="[784,891,1416,1440]" pageId="12" pageNumber="79" type="paratype">paratypes</typeStatus>
|
||
WHT 0174A–B and WHT 1424A; respectively 57, 58, and 58 scales around midbody in WHT 1417, WHT 0174B, and WHT 1424A; subdigital lamellae on the toe III,
|
||
<quantity id="953230F17E56E50C46DBFA1EFEA2ECC7" box="[234,291,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="12" pageNumber="79" unit="in" value="15.0">15 in</quantity>
|
||
all other male
|
||
<typeStatus id="8D7123B67E56E50C47FBFA00FDB9ECC8" box="[458,568,1488,1512]" pageId="12" pageNumber="79" type="paratype">paratypes</typeStatus>
|
||
, on toe IV, 21 (WHT 1417) and 22 (WHT 0174A and WHT 1424A). Ventrals respectively 100, 101, 102, and 108 (WHT 1424, WHT 0174B, WHT 1417, and WHT 0174A).
|
||
</paragraph>
|
||
<paragraph id="52759D147E56E50D46F6F9C6FF69E863" blockId="12.[151,1437,151,2015]" lastBlockId="13.[151,1437,151,2015]" lastPageId="13" lastPageNumber="80" pageId="12" pageNumber="79">
|
||
<emphasis id="60BE41067E56E50C46F6F9C6FE51EF0F" bold="true" box="[199,464,1558,1583]" pageId="12" pageNumber="79">Description of female.</emphasis>
|
||
(Based on
|
||
<typeStatus id="8D7123B67E56E50C4465F9C8FD35EF10" box="[596,692,1560,1584]" pageId="12" pageNumber="79" type="paratype">paratype</typeStatus>
|
||
WHT 0111C). An adult female,
|
||
<quantity id="953230F17E56E50C421FF9C7FB12EF0F" box="[1070,1171,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="12" pageNumber="79" unit="mm" value="44.9">44.9 mm</quantity>
|
||
SVL; head moderately large (HL 28.3% of SVL), elongate (HW 61.9% of HL), narrow (HW 17.5% of SVL), distinct from neck; snout short (ES 59.5% of HW); snout length approximately equal to the eye diameter (ED 99.6% of ES); interorbital distance broad (IO 12.1% of HL); eye large (ED 36.7% of HL); pupil rounded; ear-opening shallow, its greatest diameter being dorsoventral, ear-opening with enlarged keeled scales, tympanum smaller than orbit (
|
||
<collectingCountry id="2ADDDD847E56E50C4321F977FACAEF9F" box="[1296,1355,1703,1727]" name="Chad" pageId="12" pageNumber="79">TYD</collectingCountry>
|
||
31.2% of ED); no spines above the tympanum but an enlarged keeled scale above the tympanum separated from the tympanum by three smaller scale rows; diameter of eyes greater than eye to ear distance (ED 166.2% of TYE); forehead concave; scales on snout keeled, more or less similar in size than those of the occipital region; scales on interorbital and supercilium area keeled; no nuchal or dorsal crest; rostral scale width greater than height, ventroposteriorly in contact with first supralabial, in contact posteriorly with the prenasal and three postrostral scales unequal in size; two supranasals around nostrils on each side, three postnasals with the middle one larger; nostril rounded located in the middle of an undivided nasal plate;
|
||
<emphasis id="60BE41067E56E50C4503F872FC6EEE9B" box="[818,1007,1954,1979]" italics="true" pageId="12" pageNumber="79">canthus rostralis</emphasis>
|
||
and superciliary edges sharp; parietal plate lozenge shaped (anterior sides shorter than posterior sides) including a grey-blue coloured “pineal eye”, parietal plate larger than adjacent plates. Mental pentagonal, approximately equal in length and width, posteriolaterally in contact with two enlarged postmentals separated by a smaller scale preventing contact between them. Throat-fan absent; throat scales elongate, keeled and overlapping; three scale rows separate orbit from supralabials; supralabials 8 (6th in mid orbit position); infralabials 10, decreasing in size towards gape; gular scales keeled.
|
||
</paragraph>
|
||
<paragraph id="52759D147E57E50D46F6FE9EFAF6EB3B" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">Body slender (AG 45.7% of SVL); mid dorsal scales equal, keeled, not elongate with pointed dorsal scales at midbody, keels straight forming longitudinal, continuous, parallel ridges; lateral body scales unequal, irregular, keeled, mucronate, smaller than dorsals and with randomly distributed enlarged, rounded, keeled scales; upper dorsolateral scales directed upwards and backwards, dorsoventral ones directed downwards and backwards; 57 scales around midbody; pectoral scales not enlarged, keeled, rounded and overlapping; abdominal scales short and rounded; ventral keels forming regular and parallel continuous ridges; no preanal or femoral pores. Ventrals 74.</paragraph>
|
||
<paragraph id="52759D147E57E50D46F6FDF8FE70EA1B" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">Upper arm longer than lower arm (UAL 19.5% of SVL; LAL 17.1% of SVL); femur shorter than tibia (FEL 25.9% of SVL; TBL 32.6% of SVL; FEL 79.4% of TBL). Dorsal scales on fore and hind limbs enlarged, elongate, keeled, overlapping, and mucronate; ventral scales on upper and lower arm enlarged, slightly keeled, overlapping, and bluntly ended; scales on dorsal and ventral surface of thigh slightly keeled, overlapping and bluntly ended; keels on dorsal and ventral arm and foot forming a series of continuous parallel ridges. Digits elongate, slender, all bearing slightly recurved claws; claws are sharp and elongate; subdigital lamellae entire and tri-mucronate, 15 subdigital lamellae on the toe III, and 25 on toe IV; inter-digital webbing absent; relative length of fingers 3> 4> 5> 2> 1, and toes 4> 3> 2> 1.</paragraph>
|
||
<paragraph id="52759D147E57E50D46F6FC96FD7AEA87" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">Tail complete and base swollen, ventral scales on tail base rounded and keeled; dorsal scales on tail keeled, elongate, overlapping, directed backwards, keels forming continuous parallel ridges; tail with subcaudals on median row not enlarged, slightly keeled, overlapping.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E57E50D46F6FC62FD2DEDCB" pageId="13" pageNumber="80" type="etymology">
|
||
<paragraph id="52759D147E57E50D46F6FC62FD2DEDCB" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">
|
||
<emphasis id="60BE41067E57E50D46F6FC62FECFEAEB" bold="true" box="[199,334,946,971]" pageId="13" pageNumber="80">Etymology.</emphasis>
|
||
The species epithet is an eponym Latinised in the genitive singular, honouring Prof. Devaka Keerthi Weerakoon, for his remarkable contributions to biodiversity conservation in
|
||
<collectingCountry id="2ADDDD847E57E50D4266FC06FB4BEACF" box="[1111,1226,982,1007]" name="Sri Lanka" pageId="13" pageNumber="80">Sri Lanka</collectingCountry>
|
||
. He worked as the Technical Advisor of the
|
||
<date id="2674BBD47E57E50D4780FC2BFDD7ED33" box="[433,598,1018,1043]" pageId="13" pageNumber="80" value="2007" valueMax="2012">2007 and 2012</date>
|
||
Red Lists of Threatened Fauna and Flora in
|
||
<collectingCountry id="2ADDDD847E57E50D4279FC2AFB34ED33" box="[1096,1205,1018,1043]" name="Sri Lanka" pageId="13" pageNumber="80">Sri Lanka</collectingCountry>
|
||
and was responsible for their publication. As a consultant to the Department of Wildlife Conservation, he has provided technical support for the preparation and implementation of protected areas, produced management plans, managed alien invasive species programs in protected areas, mapped habitats, carried out protected area gap analyses programs and conducted bird surveys. His contributions to the ecology of Asian elephants and reduction of human-elephant conflicts are remarkable. Suggested English name: Devaka’s Fan-throated Lizard; Sinhala (local) name: Devakagë Theli Katussa; French name: Sitane de Devaka.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1AD0CE9F7E57E50E46F6FB26FEE8E802" lastPageId="14" lastPageNumber="81" pageId="13" pageNumber="80" type="distribution">
|
||
<paragraph id="52759D147E57E50D46F6FB26FAD9EC13" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">
|
||
<emphasis id="60BE41067E57E50D46F6FB26FEE1EC2F" bold="true" box="[199,352,1270,1295]" pageId="13" pageNumber="80">Distribution.</emphasis>
|
||
See map
|
||
<figureCitation id="CAF181917E57E50D47FCFB26FD93EC2F" box="[461,530,1270,1295]" captionStart="FIGURE 9" captionStartId="21.[151,250,1528,1550]" captionTargetBox="[311,1267,179,1495]" captionTargetId="figure@21.[311,1275,179,1503]" captionTargetPageId="21" captionText="FIGURE 9. Distribution map of S. ponticeriana (yellow), S. deccanensis (green), Sitana cf. devakai (purple), S. cf. ponticeriana (orange), Sitana sivalensis complex (grey), S. bahiri sp. nov. (blue); S. devakai sp. nov. (black); and S. ponticeriana mucronata (red). Holotype locality of each Sri Lankan species marked with squares." httpUri="https://zenodo.org/record/287780/files/figure.png" pageId="13" pageNumber="80">Fig. 9</figureCitation>
|
||
. Drier coastal areas in the north and north-western parts of
|
||
<collectingCountry id="2ADDDD847E57E50D4298FB26FA98EC2F" box="[1193,1305,1270,1295]" name="Sri Lanka" pageId="13" pageNumber="80">Sri Lanka</collectingCountry>
|
||
, sometimes recorded at a distance of ~
|
||
<quantity id="953230F17E57E50D478FFACAFDB1EC13" box="[446,560,1306,1331]" metricMagnitude="4" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="13" pageNumber="80" unit="km" value="35.0" valueMax="60.0" valueMin="10.0">10–60 km</quantity>
|
||
from the nearest beach (e.g. Anuradhapura, Vavuniya and Kilinochchi).
|
||
</paragraph>
|
||
<paragraph id="52759D147E57E50D46F6FAEEFE35EF0F" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">
|
||
<emphasis id="60BE41067E57E50D46F6FAEEFE04EC77" bold="true" box="[199,389,1342,1367]" pageId="13" pageNumber="80">Natural history.</emphasis>
|
||
Most of the occupied habitats have similar environmental conditions to those of
|
||
<taxonomicName id="95CAE6977E57E50D4322FAEFFAF2EC77" box="[1299,1395,1342,1367]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="bahiri" status="sp. nov.">
|
||
<emphasis id="60BE41067E57E50D4322FAEFFAF2EC77" box="[1299,1395,1342,1367]" italics="true" pageId="13" pageNumber="80">S. bahiri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E57E50D434BFAEEFF49EC5B" pageId="13" pageNumber="80" rank="species">
|
||
<emphasis id="60BE41067E57E50D434BFAEEFF49EC5B" bold="true" pageId="13" pageNumber="80">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, comprising the following species of trees:
|
||
<taxonomicName id="95CAE6977E57E50D449FFAB2FB8CEC5B" box="[686,1037,1378,1403]" class="Magnoliopsida" family="Rhamnaceae" genus="Ziziphus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="rugosa">
|
||
<emphasis id="60BE41067E57E50D449FFAB2FCE3EC5B" box="[686,866,1378,1403]" italics="true" pageId="13" pageNumber="80">Ziziphus rugosa</emphasis>
|
||
(Rhamnaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D4226FAB2FA19EC5B" box="[1047,1432,1378,1403]" class="Magnoliopsida" family="Sapotaceae" genus="Manilkara" kingdom="Plantae" order="Ericales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="hexandra">
|
||
<emphasis id="60BE41067E57E50D4226FAB2FB7FEC5B" box="[1047,1278,1378,1403]" italics="true" pageId="13" pageNumber="80">Manilkara hexandra</emphasis>
|
||
(Sapotaceae)
|
||
</taxonomicName>
|
||
, and most of shrubs and herbs like
|
||
<taxonomicName id="95CAE6977E57E50D4424FA56FC36ECBF" box="[533,951,1414,1439]" class="Magnoliopsida" family="Malvaceae" genus="Abelmoschus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="angulosus">
|
||
<emphasis id="60BE41067E57E50D4424FA56FCA3ECBF" box="[533,802,1414,1439]" italics="true" pageId="13" pageNumber="80">Abelmoschus angulosus</emphasis>
|
||
(Malvaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D45F3FA56FABBECBF" box="[962,1338,1414,1439]" class="Magnoliopsida" family="Rubiaceae" genus="Benkara" kingdom="Plantae" order="Gentianales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="malabarica">
|
||
<emphasis id="60BE41067E57E50D45F3FA56FB28ECBF" box="[962,1193,1414,1439]" italics="true" pageId="13" pageNumber="80">Benkara malabarica</emphasis>
|
||
(Rubiaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D4374FA57FE3EECE3" class="Magnoliopsida" family="Apocynaceae" genus="Carissa" kingdom="Plantae" order="Gentianales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="spinarum">
|
||
<emphasis id="60BE41067E57E50D4374FA57FE80ECE3" italics="true" pageId="13" pageNumber="80">Carissa spinarum</emphasis>
|
||
(Apocynaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D47E4FA7AFCFAECE3" box="[469,891,1450,1475]" class="Magnoliopsida" family="Rubiaceae" genus="Hydrophylax" kingdom="Animalia" order="Gentianales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="maritima">
|
||
<emphasis id="60BE41067E57E50D47E4FA7AFD5FECE3" box="[469,734,1450,1475]" italics="true" pageId="13" pageNumber="80">Hydrophylax maritima</emphasis>
|
||
(Rubiaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D45A0FA7BFAADECE3" box="[913,1324,1450,1475]" class="Magnoliopsida" family="Capparaceae" genus="Capparis" kingdom="Plantae" order="Capparales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="zeylanica">
|
||
<emphasis id="60BE41067E57E50D45A0FA7BFBF4ECE3" box="[913,1141,1450,1475]" italics="true" pageId="13" pageNumber="80">Capparis zeylanica</emphasis>
|
||
(Capparaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D4373FA7BFEF4ECC7" class="Liliopsida" family="Poaceae" genus="Spinifex" kingdom="Plantae" order="Poales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="littoreus">
|
||
<emphasis id="60BE41067E57E50D4373FA7BFF75ECC7" italics="true" pageId="13" pageNumber="80">Spinifex littoreus</emphasis>
|
||
(Poaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D47B8FA1EFCDFECC7" box="[393,862,1486,1511]" class="Magnoliopsida" family="Rubiaceae" genus="Canthium" kingdom="Plantae" order="Gentianales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="coromandelicum">
|
||
<emphasis id="60BE41067E57E50D47B8FA1EFD42ECC7" box="[393,707,1486,1511]" italics="true" pageId="13" pageNumber="80">Canthium coromandelicum</emphasis>
|
||
(Rubiaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D4542FA1EFA90ECC7" box="[883,1297,1486,1511]" class="Reptilia" family="Colubridae" genus="Salvadora" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="persica">
|
||
<emphasis id="60BE41067E57E50D4542FA1EFBC8ECC7" box="[883,1097,1486,1511]" italics="true" pageId="13" pageNumber="80">
|
||
<collectingCountry id="2ADDDD847E57E50D4542FA1EFC67ECC7" box="[883,998,1486,1511]" name="El Salvador" pageId="13" pageNumber="80">Salvadora</collectingCountry>
|
||
persica
|
||
</emphasis>
|
||
(Salvadoraceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D4314FA1EFE35EF2B" class="Magnoliopsida" family="Apocynaceae" genus="Calotropis" kingdom="Plantae" order="Gentianales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="gigantea">
|
||
<emphasis id="60BE41067E57E50D4314FA1EFF78EF2B" italics="true" pageId="13" pageNumber="80">Calotropis gigantea</emphasis>
|
||
(Apocynaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D47F7FA23FC05EF2B" box="[454,900,1522,1547]" class="Magnoliopsida" family="Aizoaceae" genus="Sesuvium" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="portulacastrum">
|
||
<emphasis id="60BE41067E57E50D47F7FA23FD6DEF2B" box="[454,748,1522,1547]" italics="true" pageId="13" pageNumber="80">Sesuvium portulacastrum</emphasis>
|
||
(Aizoaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="95CAE6977E57E50D45A6FA22FB77EF2B" box="[919,1270,1522,1547]" class="Magnoliopsida" family="Fabaceae" genus="Indigofera" kingdom="Plantae" order="Fabales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="linnaei">
|
||
<emphasis id="60BE41067E57E50D45A6FA22FBE8EF2B" box="[919,1129,1522,1547]" italics="true" pageId="13" pageNumber="80">Indigofera linnaei</emphasis>
|
||
(Fabaceae)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="95CAE6977E57E50D4371FA23FE31EF0F" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="80" phylum="Tracheophyta" rank="species" species="monacantha">
|
||
<emphasis id="60BE41067E57E50D4371FA23FEA3EF0F" italics="true" pageId="13" pageNumber="80">Opuntia monacantha</emphasis>
|
||
(Cactaceae)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="52759D147E57E50D46F6F9EBFBA6EEFF" blockId="13.[151,1437,151,2015]" pageId="13" pageNumber="80">
|
||
<taxonomicName id="95CAE6977E57E50D46F6F9EBFEF1EF73" box="[199,368,1594,1619]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E57E50D46F6F9EBFEF1EF73" box="[199,368,1594,1619]" italics="true" pageId="13" pageNumber="80">Sitana devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E57E50D474BF9EAFE56EF73" box="[378,471,1594,1619]" pageId="13" pageNumber="80" rank="species">
|
||
<emphasis id="60BE41067E57E50D474BF9EAFE56EF73" bold="true" box="[378,471,1594,1619]" pageId="13" pageNumber="80">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is a diurnal lizard completely suited to harsh terrestrial habitats such as dry sandy coasts (
|
||
<figureCitation id="CAF181917E57E50D46C0F98EFECAEF57" box="[241,331,1630,1655]" captionStart="FIGURE 10" captionStartId="22.[151,250,1928,1950]" captionTargetBox="[160,1395,222,1889]" captionTargetId="figure@22.[160,1396,193,1889]" captionTargetPageId="22" captionText="FIGURE 10. Habitat of Sitana bahiri sp. nov. (A) dry coastal area in Palatupana, Kirinda, Sri Lanka; (B) coast covered with Spinifex littorus (photo: A. A. T. Amarasinghe); habitat of Sitana devakai sp. nov. (C) dry warm sandy area in Puttalam, Sri Lanka; (D) area covered with Abelmoschus angulosus (photo: D. M. S. S. Karunarathna)." httpUri="https://zenodo.org/record/287781/files/figure.png" pageId="13" pageNumber="80">Fig. 10</figureCitation>
|
||
). Copulation is limited to a 5–10 second activity. Egg laying activities were observed from September to November. They lay 3–
|
||
<specimenCount id="44CC569D7E57E50D4473F952FDD1EFBB" box="[578,592,1666,1691]" pageId="13" pageNumber="80" type="egg">6</specimenCount>
|
||
eggs per clutch with a mean size of 9.2 ×
|
||
<quantity id="953230F17E57E50D4206F953FB0FEFBA" box="[1079,1166,1667,1691]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="13" pageNumber="80" unit="mm" value="4.7">4.7 mm</quantity>
|
||
(
|
||
<emphasis id="60BE41067E57E50D42AEF954FB2CEFBB" box="[1183,1197,1668,1691]" italics="true" pageId="13" pageNumber="80">n</emphasis>
|
||
=18). The incubation period is between 40–50 days. Hatchlings always live hidden inside shrubs during the first month after hatching.
|
||
<taxonomicName id="95CAE6977E57E50D46A6F91AFED9EFC3" box="[151,344,1738,1763]" class="Aves" family="Leiothrichidae" genus="Turdoides" kingdom="Animalia" order="Passeriformes" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="60BE41067E57E50D46A6F91AFED9EFC3" box="[151,344,1738,1763]" italics="true" pageId="13" pageNumber="80">Turdoides affinis</emphasis>
|
||
</taxonomicName>
|
||
(common babbler),
|
||
<taxonomicName id="95CAE6977E57E50D447CF91BFCA7EFC3" box="[589,806,1739,1763]" class="Aves" family="Cuculidae" genus="Centropus" kingdom="Animalia" order="Cuculiformes" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="60BE41067E57E50D447CF91BFCA7EFC3" box="[589,806,1739,1763]" italics="true" pageId="13" pageNumber="80">Centropus sinensis</emphasis>
|
||
</taxonomicName>
|
||
(greater coucal),
|
||
<taxonomicName id="95CAE6977E57E50D45CAF91AFB62EFC3" box="[1019,1251,1738,1763]" class="Aves" family="Alcedinidae" genus="Halcyon" kingdom="Animalia" order="Coraciiformes" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="smyrnensis">
|
||
<emphasis id="60BE41067E57E50D45CAF91AFB62EFC3" box="[1019,1251,1738,1763]" italics="true" pageId="13" pageNumber="80">Halcyon smyrnensis</emphasis>
|
||
</taxonomicName>
|
||
(white-throated kingfisher),
|
||
<taxonomicName id="95CAE6977E57E50D4714F93EFE43EE27" box="[293,450,1774,1799]" class="Aves" family="Ardeidae" genus="Bubulcus" kingdom="Animalia" order="Pelecaniformes" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="ibis">
|
||
<emphasis id="60BE41067E57E50D4714F93EFE43EE27" box="[293,450,1774,1799]" italics="true" pageId="13" pageNumber="80">Bubulcus ibis</emphasis>
|
||
</taxonomicName>
|
||
(cattle egret),
|
||
<taxonomicName id="95CAE6977E57E50D4444F920FCDFEE27" box="[629,862,1774,1799]" class="Mammalia" family="Herpestidae" genus="Herpestes" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="edwardsii">
|
||
<emphasis id="60BE41067E57E50D4444F920FCDFEE27" box="[629,862,1774,1799]" italics="true" pageId="13" pageNumber="80">Herpestes edwardsii</emphasis>
|
||
</taxonomicName>
|
||
(grey mongoose),
|
||
<taxonomicName id="95CAE6977E57E50D4273F93EFA93EE27" box="[1090,1298,1774,1799]" class="Mammalia" family="Viverridae" genus="Viverricula" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="indica">
|
||
<emphasis id="60BE41067E57E50D4273F93EFA93EE27" box="[1090,1298,1774,1799]" italics="true" pageId="13" pageNumber="80">Viverricula indica</emphasis>
|
||
</taxonomicName>
|
||
(ring-tailed civet),
|
||
<taxonomicName id="95CAE6977E57E50D46D6F8C4FE51EE0B" box="[231,464,1810,1835]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="bengalensis">
|
||
<emphasis id="60BE41067E57E50D46D6F8C4FE51EE0B" box="[231,464,1810,1835]" italics="true" pageId="13" pageNumber="80">Varanus bengalensis</emphasis>
|
||
</taxonomicName>
|
||
(land monitor) and
|
||
<taxonomicName id="95CAE6977E57E50D4489F8C2FC18EE0B" box="[696,921,1810,1835]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="tristis">
|
||
<emphasis id="60BE41067E57E50D4489F8C2FC18EE0B" box="[696,921,1810,1835]" italics="true" pageId="13" pageNumber="80">Dendrelaphis tristis</emphasis>
|
||
</taxonomicName>
|
||
(common bronzeback snake) are their major predators. Territorial behaviours are similar to those of
|
||
<taxonomicName id="95CAE6977E57E50D4570F8E7FC28EE6F" box="[833,937,1846,1871]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="bahiri" status="sp. nov.">
|
||
<emphasis id="60BE41067E57E50D4570F8E7FC28EE6F" box="[833,937,1846,1871]" italics="true" pageId="13" pageNumber="80">S. bahiri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E57E50D4589F8E6FB9BEE6F" box="[952,1050,1846,1871]" pageId="13" pageNumber="80" rank="species">
|
||
<emphasis id="60BE41067E57E50D4589F8E6FB9BEE6F" bold="true" box="[952,1050,1846,1871]" pageId="13" pageNumber="80">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="95CAE6977E57E50D4251F8E7FAE1EE6F" box="[1120,1376,1847,1871]" class="Reptilia" family="Agamidae" genus="Otocryptis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="wiegmanni">
|
||
<emphasis id="60BE41067E57E50D4251F8E7FAE1EE6F" box="[1120,1376,1847,1871]" italics="true" pageId="13" pageNumber="80">Otocryptis wiegmanni</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="365BE0E57E57E50D46A6F88AFDBBEE53" author="Karunarathna" box="[151,570,1882,1907]" pageId="13" pageNumber="80" refString="Karunarathna, D. M. S. S. & Amarasinghe, A. A. T. (2008) Notes on the territorial behaviour of Otocryptis wiegmanni Wagler, 1830 (Reptilia: Agamidae: Draconinae). Herpetotropicos, 4, 79 - 83." type="journal article" year="2008">Karunarathna & Amarasinghe, 2008</bibRefCitation>
|
||
). They also open their mouths to threaten invaders, but the mouth cavity is purple in colour (vs.
|
||
<taxonomicName id="95CAE6977E57E50D474EF8AFFE5EEEB7" box="[383,479,1918,1943]" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="80" phylum="Chordata" rank="species" species="bahiri" status="sp. nov.">
|
||
<emphasis id="60BE41067E57E50D474EF8AFFE5EEEB7" box="[383,479,1918,1943]" italics="true" pageId="13" pageNumber="80">S. bahiri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E57E50D47D4F8AEFDBFEEB7" box="[485,574,1918,1943]" pageId="13" pageNumber="80" rank="species">
|
||
<emphasis id="60BE41067E57E50D47D4F8AEFDBFEEB7" bold="true" box="[485,574,1918,1943]" pageId="13" pageNumber="80">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
blue in color). At midday when it is warm, they keep their tails erected while walking, possibly to avoid contact with the hot sandy ground. We observed strategic feeding habits such as following ants to find their nests and using “sit and wait” strategies for predation.
|
||
</paragraph>
|
||
<paragraph id="52759D147E54E50E46F6FF47FEE8E802" blockId="14.[151,1436,151,290]" pageId="14" pageNumber="81">
|
||
<emphasis id="60BE41067E54E50E46F6FF47FE3DE990" bold="true" box="[199,444,151,176]" pageId="14" pageNumber="81">Conservation status.</emphasis>
|
||
The result of the application of the IUCN Red List (2013) criteria B1-b (i) shows that
|
||
<taxonomicName id="95CAE6977E54E50E43B6FF48FF6FE9F7" class="Reptilia" family="Agamidae" genus="Sitana" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="81" phylum="Chordata" rank="species" species="devakai" status="sp. nov.">
|
||
<emphasis id="60BE41067E54E50E43B6FF48FF6FE9F7" italics="true" pageId="14" pageNumber="81">S. devakai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7B8DFC7D7E54E50E46C7FF6EFED3E9F7" box="[246,338,190,215]" pageId="14" pageNumber="81" rank="species">
|
||
<emphasis id="60BE41067E54E50E46C7FF6EFED3E9F7" bold="true" box="[246,338,190,215]" pageId="14" pageNumber="81">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is Vulnerable (VU). It is restricted to an area of occupancy (AOO) <
|
||
<quantity id="953230F17E54E50E425BFF6EFB40E9F7" box="[1130,1217,190,215]" metricMagnitude="5" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="81" unit="km" value="850.0">850 km</quantity>
|
||
2 (11 spotting sites) and its extent of occurrence (EOO) <
|
||
<quantity id="953230F17E54E50E4476FF35FD43E9DE" box="[583,706,229,254]" metricMagnitude="7" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="81" unit="km" value="11500.0">11,500 km</quantity>
|
||
<quantity id="953230F17E54E50E44F3FF32FD68E9DE" box="[706,745,226,254]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="81" unit="in" value="2.0">2 in</quantity>
|
||
the north and north-west dry zone within different forested areas in
|
||
<collectingCountry id="2ADDDD847E54E50E46C5FED9FEE7E802" box="[244,358,265,290]" name="Sri Lanka" pageId="14" pageNumber="81">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |