103 lines
11 KiB
XML
103 lines
11 KiB
XML
<document id="FBA7BEDAF2A1B57E52FFBC5D010CCD43" ID-CLB-Dataset="29652" ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZooBank="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva" docDate="2018" docId="08793DBABED289C5A14073A43198304E" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaginus scaphaner Szymczakowski 1972" docType="treatment" docVersion="6" lastPageNumber="75" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="75" updateTime="1701383097238" updateUser="ExternalLinkService">
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<mods:titleInfo id="170396E2A80B9058C88C1309F52531FB">
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<mods:title id="73DE53F044E2299B901C86101A53B988">A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
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<mods:name id="0ECB595E873CAD1686F44916DAE0DDFF" type="personal">
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<mods:namePart id="30554305C7FE7649F12732A579A8C604">Schilthuizen, Menno</mods:namePart>
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<mods:namePart id="B0715E5D62CB1982CC4669EC7F51BCBC">Perreau, Michel</mods:namePart>
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<mods:roleTerm id="5E3068AC866260D5CB3BCC4965A3171E">Author</mods:roleTerm>
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<mods:namePart id="241F3FEAFC5F8438C0C55E272CB99E0B">Njunjic, Iva</mods:namePart>
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<mods:typeOfResource id="CC555A136826F3B24251698AA8630692">text</mods:typeOfResource>
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<mods:title id="E22B15B48C064F809F57BB3137E3D5CB">ZooKeys</mods:title>
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<mods:part id="0F94FD7480196A99B32D033EEFE4733C">
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<mods:date id="24B83163148A7833A5112A2D0ACB4263">2018</mods:date>
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<mods:detail id="FCD82DAF03129A6AA819B6CAEC413614" type="volume">
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<mods:number id="89EBE6A6DF425775B4E52DEEEF88E9CA">777</mods:number>
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<mods:start id="68D6790AB82D3DF9AB090C1207364EDC">57</mods:start>
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<mods:url id="873AE8621F1C393E8140C324E0001D4D">http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
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<mods:classification id="35E68F9610D8645A27C2633AE25E58EE">journal article</mods:classification>
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<mods:identifier id="9B4904C4FD4CF03AF4D00C0B7BD4D275" type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
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<mods:identifier id="0FE429CF789930DF1D94A49582943966" type="Pensoft-Pub">1313-2970-777-57</mods:identifier>
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<treatment id="08793DBABED289C5A14073A43198304E" ID-GBIF-Taxon="146746181" LSID="urn:lsid:plazi:treatment:08793DBABED289C5A14073A43198304E" httpUri="http://treatment.plazi.org/id/08793DBABED289C5A14073A43198304E" lastPageNumber="75" pageId="18" pageNumber="75">
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<subSubSection id="649D64F8A825BA8085AAFC823765B974" pageId="18" pageNumber="75" type="nomenclature">
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<paragraph id="E8306C94055E2542AB4A11979465BFFC" pageId="18" pageNumber="75">
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<taxonomicName id="E0706BF57DC6FFCFF65FE7DA5D1ADD78" ID-CoL="78LQ3" authority="Szymczakowski, 1972" authorityName="Szymczakowski" authorityYear="1972" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus scaphaner" order="Coleoptera" pageId="18" pageNumber="75" phylum="Arthropoda" rank="species" species="scaphaner">Ptomaphaginus scaphaner Szymczakowski, 1972</taxonomicName>
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Figures 5c, 7c, d, 9i, 10f
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</paragraph>
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</subSubSection>
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<subSubSection id="FFD4C1BCFA97A65CCAB59C92A79DBCAD" pageId="18" pageNumber="75" type="reference_group">
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<paragraph id="50F94528F9D9FA7D573C9128D631B1C4" pageId="18" pageNumber="75">
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<taxonomicName id="AC9BF498C9C0858DB7FE9BC5814CF4F4" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus scaphaner" order="Coleoptera" pageId="18" pageNumber="75" phylum="Arthropoda" rank="species" species="scaphaner">Ptomaphaginus scaphaner</taxonomicName>
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Szymczakowski, 1972: 279-300, figs 28-33); type from Cue phuong Ninh binh, Vietnam (in HNHM).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3567D966C0F1AA7F0C56962939DD5B5" pageId="18" pageNumber="75" type="materials_examined">
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<paragraph id="CB946DA0EC1C6A3DEF9576BBEA2008D6" pageId="18" pageNumber="75">Material.</paragraph>
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<paragraph id="35186895071DD2774F16E067D4164E5A" pageId="18" pageNumber="75">
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(in addition to that given in
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<bibRefCitation id="BF34787560D2158FAF95B26045C01437" author="Schilthuizen, M" journalOrPublisher="Zoologische Mededelingen Leiden" pageId="37" pageNumber="94" pagination="189 - 210" title="New species and new records of Ptomaphaginus Portevin from the Sunda region (Coleoptera: Leiodidae: Cholevinae)." volume="82" year="2008">Schilthuizen and Perreau (2008)</bibRefCitation>
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): Sabah. Kinabalu Park, Silau-Silau (low), 1530 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086209-1086214), 5 males, 1 female; Kinabalu Park, Bukit Ular trail (low), 1800 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086224-1086231), 3 males, 5 females; Kinabalu Park, Mempening trail (high), 1700 m elev., human excrement traps, 7-11.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086233), 1 male; Crocker Range, Kota Kinabalu-Tambunan road (km 56), 1350 m elev., fish traps, 21-24.xi.1987 (leg. Krikken & Rombaut, RMNH.INS.1086215-1086223), 2 males, 7 females; Kinabalu Park, Headquarters Area, Liwagu Trail,
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<geoCoordinate id="43E261EE409739F3D0D1389F4DF1C728" direction="north" orientation="latitude" precision="1" value="6.008111">6°00'29.2"N</geoCoordinate>
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<geoCoordinate id="B76B8C13B2675F5643D2213F4E1AB35C" direction="east" orientation="longitude" precision="1" value="116.5455">116°32'43.8"E</geoCoordinate>
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, pitfall with chicken, 2.iv.2016 (leg. M. Schilthuizen, I.
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<normalizedToken id="3DD28867C12C7EC693D0D01EEE3225B7" originalValue="Njunjić">Njunjic</normalizedToken>
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& F. Feijen) 2 males (BORN), 1 male and 1 female (RMNH.INS.1086232). Sarawak. Mulu National Park, Limestone 6.4, Tp 113, 400 m elev., 6.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male; Mulu National Park, Limestone 6.4, Tp 110, 650 m elev., 6.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male, 2 females; Mulu National Park, mixed dipterocarp forest litter, TPS 5-6,
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<normalizedToken id="80BAA585CE7B2A59DC29FD810275972F" originalValue="v–viii">v-viii</normalizedToken>
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.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 4 males, 1 female; Mulu National Park, mixed dipterocarp forest litter, Tp 7, 21.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 male; Mulu National Park, slope, baited traps, 7.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 3 females; Mulu National Park, slope, TPS 4-6, 29.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 female; Mulu National Park, TPS 7-13, 4.v.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 1 female.
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</paragraph>
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</subSubSection>
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<subSubSection id="8AE1AD37A940E061B1B629979AC2FA6F" pageId="18" pageNumber="75" type="description">
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<paragraph id="5D899BA9846820B78907EBA054AE9EB0" pageId="18" pageNumber="75">Description.</paragraph>
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<paragraph id="20D94458AEE420FD0A7B4E3AA559EFBB" pageId="18" pageNumber="75">
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Length 1.8-2.5 mm. Habitus broad and short, colouration light reddish-brown. Pronotum 1.75 times as wide as long, slightly narrower than the elytra. Elytra 1.2 times as long as jointly wide (length measured from the caudal tip of the scutellum to the caudal tip of the elytra). Wings present. Antennae very broad and short, with antennomeres 9 and 10 twice as broad as long. Aedeagus strongly curved, extremely convex and swollen, ending in a flattened
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<normalizedToken id="248067ECC7D4B19621EDC7A450DDF0F5" originalValue="“beak”">"beak"</normalizedToken>
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; stylet long and thin, hair-like, running along the inside of the roof of the convex part of the aedeagus. Spiculum gastrale short, triangular, as long as wide. Spermatheca U-shaped, with 6-7 broad rings along the proximal part, and ca. 5 much narrower rings in the terminal one-quarter. Spermiduct very long and very thin.
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</paragraph>
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</subSubSection>
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<subSubSection id="D2358F5E5AA94534E6C2904B16CAC368" pageId="18" pageNumber="75" type="differential diagnosis">
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<paragraph id="D7613CD88EC6271E1F5A99055833BB70" pageId="18" pageNumber="75">Differential diagnosis.</paragraph>
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<paragraph id="A82C49E9C3DFBD0151F4A2EFA541D051" pageId="18" pageNumber="75">
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Unique among the Bornean
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<taxonomicName id="7FC5E009DE96EAE5DBF7F5ACE3D6932C" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus" order="Coleoptera" pageId="18" pageNumber="75" phylum="Arthropoda" rank="genus">Ptomaphaginus</taxonomicName>
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by the condensed antennae and the inflated aedeagus with flattened
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<normalizedToken id="7317F0D26ACE27B5873EDF4C245210A7" originalValue="“beak”">"beak"</normalizedToken>
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and long stylet.
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</paragraph>
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</subSubSection>
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<subSubSection id="265C966F2AD41CCAB06A9FE717067709" pageId="18" pageNumber="75" type="habitat">
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<paragraph id="776B94780576703499CC531E83327CC3" pageId="18" pageNumber="75">Habitat and distribution.</paragraph>
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<paragraph id="FB8A8D842DEC168FD880F18B98298ECE" pageId="18" pageNumber="75">
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This appears to be a very widespread species, now known from Vietnam, Peninsular Malaysia, Borneo, and Java. It is possible, however, that the species consists of a complex of closely related species, given the geographic variation in secondary sexual characters (
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<bibRefCitation id="93FB4B2AECB8E19A501C8215C20D1169" author="Schilthuizen, M" journalOrPublisher="Zoologische Mededelingen Leiden" pageId="37" pageNumber="94" pagination="189 - 210" title="New species and new records of Ptomaphaginus Portevin from the Sunda region (Coleoptera: Leiodidae: Cholevinae)." volume="82" year="2008">Schilthuizen and Perreau 2008</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |