482 lines
92 KiB
XML
482 lines
92 KiB
XML
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<mods:title id="FDFFC638513CBED619A111B85EDF53A6">First Systematic Study using the Variability of the Residual Colour Patterns: The Case of the Paleogene Seraphsidae (Mollusca, Gastropoda, Stromboidea)</mods:title>
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<mods:namePart id="EA0CAF8B1CD959C3F1079CC5571FA7F1">Merle, Didier</mods:namePart>
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<taxonomicName id="4C0F52DDFF801E48FD3AFF2AFB3DFF27" ID-CoL="8YP2K" authority="(Cossmann, 1889)" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[732,1154,215,243]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="440" phylum="Mollusca" rank="species" species="chilophorus">
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<emphasis id="B97BF54CFF801E48FD3AFF2AFC0EFF26" bold="true" box="[732,945,215,242]" italics="true" pageId="23" pageNumber="440">Seraphs chilophorus</emphasis>
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(
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<bibRefCitation id="EF9E54AFFF801E48FC26FF25FBC5FF27" author="COSSMANN M." box="[960,1146,216,243]" pageId="23" pageNumber="440" pagination="3 - 385" refId="ref23409" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris (4 e fascicule). Annales de la Societe royale malacologique de Belgique 24: 3 - 385." type="journal article" year="1889">Cossmann, 1889</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="133435DBFF801E48FCACFF0AFC28FEC6" box="[842,919,247,274]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="23" pageNumber="440">Figs 15</figureCitation>
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A-I; 16; 17)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3157AD5FF801E48FD43FECDFBFAFDAF" pageId="23" pageNumber="440" type="reference_group">
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<treatmentCitation id="0AAE0F4FFF801E48FD43FECDFB0AFE9D" author="COSSMANN M." box="[677,1205,304,329]" page="98" pageId="23" pageNumber="440" year="1889">
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<taxonomicName id="4C0F52DDFF801E48FD43FECDFB0AFE9D" ID-CoL="8ZJ3F" authority="Cossmann, 1889: 98" authorityName="Cossmann" authorityPageNumber="98" authorityYear="1889" box="[677,1205,304,329]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="440" phylum="Mollusca" rank="species" species="chilophorum" subGenus="Seraphs">
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<emphasis id="B97BF54CFF801E48FD43FECDFCB4FE9C" box="[677,779,304,328]" italics="true" pageId="23" pageNumber="440">Terebellum</emphasis>
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(
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<emphasis id="B97BF54CFF801E48FCFFFECDFCE0FE9C" box="[793,863,304,328]" italics="true" pageId="23" pageNumber="440">Seraphs</emphasis>
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)
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<emphasis id="B97BF54CFF801E48FC8AFECDFC5DFE9C" box="[876,994,304,328]" italics="true" pageId="23" pageNumber="440">chilophorum</emphasis>
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<bibRefCitation id="EF9E54AFFF801E48FC0EFECCFB0AFE9D" author="COSSMANN M." box="[1000,1205,305,329]" pageId="23" pageNumber="440" pagination="3 - 385" refId="ref23409" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris (4 e fascicule). Annales de la Societe royale malacologique de Belgique 24: 3 - 385." type="journal article" year="1889">Cossmann, 1889: 98</bibRefCitation>
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</taxonomicName>
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, pl. 3, figs 5, 6;
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</paragraph>
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<paragraph id="8BB0295EFF801E48FD43FE86FCD3FE36" blockId="23.[676,1211,379,482]" pageId="23" pageNumber="440">
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<taxonomicName id="4C0F52DDFF801E48FD43FE86FC4FFE47" ID-CoL="8ZJ3F" authorityName="Cossmann" authorityYear="1889" box="[677,1008,379,403]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="440" phylum="Mollusca" rank="species" species="chilophorum" subGenus="Seraphs">
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<emphasis id="B97BF54CFF801E48FD43FE86FCB1FE47" box="[677,782,379,403]" italics="true" pageId="23" pageNumber="440">Terebellum</emphasis>
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||
(
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||
<emphasis id="B97BF54CFF801E48FCF9FE86FCD9FE47" box="[799,870,379,403]" italics="true" pageId="23" pageNumber="440">Seraphs</emphasis>
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)
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||
<emphasis id="B97BF54CFF801E48FC91FE86FC4FFE47" box="[887,1008,379,403]" italics="true" pageId="23" pageNumber="440">chilophorum</emphasis>
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</taxonomicName>
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||
–
|
||
<treatmentCitation id="0AAE0F4FFF801E48FBEBFE86FD7DFE79" author="COSSMANN M." page="45" pageId="23" pageNumber="440" year="1904">
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<bibRefCitation id="EF9E54AFFF801E48FBEBFE86FD7DFE79" author="COSSMANN M." pageId="23" pageNumber="440" refId="ref23501" refString="COSSMANN M. 1904. - Essais de Paleoconchologie comparee, sixieme livraison, Cossmann, Paris, 151 p." type="book" year="1904">Cossmann 1904: 45</bibRefCitation>
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</treatmentCitation>
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, pl. 2, fig. 6. —
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<bibRefCitation id="EF9E54AFFF801E48FC85FE68FBCCFE79" author="COSSMANN M. & PISSARRO G." box="[867,1139,405,429]" pageId="23" pageNumber="440" pagination="26 - 45" refId="ref23645" refString="COSSMANN M. & PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene des environs de Paris. Tome 2. Hermann, Paris, pls 26 - 45." type="book chapter" year="1911">Cossmann & Pissarro 1911</bibRefCitation>
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||
: pl. 32, fig. 158-5. —
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<treatmentCitation id="0AAE0F4FFF801E48FCD0FE4DFBB5FE1C" author="KOROBKOV I. A." box="[822,1034,432,456]" page="260" pageId="23" pageNumber="440" year="1955">
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<bibRefCitation id="EF9E54AFFF801E48FCD0FE4DFBB5FE1C" author="KOROBKOV I. A." box="[822,1034,432,456]" pageId="23" pageNumber="440" refId="ref24677" refString="KOROBKOV I. A. 1955. - Spravocnik i Metodiceskoe Rukovodstvo po treticnym Molluskam, Bruhonogie [Handbook and guide of Tertiary molluscs, Gastropods], Gostoptehizdat, Moscou, 795 p. (in Russian)." type="book" year="1955">Korobkov 1955: 260</bibRefCitation>
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</treatmentCitation>
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||
, pl. 57, fig. 3. —
|
||
<treatmentCitation id="0AAE0F4FFF801E48FD40FE37FCD7FE36" author="GLIBERT M." box="[678,872,458,482]" page="39" pageId="23" pageNumber="440" year="1963">
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<bibRefCitation id="EF9E54AFFF801E48FD40FE37FC96FE36" author="GLIBERT M." box="[678,809,458,482]" pageId="23" pageNumber="440" pagination="1 - 154" refId="ref24258" refString="GLIBERT M. 1963. - Les Mesogastropoda fossiles du Cenozoique etranger des collections de l'Institut royal des Sciences naturelles de Belgique. Deuxieme partie: Fossaridae a Ficidae (inclus). Memoires de l'Institut royal des Sciences naturelles de Belgique 2 (73): 1 - 154." type="journal article" year="1963">Glibert 1963</bibRefCitation>
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: p. 39
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB0295EFF801E48FD43FE07FBFAFDAF" blockId="23.[675,1210,506,635]" pageId="23" pageNumber="440">
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<treatmentCitationGroup id="AB1F0E70FF801E48FD43FE07FBFAFDAF" pageId="23" pageNumber="440">
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<taxonomicName id="4C0F52DDFF801E48FD43FE07FCE3FDC6" ID-CoL="8YP2K" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[677,860,506,530]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="440" phylum="Mollusca" rank="species" species="chilophorus">
|
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<emphasis id="B97BF54CFF801E48FD43FE07FCE3FDC6" box="[677,860,506,530]" italics="true" pageId="23" pageNumber="440">Seraphs chilophorus</emphasis>
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</taxonomicName>
|
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–
|
||
<treatmentCitation id="0AAE0F4FFF801E48FC91FE07FBB6FDC6" author="JUNG P." box="[887,1033,506,530]" page="22" pageId="23" pageNumber="440" year="1974">
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<bibRefCitation id="EF9E54AFFF801E48FC91FE07FBB6FDC6" author="JUNG P." box="[887,1033,506,530]" pageId="23" pageNumber="440" pagination="5 - 52" refId="ref24521" refString="JUNG P. 1974. - A revision of the family Seraphsidae (Gastropoda: Strombacea). Paleontographica Americana 8 (47): 5 - 52." type="journal article" year="1974">Jung 1974: 22</bibRefCitation>
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</treatmentCitation>
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, pl. 5, figs 12-21, text-fig. 15-16. —
|
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<treatmentCitation id="0AAE0F4FFF801E48FCB5FDE9FBA9FDF8" author="DOLIN L. & DOLIN C. & LE RENARD J." box="[851,1046,532,556]" page="28" pageId="23" pageNumber="440" year="1980">
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<bibRefCitation id="EF9E54AFFF801E48FCB5FDE9FBA9FDF8" author="DOLIN L. & DOLIN C. & LE RENARD J." box="[851,1046,532,556]" pageId="23" pageNumber="440" pagination="26 - 48" refId="ref23927" refString="DOLIN L., DOLIN C. & LE RENARD J. 1980. - Inventaire systematique des Mollusques de l'Auversien a " Facies charrie " de Baron (Oise), et remarques paleontologiques. Bulletin d'Information des Geologues du Bassin de Paris 17 (2): 26 - 48." type="journal article" year="1980">
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Dolin
|
||
<emphasis id="B97BF54CFF801E48FC69FDE8FC04FDF8" box="[911,955,532,556]" italics="true" pageId="23" pageNumber="440">et al.</emphasis>
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1980: 28
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</bibRefCitation>
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</treatmentCitation>
|
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. —
|
||
<treatmentCitation id="0AAE0F4FFF801E48FBDDFDE9FD74FD93" author="SAVAZZI E." page="324" pageId="23" pageNumber="440" year="1991">
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<bibRefCitation id="EF9E54AFFF801E48FBDDFDE9FD74FD93" author="SAVAZZI E." pageId="23" pageNumber="440" pagination="311 - 331" refId="ref26067" refString="SAVAZZI E. 1991. - Constructional morphology of strombid gastropods. Lethaia 24 (3): 311 - 331." type="journal article" year="1991">Savazzi 1991: 324</bibRefCitation>
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</treatmentCitation>
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, fig. 13I, J. —
|
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<treatmentCitation id="0AAE0F4FFF801E48FCBCFDD2FBAEFD93" author="LE RENARD J." box="[858,1041,559,583]" page="6" pageId="23" pageNumber="440" year="1992">
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<bibRefCitation id="EF9E54AFFF801E48FCBCFDD2FBAEFD93" author="LE RENARD J." box="[858,1041,559,583]" pageId="23" pageNumber="440" pagination="1 - 14" refId="ref25050" refString="LE RENARD J. 1992. - Notes de nomenclature sur quelques Mollusques caracteristiques de l'Eocene du Bassin de Paris. Cossmanniana 1 (2 - 4): 1 - 14." type="journal article" year="1992">Le Renard 1992: 6</bibRefCitation>
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</treatmentCitation>
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. —
|
||
<treatmentCitation id="0AAE0F4FFF801E48FBDFFDD2FCF1FDB5" author="LE RENARD J. & PACAUD J. - M." page="112" pageId="23" pageNumber="440" year="1995">
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<bibRefCitation id="EF9E54AFFF801E48FBDFFDD2FCF1FDB5" author="LE RENARD J. & PACAUD J. - M." pageId="23" pageNumber="440" pagination="65 - 132" refId="ref25125" refString="LE RENARD J. & PACAUD J. - M. 1995. - Revision des Mollusques paleogenes du Bassin de Paris. 2: Liste des references primaires des especes. Cossmanniana 3 (3): 65 - 132." type="journal article" year="1995">Le Renard & Pacaud 1995: 112</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AAE0F4FFF801E48FC94FDB4FB26FDB5" author="LE RENARD J. & PACAUD J. - M." box="[882,1177,585,609]" page="162" pageId="23" pageNumber="440" year="1995">
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<bibRefCitation id="EF9E54AFFF801E48FC94FDB4FB26FDB5" author="LE RENARD J. & PACAUD J. - M." box="[882,1177,585,609]" pageId="23" pageNumber="440" pagination="65 - 132" refId="ref25125" refString="LE RENARD J. & PACAUD J. - M. 1995. - Revision des Mollusques paleogenes du Bassin de Paris. 2: Liste des references primaires des especes. Cossmanniana 3 (3): 65 - 132." type="journal article" year="1995">Pacaud & Le Renard 1995: 162</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AAE0F4FFF801E48FD45FD9EFCC1FDAF" author="MERLE D. & PACAUD J. - M. & KRILOFF A. & LOUBRY P." box="[675,894,611,635]" page="198" pageId="23" pageNumber="440" year="2008">
|
||
<bibRefCitation id="EF9E54AFFF801E48FD45FD9EFCC1FDAF" author="MERLE D. & PACAUD J. - M. & KRILOFF A. & LOUBRY P." box="[675,894,611,635]" pageId="23" pageNumber="440" pagination="18 - 35" refId="ref25376" refString="MERLE D., PACAUD J. - M., KRILOFF A. & LOUBRY P. 2008. - Les motifs colores residuels des coquilles lutetiennes du Bassin de Paris, in MERLE D. (ed.), Stratotype lutetien. Collection Patrimoine geologique 1. Museum national d'Histoire naturelle, Paris; Biotope, Meze; BRGM, Orleans: 182 - 227, pls 18 - 35." type="book chapter" year="2008">
|
||
Merle
|
||
<emphasis id="B97BF54CFF801E48FD04FD99FCAEFDAF" box="[738,785,611,635]" italics="true" pageId="23" pageNumber="440">et al.</emphasis>
|
||
2008: 198
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 25, figs 1a, b, 2.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3157AD5FF801E48FD43FD6EFB3CFB56" pageId="23" pageNumber="440" type="materials_examined">
|
||
<paragraph id="8BB0295EFF801E48FD43FD6EFBFCFD12" blockId="23.[677,1209,659,710]" pageId="23" pageNumber="440">
|
||
<collectionCode id="ED1EB19BFF801E48FD43FD6EFC82FD7E" box="[677,829,659,682]" pageId="23" pageNumber="440">
|
||
<typeStatus id="54B497FCFF801E48FD43FD6EFD68FD7D" box="[677,727,659,682]" pageId="23" pageNumber="440">TYPE</typeStatus>
|
||
LOCALITY
|
||
</collectionCode>
|
||
. — Fontenay-en-Vexin, Bois du But (Eure,
|
||
<collectingCountry id="F31869CEFF801E48FD0FFD53FC91FD12" box="[745,814,686,710]" name="France" pageId="23" pageNumber="440">France</collectingCountry>
|
||
), Lutetian (Middle Eocene).
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF801E48FD43FD23FB1AFB99" blockId="23.[674,1212,733,1101]" pageId="23" pageNumber="440">
|
||
<collectionCode id="ED1EB19BFF801E48FD43FD23FC8DFD20" box="[677,818,734,757]" pageId="23" pageNumber="440">TYPE MATERIAL</collectionCode>
|
||
. — Th e illustrated
|
||
<typeStatus id="54B497FCFF801E48FC03FD20FB92FD21" box="[997,1069,733,757]" pageId="23" pageNumber="440" type="syntype">syntype</typeStatus>
|
||
(Bourdot coll., sample Bernay) of
|
||
<bibRefCitation id="EF9E54AFFF801E48FCAAFD05FC4BFCC4" author="COSSMANN M." box="[844,1012,760,784]" pageId="23" pageNumber="440" pagination="3 - 385" refId="ref23409" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris (4 e fascicule). Annales de la Societe royale malacologique de Belgique 24: 3 - 385." type="journal article" year="1889">Cossmann (1889)</bibRefCitation>
|
||
has been selected unambiguously by
|
||
<bibRefCitation id="EF9E54AFFF801E48FCD1FCEFFC79FCFE" author="JUNG P." box="[823,966,786,810]" pageId="23" pageNumber="440" pagination="5 - 52" refId="ref24521" refString="JUNG P. 1974. - A revision of the family Seraphsidae (Gastropoda: Strombacea). Paleontographica Americana 8 (47): 5 - 52." type="journal article" year="1974">Jung (1974:22)</bibRefCitation>
|
||
as
|
||
<typeStatus id="54B497FCFF801E48FC39FCEFFB8BFCFE" box="[991,1076,786,810]" pageId="23" pageNumber="440" type="lectotype">lectotype</typeStatus>
|
||
, using wrongly the term
|
||
<typeStatus id="54B497FCFF801E48FD1EFCD0FCF1FC91" box="[760,846,813,837]" pageId="23" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
: “Cossmann’s original figure […]. This specimen is the
|
||
<typeStatus id="54B497FCFF801E48FCD0FCBAFC33FC8B" box="[822,908,839,863]" pageId="23" pageNumber="440" type="holotype">holotype</typeStatus>
|
||
”. In accordance with article 74.5 of the
|
||
<collectionCode id="ED1EB19BFF801E48FD03FC9CFC9CFCAD" box="[741,803,865,889]" pageId="23" pageNumber="440">ICZN</collectionCode>
|
||
(1999) this mention, made before 2000, constitutes a valid
|
||
<typeStatus id="54B497FCFF801E48FCBFFC81FC10FC40" box="[857,943,892,916]" pageId="23" pageNumber="440" type="lectotype">lectotype</typeStatus>
|
||
designation. Unfortunately, the type material of this species has not been found in the Bourdot collection, now deposited at
|
||
<collectionCode id="ED1EB19BFF801E48FBEDFC4CFBF0FC1D" box="[1035,1103,945,969]" country="France" httpUri="http://grbio.org/cool/2dzr-gk4t" name="Centre de Paleontologie Stratigraphique et Paleoecologie" pageId="23" pageNumber="440">UCBL</collectionCode>
|
||
. A
|
||
<typeStatus id="54B497FCFF801E48FB89FC4CFB05FC1D" box="[1135,1210,945,969]" pageId="23" pageNumber="440" type="neotype">neotype</typeStatus>
|
||
designation is needed to clarify the taxinomic status of this species. A
|
||
<typeStatus id="54B497FCFF801E48FCE5FC1BFCF1FC2A" box="[771,846,998,1022]" pageId="23" pageNumber="440" type="neotype">neotype</typeStatus>
|
||
from the locality of Fontenay-en-Vexin (
|
||
<collectionCode id="ED1EB19BFF801E48FD4CFBFDFD43FBCC" box="[682,764,1024,1048]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="440">MNHN</collectionCode>
|
||
<accessionNumber id="945CB4BDFF801E48FCE6FBFDFCF1FBCC" box="[768,846,1024,1048]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A25009" pageId="23" pageNumber="440" type="EnaNcbi">A25009</accessionNumber>
|
||
,
|
||
<figureCitation id="133435DBFF801E48FCB3FBFDFC2CFBCC" box="[853,915,1024,1048]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="23" pageNumber="440">Fig.16</figureCitation>
|
||
C-E) is accordingly chosen here. This initiative conforms to the rules specified by the
|
||
<collectionCode id="ED1EB19BFF801E48FB9BFBE6FB05FBE7" box="[1149,1210,1051,1075]" pageId="23" pageNumber="440">ICZN</collectionCode>
|
||
(1999: art. 75), as regards the designation of a
|
||
<typeStatus id="54B497FCFF801E48FBB5FBC8FB1EFB99" box="[1107,1185,1077,1101]" pageId="23" pageNumber="440" type="neotype">neotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF801E48FD43FB96FB3CFB56" blockId="23.[677,1155,1130,1155]" box="[677,1155,1130,1155]" pageId="23" pageNumber="440">
|
||
<collectionCode id="ED1EB19BFF801E48FD43FB96FC06FB54" box="[677,953,1131,1155]" pageId="23" pageNumber="440">OTHER MATERIAL EXAMINED</collectionCode>
|
||
. — See Appendix 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3157AD5FF801E46FD43FB53FBB0F970" lastPageId="25" lastPageNumber="442" pageId="23" pageNumber="440" type="description">
|
||
<paragraph id="8BB0295EFF801E48FD43FB53FC8AFB12" blockId="23.[674,1212,1198,1700]" box="[677,821,1198,1224]" pageId="23" pageNumber="440">DESCRIPTION</paragraph>
|
||
<paragraph id="8BB0295EFF801E48FD43FB30FD68FB33" blockId="23.[674,1212,1198,1700]" box="[677,727,1229,1255]" pageId="23" pageNumber="440">
|
||
<emphasis id="B97BF54CFF801E48FD43FB30FD68FB33" box="[677,727,1229,1255]" italics="true" pageId="23" pageNumber="440">Shell</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF801E48FD43FB10FBE4F970" blockId="23.[674,1212,1198,1700]" pageId="23" pageNumber="440">
|
||
Shell up to
|
||
<quantity id="4CF784BBFF801E48FCFFFB10FCD9FADC" box="[793,870,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="440" unit="mm" value="28.0">28 mm</quantity>
|
||
in height and
|
||
<quantity id="4CF784BBFF801E48FC1FFB10FB87FADC" box="[1017,1080,1261,1288]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="440" unit="mm" value="8.0">8 mm</quantity>
|
||
in diameter, moderately slender with acute apical part. Aperture long and narrow. Callus of the inner lip well developed and well delineated, continuing adapically on the spire and forming a narrow band parallel to the outer lip (
|
||
<figureCitation id="133435DBFF801E48FCE0FA71FCDBFA73" box="[774,868,1420,1447]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="23" pageNumber="440">Fig. 15C</figureCitation>
|
||
). Columella bent backwards near the base. Outer lip, slightly thickened and sinuous in labral view, near the adapical end of the aperture. Outer lip extending to the apex and continuing on the opposite side of the apex for a short distance (
|
||
<figureCitation id="133435DBFF801E48FD48F9D7FC9EF991" box="[686,801,1578,1605]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="23" pageNumber="440">Fig. 15G, I</figureCitation>
|
||
). Jung called this apical part of the outer lip, the apical canal. Siphonal notch moderately deep on the dorsal side. Surface sculpture of narrow oblique grooves near the base (
|
||
<figureCitation id="133435DBFF801E48FC0EF974FBF1F970" box="[1000,1102,1673,1700]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="23" pageNumber="440">Fig. 15H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7079D6FF8F1E47FF97F9F5FC13F975" ID-DOI="http://doi.org/10.5281/zenodo.5381182" ID-Zenodo-Dep="5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="24" pageNumber="441" startId="24.[113,124,1544,1561]" targetBox="[126,1202,227,1464]" targetPageId="24">
|
||
<paragraph id="8BB0295EFF8F1E47FF97F9F5FC13F975" blockId="24.[113,1213,1544,1697]" pageId="24" pageNumber="441">
|
||
FIG. 15. —
|
||
<taxonomicName id="4C0F52DDFF8F1E47FF29F9F5FE28F9CD" authority="Montfort, 1810" authorityName="Montfort" authorityYear="1810" box="[207,407,1544,1561]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="441" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BF54CFF8F1E47FF29F9F5FEACF9CD" box="[207,275,1544,1561]" italics="true" pageId="24" pageNumber="441">Seraphs</emphasis>
|
||
<bibRefCitation id="EF9E54AFFF8F1E47FEFCF9F5FE28F9CD" author="MONTFORT D. DE" box="[282,407,1544,1561]" pageId="24" pageNumber="441" refId="ref25450" refString="MONTFORT D. DE 1810. - Conchyliologie systematique, et classification methodique des coquilles offrant leurs figures, leur arrangement generique, leurs descriptions caracteristiques, leurs noms, ainsi que leur synonymie en plusieurs langues. Tome second: coquilles univalves, non cloisonnees. Schoell, Paris, 676 p." type="book" year="1810">Montfort, 1810</bibRefCitation>
|
||
</taxonomicName>
|
||
shells in normal light;
|
||
<emphasis id="B97BF54CFF8F1E47FDB3F9F5FDD0F9CD" bold="true" box="[597,623,1544,1561]" pageId="24" pageNumber="441">A -I</emphasis>
|
||
,
|
||
<taxonomicName id="4C0F52DDFF8F1E47FD9CF9F5FC2BF9CD" authority="(Cossmann, 1889)" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[634,916,1544,1561]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="441" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8F1E47FD9CF9F5FD4CF9CD" box="[634,755,1544,1561]" italics="true" pageId="24" pageNumber="441">S. chilophorus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9E54AFFF8F1E47FD18F9F5FC30F9CD" author="COSSMANN M." box="[766,911,1544,1561]" pageId="24" pageNumber="441" pagination="3 - 385" refId="ref23409" refString="COSSMANN M. 1889. - Catalogue illustre des coquilles fossiles de l'Eocene des environs de Paris (4 e fascicule). Annales de la Societe royale malacologique de Belgique 24: 3 - 385." type="journal article" year="1889">Cossmann, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B97BF54CFF8F1E47FC7BF9F5FC02F9CD" bold="true" box="[925,957,1544,1561]" pageId="24" pageNumber="441">A -E</emphasis>
|
||
, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian;
|
||
<emphasis id="B97BF54CFF8F1E47FDDBF9E2FDF5F9E4" bold="true" box="[573,586,1567,1584]" pageId="24" pageNumber="441">A</emphasis>
|
||
, ventral view;
|
||
<emphasis id="B97BF54CFF8F1E47FD23F9E2FD6DF9E4" bold="true" box="[709,722,1567,1584]" pageId="24" pageNumber="441">B</emphasis>
|
||
, labral view;
|
||
<emphasis id="B97BF54CFF8F1E47FCA5F9E2FCEEF9E4" bold="true" box="[835,849,1567,1584]" pageId="24" pageNumber="441">C</emphasis>
|
||
, detailed view of the apical part;
|
||
<emphasis id="B97BF54CFF8F1E47FB8BF9E2FBC4F9E4" bold="true" box="[1133,1147,1567,1584]" pageId="24" pageNumber="441">D</emphasis>
|
||
, dorsal view;
|
||
<emphasis id="B97BF54CFF8F1E47FF46F9C8FF13F992" bold="true" box="[160,172,1589,1606]" pageId="24" pageNumber="441">E</emphasis>
|
||
, detailed view of the apical part;
|
||
<emphasis id="B97BF54CFF8F1E47FE5FF9C8FE66F992" bold="true" box="[441,473,1589,1606]" pageId="24" pageNumber="441">F -H</emphasis>
|
||
, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;
|
||
<emphasis id="B97BF54CFF8F1E47FC09F9C8FC45F992" bold="true" box="[1007,1018,1589,1606]" pageId="24" pageNumber="441">F</emphasis>
|
||
, ventral view;
|
||
<emphasis id="B97BF54CFF8F1E47FB88F9C8FBC3F992" bold="true" box="[1134,1148,1589,1606]" pageId="24" pageNumber="441">G</emphasis>
|
||
, dorsal view;
|
||
<emphasis id="B97BF54CFF8F1E47FF47F9B1FF10F989" bold="true" box="[161,175,1612,1629]" pageId="24" pageNumber="441">H</emphasis>
|
||
, detailed view of the apical part;
|
||
<emphasis id="B97BF54CFF8F1E47FE22F9B1FE75F989" bold="true" box="[452,458,1612,1629]" pageId="24" pageNumber="441">I</emphasis>
|
||
, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian;
|
||
<emphasis id="B97BF54CFF8F1E47FE95F99EFE2DF9A0" bold="true" box="[371,402,1635,1652]" pageId="24" pageNumber="441">J -N</emphasis>
|
||
,
|
||
<taxonomicName id="4C0F52DDFF8F1E47FE78F99EFDB6F9A0" authority="Caze & Merle & Pacaud & Saint Martin, 2010" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" box="[414,521,1635,1652]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="441" phylum="Mollusca" rank="species" species="peterjungi" status="sp. nov.">
|
||
<emphasis id="B97BF54CFF8F1E47FE78F99EFE15F9A0" box="[414,426,1635,1652]" italics="true" pageId="24" pageNumber="441">S</emphasis>
|
||
.
|
||
<emphasis id="B97BF54CFF8F1E47FE50F99EFDB6F9A0" box="[438,521,1635,1652]" italics="true" pageId="24" pageNumber="441">peterjungi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2484837FF8F1E47FDE9F99EFDFCF9A0" box="[527,579,1635,1652]" pageId="24" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype MNHN
|
||
<accessionNumber id="945CB4BDFF8F1E47FD3FF99EFCA6F9A0" box="[729,793,1635,1652]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A28578" pageId="24" pageNumber="441" type="EnaNcbi">A28578</accessionNumber>
|
||
(Pacaud leg.), Chaussy (Les Garennes), Lutetian;
|
||
<emphasis id="B97BF54CFF8F1E47FF95F984FFC2F95E" bold="true" box="[115,125,1657,1674]" pageId="24" pageNumber="441">J</emphasis>
|
||
, ventral view;
|
||
<emphasis id="B97BF54CFF8F1E47FF11F984FEBBF95E" bold="true" box="[247,260,1657,1674]" pageId="24" pageNumber="441">K</emphasis>
|
||
, labral view;
|
||
<emphasis id="B97BF54CFF8F1E47FE92F984FEC0F95E" bold="true" box="[372,383,1657,1674]" pageId="24" pageNumber="441">L</emphasis>
|
||
, detailed view of the apical part;
|
||
<emphasis id="B97BF54CFF8F1E47FD7FF984FD15F95E" bold="true" box="[665,682,1657,1674]" pageId="24" pageNumber="441">M</emphasis>
|
||
, detailed view the oblique grooves;
|
||
<emphasis id="B97BF54CFF8F1E47FC3CF984FC57F95E" bold="true" box="[986,1000,1657,1674]" pageId="24" pageNumber="441">N</emphasis>
|
||
, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FF25FEB5FF26" blockId="25.[111,648,216,876]" box="[113,266,216,242]" pageId="25" pageNumber="442">
|
||
<emphasis id="B97BF54CFF8E1E46FF97FF25FEB5FF26" box="[113,266,216,242]" italics="true" pageId="25" pageNumber="442">Colour pattern</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FF05FD8FFEA6" blockId="25.[111,648,216,876]" pageId="25" pageNumber="442">The colour pattern is composed of two different dark elements on a lighter background: thick rows of patches nearly perpendicular to the line of growth and tiny dots located between these rows.</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF6EFE8BFF4EFCB8" blockId="25.[111,648,216,876]" pageId="25" pageNumber="442">
|
||
The rows are composed of dark triangular patches (7-8). Th e apex of each of is orientated towards the outer lip. Th e patches are usually disconnected and separated by a small white fluorescent patch. This morphology is called a fishbone pattern (
|
||
<figureCitation id="133435DBFF8E1E46FF9FFDE8FF54FDE4" box="[121,235,533,560]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="25" pageNumber="442">Fig. 16I, J</figureCitation>
|
||
) by
|
||
<bibRefCitation id="EF9E54AFFF8E1E46FEFDFDE8FE60FDE4" author="MEINHARDT H." box="[283,479,533,560]" pageId="25" pageNumber="442" refId="ref25280" refString="MEINHARDT H. 1998. - The Algorithmic beauty of Sea shells. The Virtual Laboratory. Springer-Verlag, Berlin; Heidelberg, New-York, 236 p." type="book" year="1998">Meinhardt (1998</bibRefCitation>
|
||
: fig. 4.10). The distance between the rows and their width are variable. Sometimes, several rows are adjacent to one another (
|
||
<figureCitation id="133435DBFF8E1E46FED7FD89FE26FD5B" box="[305,409,628,655]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="25" pageNumber="442">Fig. 16F</figureCitation>
|
||
). Th e dots are small and more or less triangular (
|
||
<figureCitation id="133435DBFF8E1E46FE59FD6EFD96FD7A" box="[447,553,659,686]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="25" pageNumber="442">Fig. 16E</figureCitation>
|
||
). As for the patches, the apex is always directed towards the outer lip. Th eir size, distribution and shape are heterogenous and irregular. Above the most adapical row, the density and the size of dots are often greater as is the degree of coalescence (
|
||
<figureCitation id="133435DBFF8E1E46FF9FFCACFF5BFCB8" box="[121,228,849,876]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="25" pageNumber="442">Fig. 16H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FC6FFDFCFC7D" blockId="25.[112,647,914,1383]" box="[113,579,914,940]" pageId="25" pageNumber="442">COMPARISONS OF THE SHELL MORPHOLOGY</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FC4DFEF4FBBE" blockId="25.[112,647,914,1383]" pageId="25" pageNumber="442">
|
||
The morphologies of
|
||
<taxonomicName id="4C0F52DDFF8E1E46FE86FC4CFE4CFC1E" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[352,499,944,971]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FE86FC4CFE4CFC1E" box="[352,499,944,971]" italics="true" pageId="25" pageNumber="442">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F52DDFF8E1E46FDC8FC4CFF18FC3E" authorityName="Caze & Merle & Pacaud & Saint Martin" authorityYear="2010" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="peterjungi">
|
||
<emphasis id="B97BF54CFF8E1E46FDC8FC4CFF18FC3E" italics="true" pageId="25" pageNumber="442">S. peterjungi</emphasis>
|
||
</taxonomicName>
|
||
n. sp (
|
||
<figureCitation id="133435DBFF8E1E46FF09FC2DFE85FC3F" box="[239,314,976,1003]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="25" pageNumber="442">Fig. 15</figureCitation>
|
||
J-N) are very similar with their thickened callus of the inner lip and their narrow band on the adapical part of the shell. Moreover, they share an apical canal (
|
||
<figureCitation id="133435DBFF8E1E46FE68FBD2FD99FB9E" box="[398,550,1071,1098]" captionStart="FIG" captionStartId="24.[113,124,1544,1561]" captionTargetBox="[126,1202,227,1464]" captionTargetId="figure-192@24.[113,1210,214,1500]" captionTargetPageId="24" captionText="FIG. 15. — Seraphs Montfort, 1810 shells in normal light; A-I, S. chilophorus (Cossmann, 1889); A-E, MNHN A28779 (Ledon coll.), Chaussy (Les Garennes), Val d’Oise, France, Lutetian; A, ventral view; B, labral view; C, detailed view of the apical part; D, dorsal view;E, detailed view of the apical part; F-H, MNHN A28780 (Ledon coll.), Chaussy (Les Garennes), Lutetian;F, ventral view;G, dorsal view; H, detailed view of the apical part; I, MNHN A27783, detailed view the oblique grooves (leg. Pacaud), Saint-Lubin-de-la-Haye, Eure-et-Loir, France, Lutetian; J-N, S. peterjungi n. sp., holotype MNHN A28578 (Pacaud leg.), Chaussy (Les Garennes), Lutetian; J, ventral view; K, labral view; L, detailed view of the apical part; M, detailed view the oblique grooves; N, dorsal view. Scale bars: A-B, D, F-G, J-K, N, 10 mm; C, I, 5 mm; E, H, L, M, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381182" httpUri="https://zenodo.org/record/5381182/files/figure.png" pageId="25" pageNumber="442">Fig. 15G, I, L</figureCitation>
|
||
), lacking in the other
|
||
<taxonomicName id="4C0F52DDFF8E1E46FF11FBB2FEFBFBBD" authorityName="Montfort" authorityYear="1810" box="[247,324,1103,1129]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97BF54CFF8E1E46FF11FBB2FEFBFBBD" box="[247,324,1103,1129]" italics="true" pageId="25" pageNumber="442">Seraphs</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF6EFB93FF67FAB3" blockId="25.[112,647,914,1383]" pageId="25" pageNumber="442">
|
||
The shell of
|
||
<taxonomicName id="4C0F52DDFF8E1E46FEEBFB92FE1EFB5C" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[269,417,1134,1161]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FEEBFB92FE1EFB5C" box="[269,417,1134,1161]" italics="true" pageId="25" pageNumber="442">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
is more slender than those of
|
||
<taxonomicName id="4C0F52DDFF8E1E46FF30FB72FEE8FB7C" baseAuthorityName="Solander & Brander" baseAuthorityYear="1766" box="[214,343,1166,1193]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="volutatus">
|
||
<emphasis id="B97BF54CFF8E1E46FF30FB72FF5CFB7D" box="[214,227,1167,1193]" italics="true" pageId="25" pageNumber="442">S</emphasis>
|
||
.
|
||
<emphasis id="B97BF54CFF8E1E46FF13FB73FEE8FB7C" box="[245,343,1166,1192]" italics="true" pageId="25" pageNumber="442">volutatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133435DBFF8E1E46FE8FFB73FE13FB7D" box="[361,428,1166,1193]" captionStart="FIG" captionStartId="10.[113,124,1544,1561]" captionTargetBox="[125,1197,227,1487]" captionTargetId="figure-170@10.[113,1210,214,1505]" captionTargetPageId="10" captionText="FIG. 6. — Seraphs Montfort, 1810 shells in normal light: A-E, S. volutatus (Solander in Brander, 1766); A-C, MNHN A28776 (ex Galerie de Zoologie coll.), Grignon, Yvelines, France, Lutetian; A, ventral view; B, labral view; C, dorsal view; D, E, lectotype of Terebellum convolutum Lamarck, 1802,MNHN B70325 (Lamarck coll.), Grignon,Lutetian; D, ventral view; E, dorsal view; F, G, S. sopitus (Solan- der in Brander,1766),MNHN A28778 (Faullummel coll.), Le Limon, Seine-et-Marne, France, Bartonian;F, ventral view; G, labral view; H-J, S. olivaceus (Cossmann,1889);H, I, MNHN A28577 (Ledon coll.),Chaussy (Les Garennes),Val d’Oise,France,Lutetian;H, ventral view; I, dorsal view; J, MNHN A29225, labral view (Ledon coll.), Chaussy (Les Garennes), Lutetian. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381164" httpUri="https://zenodo.org/record/5381164/files/figure.png" pageId="25" pageNumber="442">Fig. 6</figureCitation>
|
||
A-E) and
|
||
<taxonomicName id="4C0F52DDFF8E1E46FDC6FB72FD38FB7C" authorityName="sensu Jung" authorityYear="1974" box="[544,647,1167,1193]" class="Gastropoda" family="Bullidae" genus="Bulla" kingdom="Animalia" order="Cephalaspidea" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="sopitus">
|
||
<emphasis id="B97BF54CFF8E1E46FDC6FB72FD38FB7C" box="[544,647,1167,1193]" italics="true" pageId="25" pageNumber="442">S. sopitus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133435DBFF8E1E46FF9FFB53FF51FB1D" box="[121,238,1198,1225]" captionStart="FIG" captionStartId="10.[113,124,1544,1561]" captionTargetBox="[125,1197,227,1487]" captionTargetId="figure-170@10.[113,1210,214,1505]" captionTargetPageId="10" captionText="FIG. 6. — Seraphs Montfort, 1810 shells in normal light: A-E, S. volutatus (Solander in Brander, 1766); A-C, MNHN A28776 (ex Galerie de Zoologie coll.), Grignon, Yvelines, France, Lutetian; A, ventral view; B, labral view; C, dorsal view; D, E, lectotype of Terebellum convolutum Lamarck, 1802,MNHN B70325 (Lamarck coll.), Grignon,Lutetian; D, ventral view; E, dorsal view; F, G, S. sopitus (Solan- der in Brander,1766),MNHN A28778 (Faullummel coll.), Le Limon, Seine-et-Marne, France, Bartonian;F, ventral view; G, labral view; H-J, S. olivaceus (Cossmann,1889);H, I, MNHN A28577 (Ledon coll.),Chaussy (Les Garennes),Val d’Oise,France,Lutetian;H, ventral view; I, dorsal view; J, MNHN A29225, labral view (Ledon coll.), Chaussy (Les Garennes), Lutetian. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381164" httpUri="https://zenodo.org/record/5381164/files/figure.png" pageId="25" pageNumber="442">Fig. 6F, G</figureCitation>
|
||
). Th is species is generally more slen- der than
|
||
<taxonomicName id="4C0F52DDFF8E1E46FF30FB33FEEFFB33" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[214,336,1229,1256]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="olivaceus">
|
||
<emphasis id="B97BF54CFF8E1E46FF30FB33FEEFFB33" box="[214,336,1229,1256]" italics="true" pageId="25" pageNumber="442">S. olivaceus</emphasis>
|
||
</taxonomicName>
|
||
, although both can have an acute apex. Also,
|
||
<taxonomicName id="4C0F52DDFF8E1E46FEA4FB13FE5FFAD3" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[322,480,1261,1288]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FEA4FB13FE5FFAD3" box="[322,480,1261,1288]" italics="true" pageId="25" pageNumber="442">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from
|
||
<taxonomicName id="4C0F52DDFF8E1E46FF49FAF0FE97FAF3" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[175,296,1293,1319]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="olivaceus">
|
||
<emphasis id="B97BF54CFF8E1E46FF49FAF0FE97FAF3" box="[175,296,1293,1319]" italics="true" pageId="25" pageNumber="442">S. olivaceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F52DDFF8E1E46FED1FAF0FE0CFAF3" baseAuthorityName="Solander & Brander" baseAuthorityYear="1766" box="[311,435,1293,1319]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="volutatus">
|
||
<emphasis id="B97BF54CFF8E1E46FED1FAF0FE0CFAF3" box="[311,435,1293,1319]" italics="true" pageId="25" pageNumber="442">S. volutatus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C0F52DDFF8E1E46FE3EFAF0FD84FAF3" authorityName="sensu Jung" authorityYear="1974" box="[472,571,1293,1319]" class="Gastropoda" family="Bullidae" genus="Bulla" kingdom="Animalia" order="Cephalaspidea" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="sopitus">
|
||
<emphasis id="B97BF54CFF8E1E46FE3EFAF0FD84FAF3" box="[472,571,1293,1319]" italics="true" pageId="25" pageNumber="442">S. sopitus</emphasis>
|
||
</taxonomicName>
|
||
by the morphology of its outer lip and its callus of the inner lip.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FA71FDB3FA70" blockId="25.[112,648,1420,1700]" box="[113,524,1420,1446]" pageId="25" pageNumber="442">VARIABILITY OF THE COLOUR PATTERN</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF97FA56FF5DF9B1" blockId="25.[112,648,1420,1700]" pageId="25" pageNumber="442">
|
||
Among
|
||
<specimenCount id="9D09E2D7FF8E1E46FF2FFA56FED8FA12" box="[201,359,1451,1478]" pageId="25" pageNumber="442" type="generic">163 specimens</specimenCount>
|
||
from various localities (11 from the Lutetian and 1 from the Bartonian), 160 (98.16%) show a residual colour pattern under UV light (
|
||
<tableCitation id="C68D1CE5FF8E1E46FF3BF9F7FE94F9F1" box="[221,299,1546,1573]" captionStart="TABLE" captionStartId="9.[285,294,219,236]" captionText="TABLE 1. — Comparison between positive and" pageId="25" pageNumber="442">Table 1</tableCitation>
|
||
). Accordingly, these observations provide a good general survey of the intraspecific variability.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FF6EF994FB81FEC7" blockId="25.[112,648,1420,1700]" lastBlockId="25.[674,1211,216,1447]" pageId="25" pageNumber="442">
|
||
The residual pattern of
|
||
<taxonomicName id="4C0F52DDFF8E1E46FE75F997FD95F957" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[403,554,1641,1668]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FE75F997FD95F957" box="[403,554,1641,1668]" italics="true" pageId="25" pageNumber="442">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
is quite variable (
|
||
<figureCitation id="133435DBFF8E1E46FF35F974FEA0F970" box="[211,287,1673,1700]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17</figureCitation>
|
||
) and is often subject to significant distortion of the elements on the shell extremities, especially the apical one (
|
||
<figureCitation id="133435DBFF8E1E46FC53FF05FBBEFEC7" box="[949,1025,248,275]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17</figureCitation>
|
||
C-F).
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FD5DFEEAFCF1FCB8" blockId="25.[674,1211,216,1447]" pageId="25" pageNumber="442">
|
||
The huge variability is visible on every element of the residual pattern. Th e number of spiral rows varies from 0 to 6. Several shells show partially or totally connected rows (
|
||
<figureCitation id="133435DBFF8E1E46FC47FE8BFB9EFE45" box="[929,1057,374,401]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17B, D</figureCitation>
|
||
). Th e shape of the dark patches can show important variations from triangular (
|
||
<figureCitation id="133435DBFF8E1E46FCFFFE4BFC29FE05" box="[793,918,438,465]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17A, C</figureCitation>
|
||
) to completely distorted or shapeless (
|
||
<figureCitation id="133435DBFF8E1E46FCF3FE28FC34FE24" box="[789,907,469,496]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17F, J</figureCitation>
|
||
). Although they are usually separated by a small fluorescent patch, they may be connected with each other (
|
||
<figureCitation id="133435DBFF8E1E46FC2CFDE8FB95FDE4" box="[970,1066,533,560]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17E</figureCitation>
|
||
). The density of dots fluctuates from low values (
|
||
<figureCitation id="133435DBFF8E1E46FBFEFDC9FBC7FD9B" box="[1048,1144,564,591]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17A</figureCitation>
|
||
, D-F) to very high ones (
|
||
<figureCitation id="133435DBFF8E1E46FC8CFDA9FC72FDBB" box="[874,973,596,623]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17K</figureCitation>
|
||
). In this case, the dots form axial rows more or less coalescent (
|
||
<figureCitation id="133435DBFF8E1E46FBB7FD89FB23FD5B" box="[1105,1180,628,655]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17</figureCitation>
|
||
K- N). Th is coalescence sometimes increases until the dots disappear and only axial segments are observable (
|
||
<figureCitation id="133435DBFF8E1E46FD38FD2EFC31FD3A" box="[734,910,723,750]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17H, M, N</figureCitation>
|
||
). Th ere are often some large triangular and coalescent patches near the edge of the outer lip. Finally, there are sometimes small axial segments between the dark patches of the rows (
|
||
<figureCitation id="133435DBFF8E1E46FD01FCACFCFEFCB8" box="[743,833,849,876]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FD5DFC8CFBB1FA73" blockId="25.[674,1211,216,1447]" pageId="25" pageNumber="442">
|
||
In spite of this variability, we can distinguish three morphological
|
||
<typeStatus id="54B497FCFF8E1E46FC60FC6CFC7EFC78" box="[902,961,913,940]" pageId="25" pageNumber="442">types</typeStatus>
|
||
. Th e most abundant an representative
|
||
<typeStatus id="54B497FCFF8E1E46FCD9FC4DFCD2FC1F" box="[831,877,944,971]" pageId="25" pageNumber="442">type</typeStatus>
|
||
1 is composed of spiral rows of dark triangles directed towards the outer lip and small dots between them (
|
||
<figureCitation id="133435DBFF8E1E46FC57FC0DFC48FBDF" box="[945,1015,1008,1035]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17</figureCitation>
|
||
A-C, I). Th e
|
||
<typeStatus id="54B497FCFF8E1E46FB9AFC0DFB16FBDF" box="[1148,1193,1008,1035]" pageId="25" pageNumber="442">type</typeStatus>
|
||
2 shows some thin and tightened axial segments with, often, remnants of spiral rows of triangular patches (
|
||
<figureCitation id="133435DBFF8E1E46FD4CFBB2FCECFBBE" box="[682,851,1103,1130]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17H, N, O</figureCitation>
|
||
). Th e
|
||
<typeStatus id="54B497FCFF8E1E46FC73FBB2FC7CFBBE" box="[917,963,1103,1130]" pageId="25" pageNumber="442">type</typeStatus>
|
||
3 shows only large and more or less coalescent dark triangles (
|
||
<figureCitation id="133435DBFF8E1E46FBAAFB92FD05FB7D" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17D, E</figureCitation>
|
||
). Several shells bear a morphology intermediate (
|
||
<figureCitation id="133435DBFF8E1E46FD34FB53FC9FFB1D" box="[722,800,1198,1225]" captionStart="FIG" captionStartId="28.[677,688,833,850]" captionTargetBox="[682,1206,216,790]" captionTargetId="graphics-597@28.[740,1206,221,688]" captionTargetPageId="28" captionText="FIG. 18.– Distribution of the pattern types with their intermediaries (Inter.) in Seraphs chilophorus (Cossmann, 1889)." figureDoi="http://doi.org/10.5281/zenodo.5381188" httpUri="https://zenodo.org/record/5381188/files/figure.png" pageId="25" pageNumber="442">Fig. 18</figureCitation>
|
||
) between
|
||
<typeStatus id="54B497FCFF8E1E46FC76FB53FC78FB1D" box="[912,967,1198,1225]" pageId="25" pageNumber="442">types</typeStatus>
|
||
1 and 2 (
|
||
<figureCitation id="133435DBFF8E1E46FBD6FB53FBC4FB1D" box="[1072,1147,1198,1225]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17</figureCitation>
|
||
J-M), between
|
||
<typeStatus id="54B497FCFF8E1E46FCE5FB33FC85FB3D" box="[771,826,1230,1257]" pageId="25" pageNumber="442">types</typeStatus>
|
||
1 and 3 (
|
||
<figureCitation id="133435DBFF8E1E46FC78FB33FC43FB3D" box="[926,1020,1230,1257]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17F</figureCitation>
|
||
), between
|
||
<typeStatus id="54B497FCFF8E1E46FB88FB33FB1AFB3D" box="[1134,1189,1230,1257]" pageId="25" pageNumber="442">types</typeStatus>
|
||
2 and 3 (
|
||
<figureCitation id="133435DBFF8E1E46FD16FB10FCEBFADC" box="[752,852,1261,1288]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Fig. 17G</figureCitation>
|
||
). Th e relative abundance of these morphologies is given in
|
||
<figureCitation id="133435DBFF8E1E46FC5FFAF0FB9AFAFC" box="[953,1061,1293,1320]" captionStart="FIG" captionStartId="28.[677,688,833,850]" captionTargetBox="[682,1206,216,790]" captionTargetId="graphics-597@28.[740,1206,221,688]" captionTargetPageId="28" captionText="FIG. 18.– Distribution of the pattern types with their intermediaries (Inter.) in Seraphs chilophorus (Cossmann, 1889)." figureDoi="http://doi.org/10.5281/zenodo.5381188" httpUri="https://zenodo.org/record/5381188/files/figure.png" pageId="25" pageNumber="442">Figure 18</figureCitation>
|
||
.
|
||
<figureCitation id="133435DBFF8E1E46FBD7FAF0FB25FAFC" box="[1073,1178,1293,1320]" captionStart="FIG" captionStartId="27.[113,124,1521,1538]" captionTargetBox="[119,1195,220,1436]" captionTargetId="figure-199@27.[113,1210,214,1500]" captionTargetPageId="27" captionText="FIG. 17. — Variability of Seraphs chilophorus (Cossmann, 1889) in dorsal view and under UV light: A, I, MNHN A28926 (leg. Pacaud), Fercourt, Oise, France, Lutetian; B, MNHN A28924 (leg. Pacaud), Fercourt, Lutetian; C, MNHN A28928 (leg. Pacaud), Parnes, Oise, France,Lutetian;D, MNHN A28922 (leg.Pacaud),Fercourt,Lutetian;E, MNHN A28921 (1965-11 coll.),Saulxmarchais,Yvelines,France, Lutetian; F, MNHN A28923 (leg. Pacaud), Fercourt, Lutetian; G, MNHN A28919 (leg. Pacaud), Parnes, Lutetian; H, MNHN A28891 (leg. Pacaud), Parnes, Lutetian; J, MNHN A28925 (leg. Pacaud), Fercourt,Lutetian; K, MNHN A28888 (leg. Pacaud), Parnes,Lutetian; L, MNHN A28929 (leg. Pacaud), Parnes, Lutetian; M, MNHN A28889 (leg. Pacaud), Parnes, Lutetian; N, MNHN A28890 (leg. Pacaud), Parnes, Lutetian; O, MNHN A28920 (leg. Pacaud), Parnes, Lutetian; A-C, I, Type 1; H, N, O, Type 2; D, E, Type 3; J-M, intermediaries 1-2; F, intermediary 1-3; G, intermediary 2-3. Scale bars: 10 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381186" httpUri="https://zenodo.org/record/5381186/files/figure.png" pageId="25" pageNumber="442">Figure 17</figureCitation>
|
||
illustrates this variability with a certain disparity, but the observation of numerous intermediate specimens suggest that it is impossible to separate
|
||
<specimenCount id="9D09E2D7FF8E1E46FBBFFA91FB08FA53" box="[1113,1207,1388,1415]" pageId="25" pageNumber="442" type="generic" typeStatus="type">one type</specimenCount>
|
||
from another, as different species.
|
||
</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FD43FA31FBE8FA30" blockId="25.[674,1210,1484,1700]" box="[677,1111,1484,1510]" pageId="25" pageNumber="442">COMPARISONS OF THE COLOUR PATTERN</paragraph>
|
||
<paragraph id="8BB0295EFF8E1E46FD43FA16FBB0F970" blockId="25.[674,1210,1484,1700]" pageId="25" pageNumber="442">
|
||
<taxonomicName id="4C0F52DDFF8E1E46FD43FA16FCCEF9D1" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[677,881,1515,1541]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FD43FA16FCCEF9D1" box="[677,881,1515,1541]" italics="true" pageId="25" pageNumber="442">Seraphs chilophorus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133435DBFF8E1E46FC67FA16FC6EF9D2" box="[897,977,1515,1542]" captionStart="FIG" captionStartId="26.[113,124,1589,1606]" captionTargetBox="[114,1210,215,1525]" captionTargetId="figure-144@26.[113,1210,240,1526]" captionTargetPageId="26" captionText="FIG. 16. — Seraphs chilophorus (Cossmann, 1889) under UV light: A, B, MNHN A28927 (Roissy coll.), Parnes, Oise, France, Lutetian; A, ventral view; B, dorsal view; C-E, neotype MNHN A25009, Fontenay-en-Vexin, Eure, France, Lutetian; C, ventral view; D, dorsal view; E, detailed view of the dots; F-H, MNHN A28924 (leg. Pacaud), Fercourt, Oise, France, Lutetian; F, dorsal view; G, H, detailed views of the dots; I-K, MNHN A28928 (leg. Pacaud), Parnes, Oise, France, Lutetian; I, ventral view; J, K, dorsal views (K negative view). Scale bars: A-D, F, I-K, 10 mm; E, G, H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.5381184" httpUri="https://zenodo.org/record/5381184/files/figure.png" pageId="25" pageNumber="442">Fig. 16</figureCitation>
|
||
) bears dots, as
|
||
<taxonomicName id="4C0F52DDFF8E1E46FB9AFA16FD52F9F0" baseAuthorityName="Solander & Brander" baseAuthorityYear="1766" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="volutatus">
|
||
<emphasis id="B97BF54CFF8E1E46FB9AFA16FD52F9F0" italics="true" pageId="25" pageNumber="442">S. volutatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F52DDFF8E1E46FD1BF9F6FCDEF9F0" authorityName="sensu Jung" authorityYear="1974" box="[765,865,1547,1573]" class="Gastropoda" family="Bullidae" genus="Bulla" kingdom="Animalia" order="Cephalaspidea" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="sopitus">
|
||
<emphasis id="B97BF54CFF8E1E46FD1BF9F6FCDEF9F0" box="[765,865,1547,1573]" italics="true" pageId="25" pageNumber="442">S. sopitus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0F52DDFF8E1E46FC94F9F6FC53F9F0" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[882,1004,1546,1573]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="olivaceus">
|
||
<emphasis id="B97BF54CFF8E1E46FC94F9F6FC53F9F0" box="[882,1004,1546,1573]" italics="true" pageId="25" pageNumber="442">S. olivaceus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0F52DDFF8E1E46FBCEF9F6FB05F9F0" authorityName="Jung" authorityYear="1974" box="[1064,1210,1546,1573]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="leukoleptus">
|
||
<emphasis id="B97BF54CFF8E1E46FBCEF9F6FB05F9F0" box="[1064,1210,1546,1573]" italics="true" pageId="25" pageNumber="442">S. leukoleptus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133435DBFF8E1E46FD4DF9D7FCB0F991" box="[683,783,1578,1605]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="12.[113,124,1589,1606]" captionStartId-1="13.[112,123,979,996]" captionStartId-2="14.[113,124,1589,1606]" captionStartId-3="16.[113,124,1589,1606]" captionTargetBox-0="[114,1210,215,1525]" captionTargetBox-1="[114,1209,214,931]" captionTargetBox-2="[114,1210,215,1525]" captionTargetBox-3="[113,1210,215,1531]" captionTargetId-0="figure-126@12.[113,1210,240,1526]" captionTargetId-1="figure-372@13.[130,1219,214,970]" captionTargetId-2="figure-143@14.[113,1210,240,1526]" captionTargetId-3="figure-126@16.[113,1210,245,1531]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="16" captionText-0="FIG. 7. — Seraphs volutatus (Solander in Brander, 1766) under UV light: A, B, G, MNHN A28873, Grignon, Yvelines, France, Lutetian; A, ventral view; B, dorsal view; G, detailed view of the dots; C, D, MNHN A28876, Grignon, Lutetian; C, ventral view; D, dorsal view; E, F, MNHN A28875 (Valenciennes coll.), Fontenay-en-Vexin, Eure, France, Lutetian; E, ventral view; F, dorsal view; H, I, MNHN A28874, Grignon, Lutetian; H, ventral view; I, dorsal view. Scale bars: A-F, H, I, 10 mm; G, 5 mm. Photographs by C. Lemzaouda (MNHN)." captionText-1="FIG. 8. — Oldest known specimens of Seraphs volutatus (Solander in Brander, 1766) under UV light: A-D, MNHN A30419, Sapicourt, Marne, France, Ypresian (Cuisian); A, dorsal view; B, C, detailed view of the dots; D, labral view; E, F, MNHN A30420, Sapicourt, Ypresian (Cuisian); E, dorsal view; F, labral view. Scale bars: A, D, E, F, 5 mm; B, C, 2 mm. Photographs by C. Lemzaouda (MNHN)." captionText-2="FIG. 9. — Seraphs sopitus (Solander in Brander,1766) under UV light:A-D, MNHN A28884,Saint-Witz (Le Guépelle),Val d’Oise,France, Bartonian; A, dorsal view; B, labral view; C, D, detailed views of the dots; E-H, MNHN A27526, Saint-Witz (Le Guépelle), Bartonian; E, dorsal view; F, abapertural view; G, H, detailed views of the dots; I-K, MNHN A28778, Le Limon, Seine-et-Marne, France, Bartonian; I, dorsal view; J, detailed view of the dots; K, abapertural view. Scale bars: A, B, E, F, I, K, 10 mm; C, D, J, 2 mm; G, H, 5 mm. Photographs by C. Lemzaouda and P. Loubry (MNHN)." captionText-3="FIG. 10. — Seraphs olivaceus (Cossmann, 1889) under UV light: A-C, MNHN A28560 (leg. Pacaud), Chaussy (Les Garennes), Val d’Oise,France,Lutetian; A, dorsal view; B, abapertural view; C, detailed view of the dots;D, E, MNHN A29057 (Houdas coll.),Chaussy (Les Garennes), Lutetian; D, dorsal view; E, abapertural view; F-H, neotype, MNHN A28885 (leg. Pacaud), Chaussy (Les Garennes), Lutetian; F, dorsal view; G, labral view; H, detailed view of the dots. Scale bars: A, B, D-G, 10 mm; C, 5 mm; H, 2 mm. Photographs by C. Lemzaouda (MNHN)." figureDoi-0="http://doi.org/10.5281/zenodo.5381166" figureDoi-1="http://doi.org/10.5281/zenodo.5381168" figureDoi-2="http://doi.org/10.5281/zenodo.5381170" figureDoi-3="http://doi.org/10.5281/zenodo.5381172" httpUri-0="https://zenodo.org/record/5381166/files/figure.png" httpUri-1="https://zenodo.org/record/5381168/files/figure.png" httpUri-2="https://zenodo.org/record/5381170/files/figure.png" httpUri-3="https://zenodo.org/record/5381172/files/figure.png" pageId="25" pageNumber="442">Figs 7-10</figureCitation>
|
||
). Th e pattern of
|
||
<taxonomicName id="4C0F52DDFF8E1E46FC50F9D7FB81F990" baseAuthorityName="Cossmann" baseAuthorityYear="1889" box="[950,1086,1578,1604]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="442" phylum="Mollusca" rank="species" species="chilophorus">
|
||
<emphasis id="B97BF54CFF8E1E46FC50F9D7FB81F990" box="[950,1086,1578,1604]" italics="true" pageId="25" pageNumber="442">S. chilophorus</emphasis>
|
||
</taxonomicName>
|
||
is, however, easily distinguishable as the dots are composed of only one residual colour and by the occurrence of broad spiral rows of dark patches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |