493 lines
75 KiB
XML
493 lines
75 KiB
XML
<document id="3682730056D5C5D866ABBD04B21FB586" ID-CLB-Dataset="9554" ID-DOI="10.5281/zenodo.4272467" ID-GBIF-Dataset="59fc10fe-e8c3-4537-abb9-049fbda4c7ed" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272467" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298451113" checkinUser="felipe" docAuthor="Hoshina, Hideto" docDate="2012" docId="03E98224323C072976E041A8FE5EA0AF" docLanguage="en" docName="ActaEntMusNatPra.52.suppl.1-168.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl. 1) 52" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Leiodes toyoshimai Hoshina 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="61" masterDocId="FFD0FA5C3200076977334233FFA5A45C" masterDocTitle="Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)" masterLastPageNumber="168" masterPageNumber="1" pageNumber="57" updateTime="1698846259177" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="38DFE8B2F2E2C2F1734C3BAFF1E03BAB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="78B0B9F1601669F88F427F71BD049EFB">
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<mods:title id="22FA46C26555544F3FFDA5519D53F0B2">Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="18EA8BFCA8A87283F87052E46095729C" type="personal">
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<mods:role id="0C407FF38BA6E3084D1CB2AF93F52AFC">
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<mods:roleTerm id="9B4CB41E43F8DD586376C0A26B201422">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A4F74A75C47F530493EBDF80174CD056">Hoshina, Hideto</mods:namePart>
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<mods:typeOfResource id="D4AEA39744FDC89830DC2A631FAE6F0E">text</mods:typeOfResource>
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<mods:title id="A399C4524F8F7BABA6AB97DCBE45D555">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="0E79461B820212E6183D902597F0842F">
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<mods:date id="446C333610B26E7B8318DFDD7E7E64B3">2012</mods:date>
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<mods:detail id="A77F2457FACC175652183AE6E25D091A" type="series">
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<mods:title id="E6DC3DB3C272AED63BB75A4F6710A544">suppl. 1</mods:title>
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</mods:detail>
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<mods:detail id="3814955CC12EB5779DD1FD5F0209FE45" type="pubDate">
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<mods:number id="14A1B0B1EFC96CF2765758261476A701">2012-09-15</mods:number>
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</mods:detail>
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<mods:detail id="5BF1145803F1120E53EDE13BEE04F93F" type="volume">
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<mods:number id="B9701094F73822CA4C8342852BBAF6F6">52</mods:number>
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</mods:detail>
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<mods:extent id="D57AA9F42459A5A4E36E0B39D38043B6" unit="page">
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<mods:start id="271FBC8E28C96F0FCB9BA362E1FBB265">1</mods:start>
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<mods:end id="341C3E62A1FA32759DC4EBA43DBB77C5">168</mods:end>
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<mods:classification id="AB81D396AEEAC450EDE1E610A7FFEE70">journal article</mods:classification>
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<mods:identifier id="0EDDBD88D7D45323A630BF31507BFD3C" type="CLB-Dataset">9554</mods:identifier>
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<mods:identifier id="8CED8C0D9F1B8579317E90B126E09D45" type="DOI">10.5281/zenodo.4272467</mods:identifier>
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<mods:identifier id="4823DE4B87D15B2AC9A8413EE226B0F3" type="GBIF-Dataset">59fc10fe-e8c3-4537-abb9-049fbda4c7ed</mods:identifier>
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<mods:identifier id="8283EEFACF6C68833271F787FAC759D3" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="0FAF9E0A7188821D441DB07A826B3F18" type="Zenodo-Dep">4272467</mods:identifier>
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<treatment id="03E98224323C072976E041A8FE5EA0AF" ID-DOI="http://doi.org/10.5281/zenodo.4339396" ID-GBIF-Taxon="170447922" ID-Zenodo-Dep="4339396" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF" httpUri="http://treatment.plazi.org/id/03E98224323C072976E041A8FE5EA0AF" lastPageId="64" lastPageNumber="61" pageId="60" pageNumber="57">
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<subSubSection id="C35A60B9323C075576E041A8FC8EA7E8" box="[467,811,922,948]" pageId="60" pageNumber="57" type="nomenclature">
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<paragraph id="8BFF3332323C075576E041A8FC8EA7E8" blockId="60.[434,844,922,1000]" box="[467,811,922,948]" pageId="60" pageNumber="57">
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<heading id="D0B7845E323C075576E041A8FC8EA7E8" bold="true" box="[467,811,922,948]" centered="true" fontSize="10" level="2" pageId="60" pageNumber="57" reason="2">
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<emphasis id="B934EF20323C075576E041A8FC8EA7E8" bold="true" box="[467,811,922,948]" pageId="60" pageNumber="57">
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||
10.
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<taxonomicName id="4C4048B1323C075576C841A9FD6BA7EF" ID-CoL="3SYK6" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[507,718,922,947]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="57" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
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<emphasis id="B934EF20323C075576C841A9FD6BA7EF" bold="true" box="[507,718,922,947]" italics="true" pageId="60" pageNumber="57">Leiodes toyoshimai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A207525B323C075575E641AFFC8EA7E8" box="[725,811,924,948]" pageId="60" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35A60B9323C075576814189FD68A7B4" pageId="60" pageNumber="57" type="description">
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||
<paragraph id="8BFF3332323C075576814189FD68A7B4" blockId="60.[434,844,922,1000]" pageId="60" pageNumber="57">
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||
Japanese name: Tsumebuto-ô-tamakinokomushi (
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<figureCitation id="137B2FB7323C0755750441E7FD3AA7BB" box="[567,671,980,999]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="61.[114,145,1223,1242]" captionStartId-1="62.[151,182,1279,1298]" captionStartId-2="63.[113,144,1495,1514]" captionTargetBox-0="[170,1031,228,1201]" captionTargetBox-1="[208,1066,232,1252]" captionTargetBox-2="[145,1059,591,1463]" captionTargetId-0="figure-126@61.[164,1038,219,1209]" captionTargetId-1="figure-11@62.[203,1074,219,1262]" captionTargetId-2="figure-153@63.[137,1065,577,1476]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionTargetPageId-2="63" captionText-0="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." captionText-1="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." captionText-2="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi-0="http://doi.org/10.5281/zenodo.4272535" figureDoi-1="http://doi.org/10.5281/zenodo.4272537" figureDoi-2="http://doi.org/10.5281/zenodo.4272539" httpUri-0="https://zenodo.org/record/4272535/files/figure.png" httpUri-1="https://zenodo.org/record/4272537/files/figure.png" httpUri-2="https://zenodo.org/record/4272539/files/figure.png" pageId="60" pageNumber="57">Figs. 33–35</figureCitation>
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,
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<figureCitation id="137B2FB7323C0755759441E6FD62A7B4" box="[679,711,981,1000]" captionStart="Fig" captionStartId="163.[113,145,1438,1457]" captionTargetBox="[113,579,941,1423]" captionTargetId="figure-125@163.[114,577,942,1419]" captionTargetPageId="163" captionText="Fig. 112. Distribution of Leiodes koreana group in Japan. Solid line – L. masatsugui sp. nov.; broken line – L. koreana Park & Ahn, 2007; (●) L. toyoshimai sp. nov.; arrow – suggested migration pathway of L.koreana group from the continent to Japan." figureDoi="http://doi.org/10.5281/zenodo.4272691" httpUri="https://zenodo.org/record/4272691/files/figure.png" pageId="60" pageNumber="57">112</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35A60B9323C075577A441CDFC18A15D" pageId="60" pageNumber="57" type="materials_examined">
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||
<paragraph id="8BFF3332323C075577A441CDFCB8A04D" blockId="60.[151,1127,1021,1282]" box="[151,797,1021,1042]" pageId="60" pageNumber="57">
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||
<emphasis id="B934EF20323C075577A441CDFEAEA04D" bold="true" box="[151,267,1021,1041]" pageId="60" pageNumber="57">
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||
<typeStatus id="54FB8D90323C075577A441CDFF61A04D" box="[151,196,1022,1041]" pageId="60" pageNumber="57">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingCountry id="F35773A2323C0755762541CCFEECA04E" box="[278,329,1023,1042]" name="Japan" pageId="60" pageNumber="57">Japan</collectingCountry>
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, Honshu,
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<collectingRegion id="4984FDD0323C075576AD41CDFE52A04D" box="[414,503,1022,1041]" country="Japan" name="Gifu" pageId="60" pageNumber="57">Gifu Pref.</collectingRegion>
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, Shirakawa Village, Ô–shirakawa.
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</paragraph>
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<paragraph id="8BFF3332323C075577A4462AFC18A15D" blockId="60.[151,1127,1021,1282]" pageId="60" pageNumber="57">
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||
<emphasis id="B934EF20323C075577A4462AFEBAA070" bold="true" box="[151,287,1048,1068]" pageId="60" pageNumber="57">Type material.</emphasis>
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||
<materialsCitation id="3B28396F323C07557614462BFCD4A01B" ID-GBIF-Occurrence="2984087339" collectingDate="2004-07-31" collectionCode="FIT, MNHAH, FUFJ" collectorName="K. Toyoshima" country="Japan" location="Shirakawa Village" pageId="60" pageNumber="57" specimenCount="5" specimenCount-female="5" stateProvince="Gifu" typeStatus="holotype">
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||
<emphasis id="B934EF20323C07557614462BFE6DA070" bold="true" box="[295,456,1048,1068]" pageId="60" pageNumber="57">
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||
<collectingCountry id="F35773A2323C07557614462BFECAA070" box="[295,367,1048,1068]" name="Japan" pageId="60" pageNumber="57">JAPAN</collectingCountry>
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||
: HONSHU:
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||
</emphasis>
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||
<typeStatus id="54FB8D90323C075576E2462AFD81A077" box="[465,548,1049,1068]" pageId="60" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
|
||
, ♂,
|
||
<collectingRegion id="4984FDD0323C0755757C462AFD0EA070" box="[591,683,1049,1068]" country="Japan" name="Gifu" pageId="60" pageNumber="57">Gifu Pref.</collectingRegion>
|
||
,
|
||
<location id="8E9F65E9323C07557586462AFCFDA070" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C07557586462AFCFDA070" box="[693,856,1049,1068]" country="Japan" name="Shirakawa Village" pageId="60" pageNumber="57" stateProvince="Gifu">Shirakawa Village</location>
|
||
, Ô–shirakawa,
|
||
<date id="FFFE15F2323C075574D1462AFBE3A070" box="[994,1094,1049,1068]" pageId="60" pageNumber="57" value="2004-07-31">
|
||
<collectingDate id="EFBAEC1A323C075574D1462AFBE3A070" box="[994,1094,1049,1068]" pageId="60" pageNumber="57" value="2004-07-31">31.vii.2004</collectingDate>
|
||
</date>
|
||
,
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||
<collectorName id="26B556E4323C07557362462AFF50A01B" pageId="60" pageNumber="57">K. Toyoshima</collectorName>
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||
leg. (
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||
<collectionCode id="ED51ABF7323C075576104607FEE0A01B" box="[291,325,1076,1095]" pageId="60" pageNumber="57">FIT</collectionCode>
|
||
) (
|
||
<collectionCode id="ED51ABF7323C075576664607FE0EA01B" box="[341,427,1076,1095]" pageId="60" pageNumber="57">MNHAH</collectionCode>
|
||
).
|
||
<typeStatus id="54FB8D90323C075576844607FDA8A019" box="[439,525,1076,1095]" pageId="60" pageNumber="57" type="paratype">PARATYPES</typeStatus>
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||
, 1♂,
|
||
<specimenCount id="9D46F8BB323C0755750D4600FDCFA01B" box="[574,618,1072,1095]" pageId="60" pageNumber="57" type="female">5♀♀</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED51ABF7323C075574044607FCCEA01B" box="[823,875,1076,1095]" pageId="60" pageNumber="57">FUFJ</collectionCode>
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||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C0755744B4607FE4DA03D" ID-GBIF-Occurrence="2984087377" collectingDate="2004-07-24" collectionCode="FUFJ" collectorName="K. Toyoshima" country="Japan" location="Shirakawa Village" pageId="60" pageNumber="57" specimenCount="2" specimenCount-female="2" stateProvince="Gifu" typeStatus="holotype">
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||
<specimenCount id="9D46F8BB323C0755744B4607FC01A01B" box="[888,932,1072,1095]" pageId="60" pageNumber="57" type="female">2♀♀</specimenCount>
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||
, same data as holotype except for the date,
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||
<date id="FFFE15F2323C07557672467DFE07A03D" box="[321,418,1102,1121]" pageId="60" pageNumber="57" value="2004-07-24">
|
||
<collectingDate id="EFBAEC1A323C07557672467DFE07A03D" box="[321,418,1102,1121]" pageId="60" pageNumber="57" value="2004-07-24">24.vii.2004</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED51ABF7323C0755769D467DFE47A03D" box="[430,482,1102,1121]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075576C2467DFBC6A03D" ID-GBIF-Occurrence="2984087369" box="[497,1123,1099,1123]" collectingDate="2004-08-08" collectionCode="FUFJ" collectorName="K. Toyoshima" country="Japan" location="Shirakawa Village" pageId="60" pageNumber="57" specimenCount="3" specimenCount-female="3" stateProvince="Gifu" typeStatus="holotype">
|
||
1 ♂,
|
||
<specimenCount id="9D46F8BB323C07557528467DFDEEA03E" box="[539,587,1099,1122]" pageId="60" pageNumber="57" type="female">3 ♀♀</specimenCount>
|
||
, same data as holotype except for the date,
|
||
<date id="FFFE15F2323C075574F2467DFBB8A03D" box="[961,1053,1102,1121]" pageId="60" pageNumber="57" value="2004-08-08">
|
||
<collectingDate id="EFBAEC1A323C075574F2467DFBB8A03D" box="[961,1053,1102,1121]" pageId="60" pageNumber="57" value="2004-08-08">8.viii.2004</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED51ABF7323C0755731A467DFBF8A03D" box="[1065,1117,1102,1121]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075577A4465AFD4CA020" ID-GBIF-Occurrence="2984087317" box="[151,745,1126,1149]" collectingDate="2004-08-22" collectionCode="FUFJ" collectorName="K. Toyoshima" country="Japan" location="Shirakawa Village" pageId="60" pageNumber="57" specimenCount="3" specimenCount-female="3" stateProvince="Gifu" typeStatus="holotype">
|
||
<specimenCount id="9D46F8BB323C075577A4465AFF63A021" box="[151,198,1126,1149]" pageId="60" pageNumber="57" type="female">3 ♀♀</specimenCount>
|
||
, same data as holotype except for the date,
|
||
<date id="FFFE15F2323C0755750E465AFD01A020" box="[573,676,1129,1148]" pageId="60" pageNumber="57" value="2004-08-22">
|
||
<collectingDate id="EFBAEC1A323C0755750E465AFD01A020" box="[573,676,1129,1148]" pageId="60" pageNumber="57" value="2004-08-22">22.viii.2004</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED51ABF7323C0755759C465AFD46A020" box="[687,739,1129,1148]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075575C1465AFBC6A0CB" ID-GBIF-Occurrence="2984087441" collectingDate="2004-07-30" collectingDateMax="2004-08-07" collectingDateMin="2004-07-30" collectionCode="FIT, FUFJ" collectorName="K. Arai & S. Arai" country="Japan" elevation="1300" location="Nakatsukawa-keikoku" municipality="Ohtaki Village" pageId="60" pageNumber="57" specimenCount="3" specimenCount-female="3" stateProvince="Saitama" typeStatus="holotype">
|
||
2 ♂♂,
|
||
<specimenCount id="9D46F8BB323C0755741F465AFCF9A021" box="[812,860,1126,1149]" pageId="60" pageNumber="57" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<collectingRegion id="4984FDD0323C07557454465AFC78A020" box="[871,989,1129,1148]" country="Japan" name="Saitama" pageId="60" pageNumber="57">Saitama Pref.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B9BA948323C075574D7465AFBC1A020" box="[996,1124,1129,1148]" pageId="60" pageNumber="57">Ohtaki Village</collectingMunicipality>
|
||
,
|
||
<location id="8E9F65E9323C075577A446B7FEFDA0CB" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C075577A446B7FEFDA0CB" box="[151,344,1156,1175]" country="Japan" municipality="Ohtaki Village" name="Nakatsukawa-keikoku" pageId="60" pageNumber="57" stateProvince="Saitama">Nakatsukawa-keikoku</location>
|
||
, Oku-Chichibu-rindô (alt.
|
||
<quantity id="4CB89ED7323C0755750946B7FDD9A0CB" box="[570,636,1156,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="60" pageNumber="57" unit="m" value="1300.0">
|
||
<elevation id="006DD401323C0755750946B7FDD9A0CB" box="[570,636,1156,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="60" pageNumber="57" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="FFFE15F2323C075575B846B7FC8EA0CB" box="[651,811,1156,1175]" pageId="60" pageNumber="57" value="2004-07-30" valueMax="2004-08-07" valueMin="2004-07-30">
|
||
<collectingDate id="EFBAEC1A323C075575B846B7FC8EA0CB" box="[651,811,1156,1175]" pageId="60" pageNumber="57" value="2004-07-30" valueMax="2004-08-07" valueMin="2004-07-30">30.vii.–7.viii.2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B556E4323C0755740046B7FCD7A0CB" box="[819,882,1156,1175]" pageId="60" pageNumber="57">K. Arai</collectorName>
|
||
&
|
||
<collectorName id="26B556E4323C075574BF46B7FC62A0CB" box="[908,967,1156,1175]" pageId="60" pageNumber="57">S. Arai</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51ABF7323C075574C546B7FBBDA0CB" box="[1014,1048,1156,1175]" pageId="60" pageNumber="57">FIT</collectionCode>
|
||
) (
|
||
<collectionCode id="ED51ABF7323C0755731A46B7FBF8A0CB" box="[1065,1117,1156,1175]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075577A446ADFF72A090" ID-GBIF-Occurrence="2984087416" collectingDate="2004-07-17" collectingDateMax="2004-07-24" collectingDateMin="2004-07-17" collectionCode="FIT, FUFJ" collectorName="K. Arai & S. Arai" country="Japan" elevation="1200" location="Mt. Arimayama" municipality="Naguri Village" pageId="60" pageNumber="57" specimenCount="1" specimenCount-female="1" stateProvince="Saitama" typeStatus="holotype">
|
||
<specimenCount id="9D46F8BB323C075577A446ADFF1DA0EE" box="[151,184,1179,1202]" pageId="60" pageNumber="57" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="4984FDD0323C075577F046ADFE9FA0ED" box="[195,314,1182,1201]" country="Japan" name="Saitama" pageId="60" pageNumber="57">Saitama Pref.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B9BA948323C0755767146ADFE66A0ED" box="[322,451,1182,1201]" pageId="60" pageNumber="57">Naguri Village</collectingMunicipality>
|
||
,
|
||
<location id="8E9F65E9323C075576FF46ACFDF1A0ED" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C075576FF46ACFDF1A0ED" box="[460,596,1182,1202]" country="Japan" municipality="Naguri Village" name="Mt. Arimayama" pageId="60" pageNumber="57" stateProvince="Saitama">Mt. Arimayama</location>
|
||
(alt.
|
||
<quantity id="4CB89ED7323C075575B246ADFD60A0ED" box="[641,709,1182,1201]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="57" unit="m" value="1200.0">
|
||
<elevation id="006DD401323C075575B246ADFD60A0ED" box="[641,709,1182,1201]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="57" unit="m" value="1200.0">1200 m</elevation>
|
||
</quantity>
|
||
),
|
||
<date id="FFFE15F2323C075575E646ADFCF8A0ED" box="[725,861,1182,1201]" pageId="60" pageNumber="57" value="2004-07-17" valueMax="2004-07-24" valueMin="2004-07-17">
|
||
<collectingDate id="EFBAEC1A323C075575E646ADFCF8A0ED" box="[725,861,1182,1201]" pageId="60" pageNumber="57" value="2004-07-17" valueMax="2004-07-24" valueMin="2004-07-17">17.–24.vii.2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B556E4323C0755745446ACFC03A0ED" box="[871,934,1182,1202]" pageId="60" pageNumber="57">K. Arai</collectorName>
|
||
and
|
||
<collectorName id="26B556E4323C075574E246ADFBA8A0ED" box="[977,1037,1182,1201]" pageId="60" pageNumber="57">S. Arai</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51ABF7323C0755730D46ADFBC5A0ED" box="[1086,1120,1182,1201]" pageId="60" pageNumber="57">FIT</collectionCode>
|
||
) (
|
||
<collectionCode id="ED51ABF7323C075577AE468AFF74A090" box="[157,209,1209,1228]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075577EC468AFC4CA090" ID-GBIF-Occurrence="2984087321" box="[223,1001,1205,1229]" collectingDate="2007-08-17" collectingDateMax="2007-08-27" collectingDateMin="2007-08-17" collectionCode="FUFJ" collectorName="T. Watanabe" country="Japan" location="Owakudani" municipality="Hakone Town" pageId="60" pageNumber="57" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
|
||
1 ♂,
|
||
<collectingRegion id="4984FDD0323C0755763B468AFE36A090" box="[264,403,1209,1228]" country="Japan" name="Kanagawa" pageId="60" pageNumber="57">Kanagawa Pref.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B9BA948323C075576AA468AFDB4A090" box="[409,529,1209,1228]" pageId="60" pageNumber="57">Hakone Town</collectingMunicipality>
|
||
,
|
||
<location id="8E9F65E9323C07557529468BFDDAA090" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C07557529468BFDDAA090" box="[538,639,1208,1228]" country="Japan" municipality="Hakone Town" name="Owakudani" pageId="60" pageNumber="57" stateProvince="Kanagawa">Ôwakudani</location>
|
||
,
|
||
<date id="FFFE15F2323C075575B5468AFCB5A090" box="[646,784,1209,1228]" pageId="60" pageNumber="57" value="2007-08-17" valueMax="2007-08-27" valueMin="2007-08-17">
|
||
<collectingDate id="EFBAEC1A323C075575B5468AFCB5A090" box="[646,784,1209,1228]" pageId="60" pageNumber="57" value="2007-08-17" valueMax="2007-08-27" valueMin="2007-08-17">17.-27.viii.2007</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B556E4323C0755742B468AFC24A090" box="[792,897,1209,1228]" pageId="60" pageNumber="57">T. Watanabe</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51ABF7323C0755749C468AFC46A090" box="[943,995,1209,1228]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C075574C2468AFCC3A0BB" ID-GBIF-Occurrence="2984087450" collectingDate="2002-08-02" collectingDateMax="2002-08-22" collectingDateMin="2002-08-02" collectionCode="FIT, FUFJ" collectorName="H. Hoshina" country="Japan" location="Sarukabe-entei" municipality="Mts. Hakusan" pageId="60" pageNumber="57" specimenCount="1" stateProvince="Ishikawa" typeStatus="holotype">
|
||
1 ♂,
|
||
<collectingRegion id="4984FDD0323C0755732A468AFF61A0BB" country="Japan" name="Ishikawa" pageId="60" pageNumber="57">Ishikawa Pref.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B9BA948323C075577F846E7FEE1A0BB" box="[203,324,1236,1255]" pageId="60" pageNumber="57">Mts. Hakusan</collectingMunicipality>
|
||
,
|
||
<location id="8E9F65E9323C0755767E46E7FE6AA0BB" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C0755767E46E7FE6AA0BB" box="[333,463,1236,1255]" country="Japan" municipality="Mts. Hakusan" name="Sarukabe-entei" pageId="60" pageNumber="57" stateProvince="Ishikawa">Sarukabe-entei</location>
|
||
,
|
||
<date id="FFFE15F2323C075576EB46E7FDF9A0BB" box="[472,604,1236,1255]" pageId="60" pageNumber="57" value="2002-08-02" valueMax="2002-08-22" valueMin="2002-08-02">
|
||
<collectingDate id="EFBAEC1A323C075576EB46E7FDF9A0BB" box="[472,604,1236,1255]" pageId="60" pageNumber="57" value="2002-08-02" valueMax="2002-08-22" valueMin="2002-08-02">2.–22.viii.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B556E4323C0755755646E7FD63A0BB" box="[613,710,1236,1255]" pageId="60" pageNumber="57">H. Hoshina</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51ABF7323C075575C446E7FCBCA0BB" box="[759,793,1236,1255]" pageId="60" pageNumber="57">FIT</collectionCode>
|
||
) (
|
||
<collectionCode id="ED51ABF7323C0755741846E7FCFAA0BB" box="[811,863,1236,1255]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B28396F323C0755745C46E7FC1CA15D" ID-GBIF-Occurrence="2984087465" collectingDate="2006-08-05" collectingDateMax="2006-08-12" collectingDateMin="2006-08-05" collectionCode="FUFJ" collectorName="K. Tanaka" country="Japan" county="Higashi-iyayama Village" location="Otome-dani Valley" municipality="Nagoro" pageId="60" pageNumber="57" specimenCount="1" specimenCount-female="1" stateProvince="Tokushima" typeStatus="holotype">
|
||
3 ♂♂,
|
||
<specimenCount id="9D46F8BB323C0755749A46E7FC6CA0BB" box="[937,969,1232,1255]" pageId="60" pageNumber="57" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="4984FDD0323C075574E746E7FBC0A0BB" box="[980,1125,1236,1255]" country="Japan" name="Tokushima" pageId="60" pageNumber="57">Tokushima Pref.</collectingRegion>
|
||
,
|
||
<collectingCounty id="629E4BBE323C075577A446DDFECBA15D" box="[151,366,1262,1281]" pageId="60" pageNumber="57">Higashi-iyayama Village</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B9BA948323C0755764546DCFE1DA15E" box="[374,440,1263,1282]" pageId="60" pageNumber="57">Nagoro</collectingMunicipality>
|
||
,
|
||
<location id="8E9F65E9323C075576F246DDFDC6A15D" LSID="urn:lsid:plazi:treatment:03E98224323C072976E041A8FE5EA0AF:8E9F65E9323C075576F246DDFDC6A15D" box="[449,611,1262,1281]" country="Japan" county="Higashi-iyayama Village" municipality="Nagoro" name="Otome-dani Valley" pageId="60" pageNumber="57" stateProvince="Tokushima">Otome-dani Valley</location>
|
||
,
|
||
<date id="FFFE15F2323C0755755846DDFD4AA15D" box="[619,751,1262,1281]" pageId="60" pageNumber="57" value="2006-08-05" valueMax="2006-08-12" valueMin="2006-08-05">
|
||
<collectingDate id="EFBAEC1A323C0755755846DDFD4AA15D" box="[619,751,1262,1281]" pageId="60" pageNumber="57" value="2006-08-05" valueMax="2006-08-12" valueMin="2006-08-05">5.–12.viii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B556E4323C075575CB46DCFCEAA15D" box="[760,847,1262,1282]" pageId="60" pageNumber="57">K. Tanaka</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED51ABF7323C0755744C46DDFC16A15D" box="[895,947,1262,1281]" pageId="60" pageNumber="57">FUFJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B9323C075577A4472AFCA0A1AE" pageId="60" pageNumber="57" type="diagnosis">
|
||
<paragraph id="8BFF3332323C075577A4472AFCA0A1AE" blockId="60.[151,1127,1305,1618]" pageId="60" pageNumber="57">
|
||
<emphasis id="B934EF20323C075577A4472AFEA9A16D" bold="true" box="[151,268,1305,1329]" pageId="60" pageNumber="57">Diagnosis.</emphasis>
|
||
Body
|
||
<quantity id="4CB89ED7323C075576664729FE70A16D" box="[341,469,1306,1330]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="60" pageNumber="57" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
long, ca. 1.8× as long as wide. Dorsum brown. Each elytron with nine distinct rows of punctures and subhumeral row as long as ca. 1/4 of elytral length. Mesoventrite with one distinct excavation between median carina and transverse carina. Median carina of mesoventrite low. Mesotibiae showing distinct sexual dimorphism. Male metafemora a little curved posteriad at posterior margins. Tarsomeres 2 and 3 of male protarsi and mesotarsi extremely expanded. Male metatibiae feebly curved inwards. Female abdominal sternite 8 with two distinct projections on anterior margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B9323C072977A447CAFD27A76F" lastPageId="64" lastPageNumber="61" pageId="60" pageNumber="57" type="description">
|
||
<paragraph id="8BFF3332323C075577A447CAFED9A20E" blockId="60.[151,1127,1305,1618]" pageId="60" pageNumber="57">
|
||
<emphasis id="B934EF20323C075577A447CAFE87A24D" bold="true" box="[151,290,1529,1553]" pageId="60" pageNumber="57">Description.</emphasis>
|
||
Measurements of
|
||
<typeStatus id="54FB8D90323C075576DB47C9FDEFA24E" box="[488,586,1530,1554]" pageId="60" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
: Body length
|
||
<quantity id="4CB89ED7323C075575EE47C9FC97A24D" box="[733,818,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="60" pageNumber="57" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; head
|
||
<quantity id="4CB89ED7323C0755744547C9FC71A24D" box="[886,980,1530,1554]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="60" pageNumber="57" unit="mm" value="0.47">0.47 mm</quantity>
|
||
in length and
|
||
<quantity id="4CB89ED7323C075577A44429FF50A26D" box="[151,245,1562,1586]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="60" pageNumber="57" unit="mm" value="0.87">0.87 mm</quantity>
|
||
in width; pronotum 1.0 mm in length and
|
||
<quantity id="4CB89ED7323C075575874429FCA0A26D" box="[692,773,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="60" pageNumber="57" unit="mm" value="1.6">1.6 mm</quantity>
|
||
in width; elytra
|
||
<quantity id="4CB89ED7323C075574814429FBA6A26D" box="[946,1027,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="60" pageNumber="57" unit="mm" value="1.9">1.9 mm</quantity>
|
||
in length and
|
||
<quantity id="4CB89ED7323C075577F74409FEB0A20D" box="[196,277,1594,1618]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="60" pageNumber="57" unit="mm" value="1.8">1.8 mm</quantity>
|
||
in width.
|
||
</paragraph>
|
||
<caption id="DF3F63BA323D0754774146F4FCC1A14C" ID-DOI="http://doi.org/10.5281/zenodo.4272535" ID-Zenodo-Dep="4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="61" pageNumber="54" startId="61.[114,145,1223,1242]" targetBox="[170,1031,228,1201]" targetPageId="61">
|
||
<paragraph id="8BFF3332323D0754774146F4FCC1A14C" blockId="61.[113,1090,1223,1296]" pageId="61" pageNumber="54">
|
||
Fig. 33.
|
||
<taxonomicName id="4C4048B1323D0754778A46FBFEC5A086" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[185,352,1224,1242]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="54" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
|
||
<emphasis id="B934EF20323D0754778A46FBFEC5A086" box="[185,352,1224,1242]" italics="true" pageId="61" pageNumber="54">Leiodes toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B323D0754765646F4FE0DA086" box="[357,424,1223,1242]" pageId="61" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF3332323D075477A34709FF5BA18E" blockId="61.[113,1089,1338,1618]" pageId="61" pageNumber="54">Coloration. Dorsum shining and almost unicolor, brown; antennomere 1 light brown; antennomeres 2–6 and 8 brown; antennomeres 7, 9, 10, and basal 2/3 of antennomere 11 blackish dark brown; apical 1/3 of antennomere 11 light brown; legs brownish; tarsi often light brown; mesoventrite and metaventrite brown; abdominal ventrites slightly paler than metaventrite.</paragraph>
|
||
<paragraph id="8BFF3332323D075477A347E9FD6CA1AE" blockId="61.[113,1089,1338,1618]" box="[144,713,1498,1522]" pageId="61" pageNumber="54">
|
||
Body
|
||
<quantity id="4CB89ED7323D075477E347E9FEF5A1AE" box="[208,336,1498,1522]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="61" pageNumber="54" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
in length, ca. 1.8× as long as wide.
|
||
</paragraph>
|
||
<paragraph id="8BFF3332323D075777A347C9FC00A184" blockId="61.[113,1089,1338,1618]" lastBlockId="62.[151,1127,1440,1624]" lastPageId="62" lastPageNumber="59" pageId="61" pageNumber="54">
|
||
Head ca. 1.8× as wide as long, ca. 0.48× as long as and 0.54× as wide as pronotum, densely and minutely punctate (
|
||
<figureCitation id="137B2FB7323D0754765A4429FE6EA26E" box="[361,459,1562,1586]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="61" pageNumber="54">Fig. 33A</figureCitation>
|
||
), usually bearing some large punctures (
|
||
<figureCitation id="137B2FB7323D075474434429FC77A26E" box="[880,978,1562,1586]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="61" pageNumber="54">Fig. 33A</figureCitation>
|
||
); antennomeres 1–3 each longer than wide; antennomeres 4 and 11 about as long as wide; remaining antennomeres each wider than long; antennomere 11 robust (
|
||
<figureCitation id="137B2FB7323E075774184793FC2BA1E4" box="[811,910,1440,1464]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="62" pageNumber="59">Fig. 33C</figureCitation>
|
||
); relative lengths of antennomeres 2 to 11 – 3.5: 3.7: 1.5: 1.5: 1.3: 2.7: 1.0: 3.8: 3.6: 3.7.
|
||
</paragraph>
|
||
<caption id="DF3F63BA323E075777A446CCFE9EA121" ID-DOI="http://doi.org/10.5281/zenodo.4272537" ID-Zenodo-Dep="4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="62" pageNumber="59" startId="62.[151,182,1279,1298]" targetBox="[208,1066,232,1252]" targetPageId="62">
|
||
<paragraph id="8BFF3332323E075777A446CCFE9EA121" blockId="62.[151,1128,1279,1405]" pageId="62" pageNumber="59">
|
||
Fig. 34.
|
||
<taxonomicName id="4C4048B1323E075777D34733FE23A14E" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[224,390,1280,1298]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="59" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
|
||
<emphasis id="B934EF20323E075777D34733FE23A14E" box="[224,390,1280,1298]" italics="true" pageId="62" pageNumber="59">Leiodes toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B323E075776BF46CCFE75A14E" box="[396,464,1279,1298]" pageId="62" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF3332323E0757778647D2FDDEA264" blockId="62.[151,1127,1440,1624]" pageId="62" pageNumber="59">
|
||
Pronotum ca. 1.6× as wide as long, ca. 0.51× as long as and 0.91× as wide as elytra, widest near base, simply and very feebly curved at posterior margin, distinctly and minutely punctate, punctation similar to that on head (
|
||
<figureCitation id="137B2FB7323E0757753F4413FDCAA264" box="[524,623,1568,1592]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="62" pageNumber="59">Fig. 33A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF3332323E075777864473FE55A204" blockId="62.[151,1127,1440,1624]" box="[181,496,1600,1624]" pageId="62" pageNumber="59">Scutellum minutely punctate.</paragraph>
|
||
<paragraph id="8BFF3332323F075677A342E8FD61A5CF" blockId="63.[113,1089,219,563]" pageId="63" pageNumber="56">
|
||
Elytra ca. 1.1× as long as wide in dorsal view, widest ca. at basal 1/4 (
|
||
<figureCitation id="137B2FB7323F0756749042E8FBACA4A8" box="[931,1033,219,244]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33A</figureCitation>
|
||
), not transversely strigose; each elytron bearing nine rows of punctures with small number of large punctures and dense very fine punctures between rows (
|
||
<figureCitation id="137B2FB7323F075675FB4328FC8CA568" box="[712,809,283,308]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33E</figureCitation>
|
||
); row 9 invisible in dorsal view, subhumeral row as long as ca. 1/4 of elytral length (
|
||
<figureCitation id="137B2FB7323F075675E94308FC99A508" box="[730,828,315,340]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33B</figureCitation>
|
||
); elytral rows composed of coarse or minute punctures, larger than those of pronotum (
|
||
<figureCitation id="137B2FB7323F075674204368FCDDA528" box="[787,888,347,372]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33A</figureCitation>
|
||
); sutural stria fine, arising from apex to ca. apical half of the elytral length.
|
||
</paragraph>
|
||
<paragraph id="8BFF3332323F075677A343A8FDBDA5EF" blockId="63.[113,1089,219,563]" box="[144,536,411,435]" pageId="63" pageNumber="56">Metathoracic wings fully developed.</paragraph>
|
||
<paragraph id="8BFF3332323F075677A34388FCCEA66F" blockId="63.[113,1089,219,563]" pageId="63" pageNumber="56">
|
||
Mesoventrite strongly microreticulate, impunctate, almost glabrous, and with one distinct excavation between median carina and transverse carina (
|
||
<figureCitation id="137B2FB7323F0756758D43E8FCBFA5A8" box="[702,794,475,500]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33F</figureCitation>
|
||
); median carina of mesoventrite low (
|
||
<figureCitation id="137B2FB7323F075677E443C8FE90A648" box="[215,309,507,532]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="63" pageNumber="56">Fig. 33F</figureCitation>
|
||
); metaventrite showing indistinct sexual dimorphism, sparsely pubescent, and distinctly microreticulate except for almost smooth middle portion.
|
||
</paragraph>
|
||
<caption id="DF3F63BA323F0756774247E4FDAFA27C" ID-DOI="http://doi.org/10.5281/zenodo.4272539" ID-Zenodo-Dep="4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="63" pageNumber="56" startId="63.[113,144,1495,1514]" targetBox="[145,1059,591,1463]" targetPageId="63">
|
||
<paragraph id="8BFF3332323F0756774247E4FDAFA27C" blockId="63.[113,1089,1495,1568]" pageId="63" pageNumber="56">
|
||
Fig. 35.
|
||
<taxonomicName id="4C4048B1323F0756778447EBFEFEA1B6" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[183,347,1496,1514]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="56" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
|
||
<emphasis id="B934EF20323F0756778447EBFEFEA1B6" box="[183,347,1496,1514]" italics="true" pageId="63" pageNumber="56">Leiodes toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B323F0756766C47E4FE04A1B6" box="[351,417,1495,1514]" pageId="63" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF333232400729778642EFFD74A568" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
Legs showing distinct sexual dimorphism on protarsi, protibiae, mesotarsi, metafemora, and metatibiae; mesotibiae showing indistinct sexual dimorphism; metafemur robust, with small dorsal posteroapical projection (
|
||
<figureCitation id="137B2FB732400729751C4328FD60A568" box="[559,709,283,308]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Figs. 34I, 34J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF33323240072977864309FD89A62F" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977864309FF50A508" bold="true" box="[181,245,314,340]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977864309FF48A50F" bold="true" box="[181,237,314,339]" italics="true" pageId="64" pageNumber="61">Male</emphasis>
|
||
.
|
||
</emphasis>
|
||
Metaventrite with relatively robust pubescence around middle portion (
|
||
<figureCitation id="137B2FB73240072974C54308FBFFA508" box="[1014,1114,315,340]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="64" pageNumber="61">Fig. 33G</figureCitation>
|
||
); protibiae gradually and strongly widening from base towards apex at internal margins (
|
||
<figureCitation id="137B2FB73240072973084368FF6CA5C8" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34G</figureCitation>
|
||
); tarsomeres 2–4 of protarsi and mesotarsi expanded and tarsomeres 2 and 3 extremely broadened (
|
||
<figureCitation id="137B2FB732400729762543A8FE64A5E8" box="[278,449,411,436]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Figs. 34A, 34C</figureCitation>
|
||
); mesotibiae relatively strongly broadening from base towards apex (
|
||
<figureCitation id="137B2FB73240072977EB4388FE9FA588" box="[216,314,443,468]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34C</figureCitation>
|
||
); metafemora a little curved posteriad at posterior margins (
|
||
<figureCitation id="137B2FB73240072974854388FBB2A588" box="[950,1047,443,468]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34E</figureCitation>
|
||
); metatibiae feebly curved inwards (
|
||
<figureCitation id="137B2FB73240072976EE43E8FD9AA5A8" box="[477,575,475,500]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34E</figureCitation>
|
||
); abdominal sternite 8 strongly curved (
|
||
<figureCitation id="137B2FB73240072974C743E8FBFFA5AF" box="[1012,1114,475,499]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35D</figureCitation>
|
||
); aedeagus relatively robust (
|
||
<figureCitation id="137B2FB732400729768C43C8FDCCA64F" box="[447,617,507,531]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Figs. 35A, 35B</figureCitation>
|
||
); median lobe triangular at apex in dorsal view (
|
||
<figureCitation id="137B2FB73240072977AC4028FEA6A66F" box="[159,259,539,563]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35A</figureCitation>
|
||
), strongly curved and pointed apically in lateral view (
|
||
<figureCitation id="137B2FB73240072974604028FC13A66F" box="[851,950,539,563]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35B</figureCitation>
|
||
); each paramere bearing two apical setae and transparent very small lobe at apex (
|
||
<figureCitation id="137B2FB73240072974644008FC19A60F" box="[855,956,571,595]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35A</figureCitation>
|
||
); inner sac with some large sclerites (
|
||
<figureCitation id="137B2FB732400729764B4068FDBAA62F" box="[376,543,603,627]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Figs. 35A, 35C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF33323240072977864049FD27A76F" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977864049FEAAA6C8" bold="true" box="[181,271,634,660]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977864049FEADA6CF" bold="true" box="[181,264,634,659]" italics="true" pageId="64" pageNumber="61">Female</emphasis>
|
||
.
|
||
</emphasis>
|
||
Metaventrite with relatively thin pubescence (
|
||
<figureCitation id="137B2FB73240072975C84048FCFAA6C8" box="[763,863,635,660]" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="64" pageNumber="61">Fig. 33H</figureCitation>
|
||
); protibiae gradually and very feebly widening from base towards apex at internal margins (
|
||
<figureCitation id="137B2FB732400729745F40A8FC74A6E8" box="[876,977,667,692]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34H</figureCitation>
|
||
); protarsi and mesotarsi slender (
|
||
<figureCitation id="137B2FB73240072976524088FDAFA688" box="[353,522,699,724]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Figs. 34B, 34D</figureCitation>
|
||
); mesotibiae relatively weakly broadening from base towards apex (
|
||
<figureCitation id="137B2FB732400729762940E8FEDEA6A8" box="[282,379,731,756]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34D</figureCitation>
|
||
); metafemora almost straight at posterior margins (
|
||
<figureCitation id="137B2FB73240072974B840E8FC42A6A8" box="[907,999,731,756]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34F</figureCitation>
|
||
); metatibiae almost straight (
|
||
<figureCitation id="137B2FB732400729767140C8FE04A748" box="[322,417,763,788]" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Fig. 34F</figureCitation>
|
||
); abdominal sternite 8 with two projections at anterior margin (
|
||
<figureCitation id="137B2FB732400729730840C8FF60A76F" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35E</figureCitation>
|
||
); coxites and stylus as shown in
|
||
<figureCitation id="137B2FB73240072975114128FDDBA76F" box="[546,638,795,819]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">
|
||
Fig.
|
||
<specimenCount id="9D46F8BB3240072975604128FDDBA76F" box="[595,638,795,819]" pageId="64" pageNumber="61" type="generic">35F</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93240072977A44108FE71A02F" pageId="64" pageNumber="61" type="diagnosis">
|
||
<paragraph id="8BFF33323240072977A44108FE71A02F" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977A44108FE2CA70F" bold="true" box="[151,393,827,851]" pageId="64" pageNumber="61">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C4048B13240072976A5410EFDC1A708" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[406,612,829,852]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
|
||
<emphasis id="B934EF203240072976A5410EFDC1A708" box="[406,612,829,852]" italics="true" pageId="64" pageNumber="61">Leiodes toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B324007297559410FFD19A70F" box="[618,700,828,851]" pageId="64" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C4048B1324007297470410EFC11A708" authorityName="Park & Ahn" authorityYear="2007" box="[835,948,829,852]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="koreana">
|
||
<emphasis id="B934EF20324007297470410EFC11A708" box="[835,948,829,852]" italics="true" pageId="64" pageNumber="61">L. koreana</emphasis>
|
||
</taxonomicName>
|
||
in elytral appearance, but can be distinguished from the latter by having a relatively small body (
|
||
<quantity id="4CB89ED73240072974EB4168FBFEA728" box="[984,1115,859,884]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="64" pageNumber="61" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
), mesoventrite with a distinct excavation between the median carina and transverse carina (
|
||
<figureCitation id="137B2FB732400729730F4148FF61A7E8" captionStart="Fig" captionStartId="61.[114,145,1223,1242]" captionTargetBox="[170,1031,228,1201]" captionTargetId="figure-126@61.[164,1038,219,1209]" captionTargetPageId="61" captionText="Fig. 33. Leiodes toyoshimai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – basal margin of pronotum; E – elytral punctures; F – mesoventrite, lateral view; G – male metaventrite; H – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for F; IV: 0.5 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272535" httpUri="https://zenodo.org/record/4272535/files/figure.png" pageId="64" pageNumber="61">Fig. 33F</figureCitation>
|
||
), and male protarsi and mesotarsi with extremely expanded tarsomeres 2 and 3 (
|
||
<figureCitation id="137B2FB732400729730241A8FEA1A788" captionStart="Fig" captionStartId="62.[151,182,1279,1298]" captionTargetBox="[208,1066,232,1252]" captionTargetId="figure-11@62.[203,1074,219,1262]" captionTargetPageId="62" captionText="Fig. 34. Leiodes toyoshimai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G – male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E and F; III: 0.2 mm for G and H; IV: 0.2 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272537" httpUri="https://zenodo.org/record/4272537/files/figure.png" pageId="64" pageNumber="61">Figs. 34A, 34C</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName id="4C4048B13240072976AA418EFDA9A788" authorityName="Park & Ahn" authorityYear="2007" box="[409,524,957,980]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="koreana">
|
||
<emphasis id="B934EF203240072976AA418EFDA9A788" box="[409,524,957,980]" italics="true" pageId="64" pageNumber="61">L. koreana</emphasis>
|
||
</taxonomicName>
|
||
has a relatively large body (
|
||
<quantity id="4CB89ED7324007297471418FFC6CA788" box="[834,969,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="64" pageNumber="61" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 mm</quantity>
|
||
), mesoventrite with a shallow excavation (
|
||
<figureCitation id="137B2FB732400729768841E8FDBCA7A8" box="[443,537,987,1012]" captionStart="Fig" captionStartId="53.[113,144,1542,1561]" captionTargetBox="[163,1048,625,1517]" captionTargetId="figure-153@53.[150,1052,609,1527]" captionTargetPageId="53" captionText="Fig. 27. Leiodes koreana Park & Ahn, 2007.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272523" httpUri="https://zenodo.org/record/4272523/files/figure.png" pageId="64" pageNumber="61">Fig. 27E</figureCitation>
|
||
) and tarsomeres 2 and 3 of male protarsi and mesotarsi moderately expanded (
|
||
<figureCitation id="137B2FB73240072976A241C8FE5DA048" box="[401,504,1019,1044]" captionStart="Fig" captionStartId="54.[151,182,1491,1510]" captionTargetBox="[208,1061,585,1469]" captionTargetId="figure-171@54.[203,1073,561,1475]" captionTargetPageId="54" captionText="Fig. 28. Leiodes koreana Park & Ahn, 2007. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male hind leg, ventral view; D – female hind leg, ventral view; E – male protibia, dorsal view; F – female protibia, dorsal view; G – male metafemur, dorsal view; H – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 0.2 mm for E and F; III: 0.2 mm for G and H." figureDoi="http://doi.org/10.5281/zenodo.4272525" httpUri="https://zenodo.org/record/4272525/files/figure.png" pageId="64" pageNumber="61">Fig. 28A</figureCitation>
|
||
).
|
||
<taxonomicName id="4C4048B132400729753E41CEFD7AA048" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[525,735,1021,1044]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="toyoshimai" status="sp. nov.">
|
||
<emphasis id="B934EF2032400729753E41CEFD7AA048" box="[525,735,1021,1044]" italics="true" pageId="64" pageNumber="61">Leiodes toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B3240072975DA41CFFC9AA04F" box="[745,831,1020,1043]" pageId="64" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is also similar to
|
||
<taxonomicName id="4C4048B132400729733941CEFEF1A068" authority="(Kugelann, 1794)" baseAuthorityName="Kugelann" baseAuthorityYear="1794" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="B934EF2032400729733941CEFBC2A048" box="[1034,1127,1021,1044]" italics="true" pageId="64" pageNumber="61">L. dubia</emphasis>
|
||
(Kugelann, 1794)
|
||
</taxonomicName>
|
||
inhabiting Europe and the Russian Far East by having a robust body, but can be separated from it by having a relatively robust aedeagus (
|
||
<figureCitation id="137B2FB73240072974654608FC19A00F" box="[854,956,1083,1107]" captionStart="Fig" captionStartId="63.[113,144,1495,1514]" captionTargetBox="[145,1059,591,1463]" captionTargetId="figure-153@63.[137,1065,577,1476]" captionTargetPageId="63" captionText="Fig. 35. Leiodes toyoshimai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – apex of aedeagus, ventral view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A–C; II: 0.2 mm for D and E; III: 0.1 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272539" httpUri="https://zenodo.org/record/4272539/files/figure.png" pageId="64" pageNumber="61">Fig. 35A</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName id="4C4048B1324007297362460EFF71A028" baseAuthorityName="Kugelann" baseAuthorityYear="1794" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="B934EF20324007297362460EFF71A028" italics="true" pageId="64" pageNumber="61">L. dubia</emphasis>
|
||
</taxonomicName>
|
||
has a slender aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93240072977A4464FFD01A0EF" pageId="64" pageNumber="61" type="etymology">
|
||
<paragraph id="8BFF33323240072977A4464FFD01A0EF" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977A4464FFEBDA0C8" bold="true" box="[151,280,1148,1172]" pageId="64" pageNumber="61">Etymology.</emphasis>
|
||
This species is dedicated to Mr. Kentarô Toyoshima, who kindly gave me many valuable specimens of
|
||
<taxonomicName id="4C4048B13240072976B846AEFE7BA0E8" authorityName=": HATCH" authorityYear="1929" box="[395,478,1181,1204]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B934EF203240072976B846AEFE7BA0E8" box="[395,478,1181,1204]" italics="true" pageId="64" pageNumber="61">Leiodes</emphasis>
|
||
</taxonomicName>
|
||
used in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93240072977A44688FE5EA0AF" pageId="64" pageNumber="61" type="distribution">
|
||
<paragraph id="8BFF33323240072977A44688FE5EA0AF" blockId="64.[151,1127,219,1267]" pageId="64" pageNumber="61">
|
||
<emphasis id="B934EF203240072977A44688FE8EA08F" bold="true" box="[151,299,1211,1235]" pageId="64" pageNumber="61">Distribution.</emphasis>
|
||
<collectingCountry id="F35773A2324007297606468FFED3A08F" box="[309,374,1212,1235]" name="Japan" pageId="64" pageNumber="61">Japan</collectingCountry>
|
||
: Honshu (
|
||
<collectingRegion id="4984FDD03240072976DA4688FDE1A08F" box="[489,580,1211,1235]" country="Japan" name="Saitama" pageId="64" pageNumber="61">Saitama</collectingRegion>
|
||
,
|
||
<collectingRegion id="4984FDD0324007297561468FFD63A088" box="[594,710,1212,1236]" country="Japan" name="Kanagawa" pageId="64" pageNumber="61">Kanagawa</collectingRegion>
|
||
,
|
||
<collectingRegion id="4984FDD03240072975E64688FCACA08F" box="[725,777,1211,1235]" country="Japan" name="Gifu" pageId="64" pageNumber="61">Gifu</collectingRegion>
|
||
, and
|
||
<collectingRegion id="4984FDD032400729747B4688FC0CA08F" box="[840,937,1211,1235]" country="Japan" name="Ishikawa" pageId="64" pageNumber="61">Ishikawa</collectingRegion>
|
||
Prefectures) and Shikoku (
|
||
<collectingRegion id="4984FDD03240072977CD46E8FE4AA0AF" box="[254,495,1243,1267]" country="Japan" name="Tokushima" pageId="64" pageNumber="61">Tokushima Prefecture</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |