328 lines
57 KiB
XML
328 lines
57 KiB
XML
<document id="86C2A33467FB8D168C5400C64A83FE10" ID-CLB-Dataset="9554" ID-DOI="10.5281/zenodo.4272467" ID-GBIF-Dataset="59fc10fe-e8c3-4537-abb9-049fbda4c7ed" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272467" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298451113" checkinUser="felipe" docAuthor="Hoshina, Hideto" docDate="2012" docId="03E98224322C075976DF46E8FD33A248" docLanguage="en" docName="ActaEntMusNatPra.52.suppl.1-168.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl. 1) 52" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Leiodes yasudai Hoshina 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="FFD0FA5C3200076977334233FFA5A45C" masterDocTitle="Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)" masterLastPageNumber="168" masterPageNumber="1" pageNumber="41" updateTime="1698846259177" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="BB0607790F776297CDE897F8920933E2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D5E42BE3592673AC1479C13C3C1118DE">
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<mods:title id="FAF6E2DF071FF4AB6CD4CFFC97CB4463">Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="E4A204BFC85F45A664D4D325BE99FB72" type="personal">
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<mods:role id="874DFAAF0EAE92B9136B0A844297F127">
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<mods:roleTerm id="41A701CCE57568F5F0CCE7F739D40CAA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="32E6B05E585AEEC19A0D84A508BFBE9B">Hoshina, Hideto</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DEE7C58239C6B6B6DF0794936377FC1F">text</mods:typeOfResource>
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<mods:titleInfo id="420CD5149625B22AB60BE62230D45EA7">
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<mods:title id="354B0648ACBD4918658F942F775645A0">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="DF8A93608C97C101AACE23CFBD969C4D">
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<mods:date id="4665541DFCB0730313291ED55E9C72C4">2012</mods:date>
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<mods:detail id="B57A137C12E53DE2F63DBE6D948E98D4" type="series">
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<mods:title id="B63F4C6A15975D78C68C43135BC4A301">suppl. 1</mods:title>
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</mods:detail>
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<mods:detail id="7B15F3F52B007E34D6B0AF000798BFBE" type="pubDate">
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<mods:number id="76EBB17EB99CB60656F089A407E70948">2012-09-15</mods:number>
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</mods:detail>
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<mods:detail id="F9F0B39EAE3924332260C522838B5CD6" type="volume">
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<mods:number id="7329A6D01588DA03EE0E58D6F92D641A">52</mods:number>
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</mods:detail>
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<mods:extent id="65EB31189850563F8FC0A65B0D486A39" unit="page">
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<mods:start id="398BBC2E72954DE05D5748AE0941C9B3">1</mods:start>
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<mods:end id="C8CB45EF6F52B47C18D77F4B54D63CED">168</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="FDCD65104B9DF74578A6016C605D1F44">journal article</mods:classification>
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<mods:identifier id="D694AE60A55125A91E89A7C70B92B8F3" type="CLB-Dataset">9554</mods:identifier>
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<mods:identifier id="E5672D0E429852EBBB62057491E20422" type="DOI">10.5281/zenodo.4272467</mods:identifier>
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<mods:identifier id="BF90AEC410EBD44030D1BAD26A79F5E3" type="GBIF-Dataset">59fc10fe-e8c3-4537-abb9-049fbda4c7ed</mods:identifier>
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<mods:identifier id="D44F3A83B36C1F44BD16079B57883A08" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="D508279FD9A6F4DFA0F6E37B157D7D07" type="Zenodo-Dep">4272467</mods:identifier>
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<treatment id="03E98224322C075976DF46E8FD33A248" ID-DOI="http://doi.org/10.5281/zenodo.4339425" ID-GBIF-Taxon="170447929" ID-Zenodo-Dep="4339425" LSID="urn:lsid:plazi:treatment:03E98224322C075976DF46E8FD33A248" httpUri="http://treatment.plazi.org/id/03E98224322C075976DF46E8FD33A248" lastPageId="48" lastPageNumber="45" pageId="44" pageNumber="41">
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<subSubSection id="C35A60B9322C074576DF46E8FCB7A0A8" box="[492,786,1242,1268]" pageId="44" pageNumber="41" type="nomenclature">
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<paragraph id="8BFF3332322C074576DF46E8FCB7A0A8" blockId="44.[444,834,1242,1320]" box="[492,786,1242,1268]" pageId="44" pageNumber="41">
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<heading id="D0B7845E322C074576DF46E8FCB7A0A8" bold="true" box="[492,786,1242,1268]" centered="true" fontSize="10" level="2" pageId="44" pageNumber="41" reason="2">
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<emphasis id="B934EF20322C074576DF46E8FCB7A0A8" bold="true" box="[492,786,1242,1268]" pageId="44" pageNumber="41">
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6.
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<taxonomicName id="4C4048B1322C0745753546E9FD13A0AF" ID-CoL="3SYKJ" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[518,694,1242,1267]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="41" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
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<emphasis id="B934EF20322C0745753546E9FD13A0AF" bold="true" box="[518,694,1242,1267]" italics="true" pageId="44" pageNumber="41">Leiodes yasudai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A207525B322C0745758E46EFFCB7A0A8" box="[701,786,1244,1268]" pageId="44" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35A60B9322C0745768F46C9FD72A174" pageId="44" pageNumber="41" type="description">
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<paragraph id="8BFF3332322C0745768F46C9FD72A174" blockId="44.[444,834,1242,1320]" pageId="44" pageNumber="41">
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Japanese name: Daisetsu-ô-tamakinokomushi (
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<figureCitation id="137B2FB7322C0745751F4727FDCCA17B" box="[556,617,1300,1319]" captionStart="Fig" captionStartId="17.[111,142,1500,1519]" captionTargetBox="[150,1053,496,1471]" captionTargetId="figure-116@17.[141,1061,491,1471]" captionTargetPageId="17" captionText="Fig. 6. Heads (A–I) and mandibles (J and K) of Leiodes spp. A – Leiodes yasudai sp. nov.; B – L. multipunctata (Rye, 1873); C – L. irregularis Hatch, 1929; D – L. osawai Nakane, 1963; E – L. yukihikoi sp. nov.; F – L. okawai Nakane, 1963; G – L. tanakai sp. nov.; H, J, K – L. lucens (Fairmaire, 1855); I – L. yoshidai sp. nov. Scale I: 0.5 mm for A–I; II: 0.2 mm for J and K." figureDoi="http://doi.org/10.5281/zenodo.4272479" httpUri="https://zenodo.org/record/4272479/files/figure.png" pageId="44" pageNumber="41">Figs. 6</figureCitation>
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,
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<figureCitation id="137B2FB7322C074575414726FD0CA174" box="[626,681,1301,1320]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="45.[112,143,1516,1535]" captionStartId-1="46.[151,182,1464,1483]" captionStartId-2="47.[113,144,1191,1210]" captionTargetBox-0="[152,1047,472,1486]" captionTargetBox-1="[177,1094,236,1406]" captionTargetBox-2="[136,1062,228,1156]" captionTargetId-0="figure-100@45.[147,1056,465,1487]" captionTargetId-1="figure-11@46.[164,1113,219,1418]" captionTargetId-2="figure-106@47.[121,1080,219,1171]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionTargetPageId-2="47" captionText-0="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." captionText-1="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." captionText-2="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi-0="http://doi.org/10.5281/zenodo.4272511" figureDoi-1="http://doi.org/10.5281/zenodo.4272513" figureDoi-2="http://doi.org/10.5281/zenodo.4272515" httpUri-0="https://zenodo.org/record/4272511/files/figure.png" httpUri-1="https://zenodo.org/record/4272513/files/figure.png" httpUri-2="https://zenodo.org/record/4272515/files/figure.png" pageId="44" pageNumber="41">21–23</figureCitation>
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,
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<figureCitation id="137B2FB7322C074575814726FD74A174" box="[690,721,1301,1320]" captionStart="Fig" captionStartId="163.[610,643,761,780]" captionTargetBox="[602,1118,210,759]" captionTargetId="figure-18@163.[608,1090,221,747]" captionTargetPageId="163" captionText="Fig. 111. Distribution of Leiodes circinipes species group: (●) L. yasudai sp. nov., (■) L. juzoi sp. nov., (▲) L. yoshitakei sp. nov., (*) L. circinipes (Rye, 1873); circle by broken line – Mts. Daisetsu; arrow – suggested migration pathway of L. circinipes group from the continent to Japan." figureDoi="http://doi.org/10.5281/zenodo.4272689" httpUri="https://zenodo.org/record/4272689/files/figure.png" pageId="44" pageNumber="41">111</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A60B9322C074577A4470DFC1FA1FD" pageId="44" pageNumber="41" type="materials_examined">
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<paragraph id="8BFF3332322C074577A4470DFD0BA10D" blockId="44.[151,1128,1341,1442]" box="[151,686,1341,1362]" pageId="44" pageNumber="41">
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<emphasis id="B934EF20322C074577A4470DFEB5A10D" bold="true" box="[151,272,1341,1361]" pageId="44" pageNumber="41">
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<typeStatus id="54FB8D90322C074577A4470DFF61A10D" box="[151,196,1342,1361]" pageId="44" pageNumber="41">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F35773A2322C07457626470CFEEDA10E" box="[277,328,1343,1362]" name="Japan" pageId="44" pageNumber="41">Japan</collectingCountry>
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,
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<collectingRegion id="4984FDD0322C07457663470DFE03A10D" box="[336,422,1342,1361]" country="Japan" name="Hokkaido" pageId="44" pageNumber="41">Hokkaido</collectingRegion>
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, Mts. Daisetsu, Mt. Kurodake.
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</paragraph>
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<paragraph id="8BFF3332322C074577A4476AFC1FA1FD" blockId="44.[151,1128,1341,1442]" pageId="44" pageNumber="41">
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<emphasis id="B934EF20322C074577A4476AFEB2A130" bold="true" box="[151,279,1368,1388]" pageId="44" pageNumber="41">Type material.</emphasis>
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<materialsCitation id="3B28396F322C07457629476BFBCDA130" ID-GBIF-Occurrence="2984087455" box="[282,1128,1366,1389]" collectingDate="1990-09-04" collectionCode="MNHAH" collectorName="N. Yasuda" country="Japan" location="Mts. Daisetsu, Mt. Kurodake" pageId="44" pageNumber="41" specimenCount="1" specimenCount-female="1" stateProvince="HOKKAIDO" typeStatus="holotype">
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||
<emphasis id="B934EF20322C07457629476BFE62A130" bold="true" box="[282,455,1368,1388]" pageId="44" pageNumber="41">
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||
<collectingCountry id="F35773A2322C07457629476BFEC4A130" box="[282,353,1368,1388]" name="Japan" pageId="44" pageNumber="41">JAPAN</collectingCountry>
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||
:
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<collectingRegion id="4984FDD0322C0745765B476AFE65A137" box="[360,448,1369,1388]" country="Japan" name="Hokkaido" pageId="44" pageNumber="41">HOKKAIDO</collectingRegion>
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||
:
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</emphasis>
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||
<typeStatus id="54FB8D90322C074576F8476AFDB9A137" box="[459,540,1369,1388]" pageId="44" pageNumber="41" type="holotype">HOLOTYPE</typeStatus>
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||
,
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<specimenCount id="9D46F8BB322C074575164765FD90A131" box="[549,565,1366,1389]" pageId="44" pageNumber="41" type="female">♀</specimenCount>
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,
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<location id="8E9F65E9322C0745750D476AFC88A130" LSID="urn:lsid:plazi:treatment:03E98224322C075976DF46E8FD33A248:8E9F65E9322C0745750D476AFC88A130" box="[574,813,1369,1388]" country="Japan" name="Mts. Daisetsu, Mt. Kurodake" pageId="44" pageNumber="41" stateProvince="HOKKAIDO">Mts.Daisetsu,Mt. Kurodake</location>
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,
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<date id="FFFE15F2322C07457400476AFC21A130" box="[819,900,1369,1388]" pageId="44" pageNumber="41" value="1990-09-04">
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<collectingDate id="EFBAEC1A322C07457400476AFC21A130" box="[819,900,1369,1388]" pageId="44" pageNumber="41" value="1990-09-04">4.ix.1990</collectingDate>
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</date>
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,
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<collectorName id="26B556E4322C074574B8476AFC7AA130" box="[907,991,1369,1388]" pageId="44" pageNumber="41">N.Yasuda</collectorName>
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leg.(
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<collectionCode id="ED51ABF7322C07457339476AFBFBA130" box="[1034,1118,1369,1388]" collectionName="MNHAH" pageId="44" pageNumber="41">MNHAH</collectionCode>
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).
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</materialsCitation>
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<materialsCitation id="3B28396F322C074577A44747FC1FA1FD" ID-GBIF-Occurrence="2984087543" collectingDate="1999-08-07" collectingDateMax="1999-08-15" collectingDateMin="1999-08-07" collectionCode="MNHAH" collectorName="S. Hori" location="Opposite" municipality="Mts. Daisetsu" pageId="44" pageNumber="41" specimenCount="5" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
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||
<typeStatus id="54FB8D90322C074577A44747FF43A1D9" box="[151,230,1396,1415]" pageId="44" pageNumber="41" type="paratype">PARATYPE</typeStatus>
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||
:
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<specimenCount id="9D46F8BB322C074577C34747FEABA1D4" box="[240,270,1392,1416]" pageId="44" pageNumber="41" type="male">1 ♂</specimenCount>
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||
,
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||
<collectingMunicipality id="6B9BA948322C0745762B4747FE2AA1DB" box="[280,399,1396,1415]" pageId="44" pageNumber="41">Mts. Daisetsu</collectingMunicipality>
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, Mikura-zawa,
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<date id="FFFE15F2322C074575224740FD30A1DA" box="[529,661,1395,1414]" pageId="44" pageNumber="41" value="1999-08-07" valueMax="1999-08-15" valueMin="1999-08-07">
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||
<collectingDate id="EFBAEC1A322C074575224740FD30A1DA" box="[529,661,1395,1414]" pageId="44" pageNumber="41" value="1999-08-07" valueMax="1999-08-15" valueMin="1999-08-07">7.–15.viii.1999</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26B556E4322C074575AF4747FD7DA1DB" box="[668,728,1396,1415]" pageId="44" pageNumber="41">S. Hori</collectorName>
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||
leg. (
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||
<collectionCode id="ED51ABF7322C074574354747FCF9A1DB" box="[774,860,1396,1415]" collectionName="MNHAH" pageId="44" pageNumber="41">MNHAH</collectionCode>
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||
).
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<location id="8E9F65E9322C0745745B4747FC10A1DB" LSID="urn:lsid:plazi:treatment:03E98224322C075976DF46E8FD33A248:8E9F65E9322C0745745B4747FC10A1DB" box="[872,949,1396,1415]" municipality="Mts. Daisetsu" name="Opposite" pageId="44" pageNumber="41">Opposite</location>
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||
to the usual practice, I have designated the
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||
<specimenCount id="9D46F8BB322C0745766247BDFE2EA1FD" box="[337,395,1422,1441]" pageId="44" pageNumber="41" type="female">female</specimenCount>
|
||
<specimenCount id="9D46F8BB322C074576A247BDFE47A1FD" box="[401,482,1422,1441]" pageId="44" pageNumber="41" type="generic">specimen</specimenCount>
|
||
as the holotype, as the above
|
||
<specimenCount id="9D46F8BB322C074575D247BDFCAFA1FD" box="[737,778,1422,1441]" pageId="44" pageNumber="41" type="male">male</specimenCount>
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||
<specimenCount id="9D46F8BB322C0745742347BDFCC4A1FD" box="[784,865,1422,1441]" pageId="44" pageNumber="41" type="generic">specimen</specimenCount>
|
||
is teneral.
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||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B9322C074477A4478AFD10A54F" lastPageId="45" lastPageNumber="38" pageId="44" pageNumber="41" type="diagnosis">
|
||
<paragraph id="8BFF3332322C074477A4478AFD10A54F" blockId="44.[151,1127,1465,1586]" lastBlockId="45.[113,1090,219,436]" lastPageId="45" lastPageNumber="38" pageId="44" pageNumber="41">
|
||
<emphasis id="B934EF20322C074577A4478AFEA9A18D" bold="true" box="[151,268,1465,1489]" pageId="44" pageNumber="41">Diagnosis.</emphasis>
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||
Body 2.8–3.0 mm long and ca. 1.8× as long as wide. Dorsum brownish. Elytra densely and strongly punctate between striae, and not transversely strigose between punctures. Mesoventrite without distinct excavation between median carina and transverse carina. Median carina of mesoventrite low. Male metaventrite bearing erect and dense pubescence medially. Male mesotibia protuberant at interoapical corner. Male metatibiae strongly arcuate. Female abdominal sternite 8 with a spiculum ventrale.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35A60B9322D075977424328FD67A148" lastPageId="48" lastPageNumber="45" pageId="45" pageNumber="38" type="description">
|
||
<paragraph id="8BFF3332322D074477424328FEF3A52F" blockId="45.[113,1090,219,436]" pageId="45" pageNumber="38">
|
||
<emphasis id="B934EF20322D074477424328FF59A56F" bold="true" box="[113,252,283,307]" pageId="45" pageNumber="38">Description.</emphasis>
|
||
Measurements of
|
||
<typeStatus id="54FB8D90322D074476F14328FD81A56F" box="[450,548,283,307]" pageId="45" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
: Body length
|
||
<quantity id="4CB89ED7322D07447584432FFCA9A56F" box="[695,780,284,308]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="45" pageNumber="38" unit="mm" value="2.8">2.8 mm</quantity>
|
||
; head
|
||
<quantity id="4CB89ED7322D074474634328FC0BA56F" box="[848,942,283,307]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="45" pageNumber="38" unit="mm" value="0.45">0.45 mm</quantity>
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||
in length and
|
||
<quantity id="4CB89ED7322D07447742430FFF6BA50F" box="[113,206,316,340]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="45" pageNumber="38" unit="mm" value="0.77">0.77 mm</quantity>
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||
in width; pronotum
|
||
<quantity id="4CB89ED7322D07447697430FFDA4A50F" box="[420,513,316,340]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="45" pageNumber="38" unit="mm" value="0.74">0.74 mm</quantity>
|
||
in length and
|
||
<quantity id="4CB89ED7322D074475A7430FFD41A50F" box="[660,740,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="45" pageNumber="38" unit="mm" value="1.3">1.3 mm</quantity>
|
||
in width; elytra
|
||
<quantity id="4CB89ED7322D074474BC430FFC7BA50F" box="[911,990,316,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="45" pageNumber="38" unit="mm" value="1.9">1.9 mm</quantity>
|
||
in length and
|
||
<quantity id="4CB89ED7322D074477AC4368FF55A52F" box="[159,240,347,371]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="45" pageNumber="38" unit="mm" value="1.6">1.6 mm</quantity>
|
||
in width.
|
||
</paragraph>
|
||
<paragraph id="8BFF3332322D074677A34348FC0BA1C8" blockId="45.[113,1090,219,436]" lastBlockId="47.[113,1089,1340,1556]" lastPageId="47" lastPageNumber="40" pageId="45" pageNumber="38">
|
||
Coloration (based on
|
||
<typeStatus id="54FB8D90322D074476454348FE76A5CF" box="[374,467,379,403]" pageId="45" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
only). Dorsum shining and brownish; elytra slightly paler than head and pronotum; antennomeres 1–6 and 8 brown; antennomere 7 a little darker than 6; antennomeres 9, 10, and basal 2/3 of antennomere 11 dark brown; apical 1/3 of antennomere 11 light brown; coxae, mesotrochanters, and metatrochanters brown; remaining parts of legs yellowish brown; mesoventrite, metaventrite, and abdominal ventrites brown.
|
||
</paragraph>
|
||
<caption id="DF3F63BA322D0744774347DFFD98A269" ID-DOI="http://doi.org/10.5281/zenodo.4272511" ID-Zenodo-Dep="4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="45" pageNumber="38" startId="45.[112,143,1516,1535]" targetBox="[152,1047,472,1486]" targetPageId="45">
|
||
<paragraph id="8BFF3332322D0744774347DFFD98A269" blockId="45.[112,1089,1516,1589]" pageId="45" pageNumber="38">
|
||
Fig. 21.
|
||
<taxonomicName id="4C4048B1322D0744778B47DEFEE7A1A3" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[184,322,1517,1535]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="38" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
|
||
<emphasis id="B934EF20322D0744778B47DEFEE7A1A3" box="[184,322,1517,1535]" italics="true" pageId="45" pageNumber="38">Leiodes yasudai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B322D0744767B47DEFE2FA25C" box="[328,394,1517,1536]" pageId="45" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3F63BA322E074777A4478BFEBEA26A" ID-DOI="http://doi.org/10.5281/zenodo.4272513" ID-Zenodo-Dep="4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="46" pageNumber="43" startId="46.[151,182,1464,1483]" targetBox="[177,1094,236,1406]" targetPageId="46">
|
||
<paragraph id="8BFF3332322E074777A4478BFEBEA26A" blockId="46.[151,1125,1464,1590]" pageId="46" pageNumber="43">
|
||
Fig. 22.
|
||
<taxonomicName id="4C4048B1322E074777D1478AFECBA197" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[226,366,1465,1483]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="43" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
|
||
<emphasis id="B934EF20322E074777D1478AFECBA197" box="[226,366,1465,1483]" italics="true" pageId="46" pageNumber="43">Leiodes yasudai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B322E07477646478BFE1FA197" box="[373,442,1464,1483]" pageId="46" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3F63BA322F074677424694FF4FA0AC" ID-DOI="http://doi.org/10.5281/zenodo.4272515" ID-Zenodo-Dep="4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="47" pageNumber="40" startId="47.[113,144,1191,1210]" targetBox="[136,1062,228,1156]" targetPageId="47">
|
||
<paragraph id="8BFF3332322F074677424694FF4FA0AC" blockId="47.[113,1089,1191,1264]" pageId="47" pageNumber="40">
|
||
Fig. 23.
|
||
<taxonomicName id="4C4048B1322F0746778B469BFEE7A0E6" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[184,322,1192,1210]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="40" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
|
||
<emphasis id="B934EF20322F0746778B469BFEE7A0E6" box="[184,322,1192,1210]" italics="true" pageId="47" pageNumber="40">Leiodes yasudai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B322F074676744694FE2FA0E6" box="[327,394,1191,1210]" pageId="47" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF3332322F074677A347AFFD6CA1E8" blockId="47.[113,1089,1340,1556]" box="[144,713,1436,1460]" pageId="47" pageNumber="40">Body 2.8–3.0 mm in length, ca. 1.8× as long as wide.</paragraph>
|
||
<paragraph id="8BFF3332322F075977A3478FFD9FA54F" blockId="47.[113,1089,1340,1556]" lastBlockId="48.[151,1127,219,1556]" lastPageId="48" lastPageNumber="45" pageId="47" pageNumber="40">
|
||
Head ca. 1.7× as wide as long, ca. 0.61× as long as and 0.56× as wide as pronotum, distinctly punctate, bearing some large punctures (
|
||
<figureCitation id="137B2FB7322F0746754A47EFFD7AA1A8" box="[633,735,1500,1524]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="47" pageNumber="40">Fig. 21A</figureCitation>
|
||
); antennomeres 1–4 each longer than wide; antennomere 11 about as long as wide; remaining antennomeres each wider than long; antennomere 11 oval (
|
||
<figureCitation id="137B2FB73230075976F042E8FD80A4A8" box="[451,549,219,244]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21C</figureCitation>
|
||
); relative lengths of antennomeres 2 to 11 – 2.8: 4.3: 1.9: 1.8: 1.6: 2.8: 1.0: 3.4: 3.1: 4.5.
|
||
</paragraph>
|
||
<paragraph id="8BFF3332323007597786432FFDA1A528" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
Pronotum ca. 1.8× as wide as long, ca. 0.40× as long as and 0.87× as wide as elytra, widest at base, simply and very feebly curved at posterior margin, distinctly punctate, punctation similar to that on head (
|
||
<figureCitation id="137B2FB73230075976A74368FE5DA528" box="[404,504,347,372]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF33323230075977864348FE55A5CF" blockId="48.[151,1127,219,1556]" box="[181,496,379,403]" pageId="48" pageNumber="45">Scutellum minutely punctate.</paragraph>
|
||
<paragraph id="8BFF333232300759778643A8FE41A66F" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
Elytra ca. 1.2× as long as wide in dorsal view, widest ca. at basal 1/3 (
|
||
<figureCitation id="137B2FB73230075974FA43A8FB8AA5E8" box="[969,1071,411,436]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21A</figureCitation>
|
||
), not transversely strigose, densely and strongly punctate between striae, therefore superficially not appearing to bear rows of punctures (
|
||
<figureCitation id="137B2FB732300759755343E8FCE3A5A8" box="[608,838,475,500]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Figs. 21A, 21B, 21F</figureCitation>
|
||
); most punctures of elytra larger than those of head and pronotum (
|
||
<figureCitation id="137B2FB732300759757943C8FD0BA648" box="[586,686,507,532]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21A</figureCitation>
|
||
); sutural stria fine, reaching from apex to ca. apical half of elytral length.
|
||
</paragraph>
|
||
<paragraph id="8BFF33323230075977864008FD98A60F" blockId="48.[151,1127,219,1556]" box="[181,573,571,595]" pageId="48" pageNumber="45">Metathoracic wings fully developed.</paragraph>
|
||
<paragraph id="8BFF33323230075977864068FC58A68F" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
Mesoventrite strongly microreticulate, impunctate, almost glabrous, without distinct excavation between median carina and transverse carina (
|
||
<figureCitation id="137B2FB732300759758F4048FCBFA6C8" box="[700,794,635,660]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21E</figureCitation>
|
||
); median carina of mesoventrite low (
|
||
<figureCitation id="137B2FB73230075977FF40A8FE8FA6E8" box="[204,298,667,692]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21E</figureCitation>
|
||
); metaventrite showing sexual dimorphism (
|
||
<figureCitation id="137B2FB73230075975C540A8FC37A6E8" box="[758,914,667,692]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Figs. 21F, 21G</figureCitation>
|
||
), sparsely and finely pubescent, and strongly microreticulate except for almost smooth middle portion.
|
||
</paragraph>
|
||
<paragraph id="8BFF333232300759778640EFFEE8A768" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
Legs showing distinct sexual dimorphism of protarsi, protibia, mesotibia, mesotarsi, metafemur, and metatibia; metafemora weakly expanded at about apical 1/4 of posterior margins (
|
||
<figureCitation id="137B2FB73230075977AC4128FEE4A768" box="[159,321,795,820]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Figs. 22E, 22F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFF33323230075977864109FBB0A008" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977864109FF50A708" bold="true" box="[181,245,826,852]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977864109FF48A70F" bold="true" box="[181,237,826,851]" italics="true" pageId="48" pageNumber="45">Male</emphasis>
|
||
.
|
||
</emphasis>
|
||
Middle portion of metaventrite with dense erect pubescence (
|
||
<figureCitation id="137B2FB73230075974A04108FC56A708" box="[915,1011,827,852]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21F</figureCitation>
|
||
); protibiae strongly curved inwards along dorsoapical margins in dorsal view (
|
||
<figureCitation id="137B2FB73230075974604168FC13A728" box="[851,950,859,884]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22G</figureCitation>
|
||
); tarsomeres 2–4 of protarsi and mesotarsi expanded (
|
||
<figureCitation id="137B2FB73230075975224148FD10A7C8" box="[529,693,891,916]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Figs. 22A, 22C</figureCitation>
|
||
); mesotibia strongly protuberant and bearing a robust spine at interoapical corner (
|
||
<figureCitation id="137B2FB732300759757E41A8FD0BA7E8" box="[589,686,923,948]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22C</figureCitation>
|
||
); metafemur with a long dorsal projection posteroapically, projection curved inwards (
|
||
<figureCitation id="137B2FB73230075975434188FD6CA788" box="[624,713,955,980]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22I</figureCitation>
|
||
); metatibiae strongly arcuately curved (
|
||
<figureCitation id="137B2FB73230075977AC41E8FEA7A7A8" box="[159,258,987,1012]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22E</figureCitation>
|
||
); abdominal sternite 8 strongly curved (
|
||
<figureCitation id="137B2FB732300759758741E8FCB2A7A8" box="[692,791,987,1012]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23C</figureCitation>
|
||
); aedeagus slender (
|
||
<figureCitation id="137B2FB73230075974C741E8FF62A048" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Figs. 23A, 23B</figureCitation>
|
||
); median lobe simply narrowed from apical 1/3 towards apex, rounded at apex in dorsal view (
|
||
<figureCitation id="137B2FB73230075977EB4628FE9CA068" box="[216,313,1051,1076]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23A</figureCitation>
|
||
), feebly curved and apically pointed in lateral view (
|
||
<figureCitation id="137B2FB732300759746F4628FC19A068" box="[860,956,1051,1076]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23B</figureCitation>
|
||
); each paramere bearing a small transparent lobe at apex (
|
||
<figureCitation id="137B2FB732300759757E4608FD14A008" box="[589,689,1083,1108]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23A</figureCitation>
|
||
); inner sac as shown in
|
||
<figureCitation id="137B2FB732300759749C4608FBB4A008" box="[943,1041,1083,1108]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23A</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFF33323230075977864669FD67A148" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977864669FEAAA028" bold="true" box="[181,271,1114,1140]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977864669FEADA02F" bold="true" box="[181,264,1114,1139]" italics="true" pageId="48" pageNumber="45">Female</emphasis>
|
||
.
|
||
</emphasis>
|
||
Middle portion of metaventrite with sparse erect pubescence (
|
||
<figureCitation id="137B2FB73230075974AC4668FBA7A028" box="[927,1026,1115,1140]" captionStart="Fig" captionStartId="45.[112,143,1516,1535]" captionTargetBox="[152,1047,472,1486]" captionTargetId="figure-100@45.[147,1056,465,1487]" captionTargetPageId="45" captionText="Fig. 21. Leiodes yasudai sp. nov. A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 0.5 mm for F and G." figureDoi="http://doi.org/10.5281/zenodo.4272511" httpUri="https://zenodo.org/record/4272511/files/figure.png" pageId="48" pageNumber="45">Fig. 21G</figureCitation>
|
||
); protibia not curved inwards on dorsoapical margin in dorsal view (
|
||
<figureCitation id="137B2FB73230075974374648FCCDA0C8" box="[772,872,1147,1172]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22H</figureCitation>
|
||
); protarsi and mesotarsi slender (
|
||
<figureCitation id="137B2FB73230075977C646A8FE33A0E8" box="[245,406,1179,1204]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22B, 22D</figureCitation>
|
||
); mesotibia with a slender spine at interoapical corner (
|
||
<figureCitation id="137B2FB73230075974C646A8FBFEA0E8" box="[1013,1115,1179,1204]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22D</figureCitation>
|
||
); metafemur with a small dorsal projection posteroapically (
|
||
<figureCitation id="137B2FB73230075975C14688FCECA088" box="[754,841,1211,1236]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22J</figureCitation>
|
||
); metatibiae almost straight (
|
||
<figureCitation id="137B2FB73230075977AC46E8FF5BA0A8" box="[159,254,1243,1268]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22F</figureCitation>
|
||
); abdominal sternite 8 with a spiculum ventrale at central point of anterior margin (
|
||
<figureCitation id="137B2FB73230075977AC46C8FEA6A148" box="[159,259,1275,1300]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23D</figureCitation>
|
||
); coxites and stylus as shown in
|
||
<figureCitation id="137B2FB732300759756C46C8FD1BA148" box="[607,702,1275,1300]" captionStart="Fig" captionStartId="47.[113,144,1191,1210]" captionTargetBox="[136,1062,228,1156]" captionTargetId="figure-106@47.[121,1080,219,1171]" captionTargetPageId="47" captionText="Fig. 23. Leiodes yasudai sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E." figureDoi="http://doi.org/10.5281/zenodo.4272515" httpUri="https://zenodo.org/record/4272515/files/figure.png" pageId="48" pageNumber="45">Fig. 23E</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93230075977A44728FEACA1E8" pageId="48" pageNumber="45" type="diagnosis">
|
||
<paragraph id="8BFF33323230075977A44728FEACA1E8" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977A44728FE29A16F" bold="true" box="[151,396,1307,1331]" pageId="48" pageNumber="45">Differential diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C4048B13230075976AF472EFDEFA168" authority="Hoshina, 2012" authorityName="Hoshina" authorityYear="2012" box="[412,586,1309,1332]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="45" phylum="Arthropoda" rank="species" species="yasudai" status="sp. nov.">
|
||
<emphasis id="B934EF203230075976AF472EFDEFA168" box="[412,586,1309,1332]" italics="true" pageId="48" pageNumber="45">Leiodes yasudai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207525B323007597560472FFD0DA16F" box="[595,680,1308,1331]" pageId="48" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C4048B1323007597409472EFBECA168" authority="(Rye, 1873)" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[826,1097,1308,1332]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="45" phylum="Arthropoda" rank="species" species="circinipes">
|
||
<emphasis id="B934EF20323007597409472EFC65A168" box="[826,960,1309,1332]" italics="true" pageId="48" pageNumber="45">L. circinipes</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD14EC33230075974E3472FFBE7A168" author="RYE E. C." box="[976,1090,1308,1332]" pageId="48" pageNumber="45" pagination="131 - 135" refId="ref74051" refString="RYE E. C. 1873: Notes on Anisotomidae, with description of three new species (two from Japan, and one from Gt. Britain [sic!]). No. 1. Entomologist's Monthly Magazine 10: 131 - 135." type="journal article" year="1873">Rye, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in elytral punctation, but can be separated from it by having almost unicolor elytra and strongly curved male metatibiae (
|
||
<figureCitation id="137B2FB73230075976934768FDA4A128" box="[416,513,1371,1396]" captionStart="Fig" captionStartId="46.[151,182,1464,1483]" captionTargetBox="[177,1094,236,1406]" captionTargetId="figure-11@46.[164,1113,219,1418]" captionTargetPageId="46" captionText="Fig. 22. Leiodes yasudai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – female hind leg, ventral view; G –male protibia, dorsal view; H – female protibia, dorsal view; I – male metafemur, dorsal view; J – female metafemur, dorsal view. Scale I: 1 mm for A–D; II: 1 mm for E and F; III: 0.2 mm for G and H; IV: 0.5 mm for I and J." figureDoi="http://doi.org/10.5281/zenodo.4272513" httpUri="https://zenodo.org/record/4272513/files/figure.png" pageId="48" pageNumber="45">Fig. 22E</figureCitation>
|
||
). In contrast,
|
||
<taxonomicName id="4C4048B13230075975A1476EFCB2A128" baseAuthorityName="Rye" baseAuthorityYear="1873" box="[658,791,1373,1396]" class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="45" phylum="Arthropoda" rank="species" species="circinipes">
|
||
<emphasis id="B934EF203230075975A1476EFCB2A128" box="[658,791,1373,1396]" italics="true" pageId="48" pageNumber="45">L. circinipes</emphasis>
|
||
</taxonomicName>
|
||
has brownish elytra with black stripes near the elytral suture (
|
||
<figureCitation id="137B2FB73230075976E54748FD90A1CF" box="[470,565,1403,1427]" captionStart="Fig" captionStartId="38.[151,182,1544,1563]" captionTargetBox="[251,1026,769,1525]" captionTargetId="figure-252@38.[244,1031,760,1529]" captionTargetPageId="38" captionText="Fig. 16. Leiodes circinipes (Rye, 1873).A – body, dorsal view; B – ditto, lateral view; C – dorsal color; D – antenna; E – elytral punctures; F – mesoventrite, lateral view. Scale I: 1 mm for A and B; II: 0.5 mm for D; III: 0.5 mm for F." figureDoi="http://doi.org/10.5281/zenodo.4272501" httpUri="https://zenodo.org/record/4272501/files/figure.png" pageId="48" pageNumber="45">Fig. 16C</figureCitation>
|
||
) and relatively weakly curved male metatibiae (
|
||
<figureCitation id="137B2FB73230075973024748FF59A1E8" captionStart="Fig" captionStartId="39.[113,144,1452,1471]" captionTargetBox="[119,1082,224,1375]" captionTargetId="figure-15@39.[113,1088,219,1384]" captionTargetPageId="39" captionText="Fig. 17. Leiodes circinipes (Rye, 1873).A – male fore leg, ventral view; B – female fore leg, ventral view; C – male middle leg, ventral view; D – female middle leg, ventral view; E – male hind leg, ventral view; F – ditto; G – female hind leg, ventral view; H – male metafemur, dorsal view; I – female metafemur, dorsal view; J – male protibiae, dorsal view; K – female protibiae, dorsal view. Scale I: 0.5 mm for A–D; II: 0.5 mm for E; III: 0.5 mm for F and G; IV: 0.2 mm for H and I; V: 0.2 mm for J and K." figureDoi="http://doi.org/10.5281/zenodo.4272503" httpUri="https://zenodo.org/record/4272503/files/figure.png" pageId="48" pageNumber="45">Figs. 17E, 17F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93230075977A4478FFF1DA1AF" pageId="48" pageNumber="45" type="etymology">
|
||
<paragraph id="8BFF33323230075977A4478FFF1DA1AF" blockId="48.[151,1127,219,1556]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977A4478FFEBDA188" bold="true" box="[151,280,1468,1492]" pageId="48" pageNumber="45">Etymology.</emphasis>
|
||
This species is dedicated to the collector of the
|
||
<typeStatus id="54FB8D903230075974154788FC23A18F" box="[806,902,1467,1491]" pageId="48" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
, Dr. Nobunori Yasuda.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A60B93230075977A447C8FD33A248" box="[151,662,1531,1556]" pageId="48" pageNumber="45" type="distribution">
|
||
<paragraph id="8BFF33323230075977A447C8FD33A248" blockId="48.[151,1127,219,1556]" box="[151,662,1531,1556]" pageId="48" pageNumber="45">
|
||
<emphasis id="B934EF203230075977A447C8FE8FA24F" bold="true" box="[151,298,1531,1555]" pageId="48" pageNumber="45">Distribution.</emphasis>
|
||
<collectingCountry id="F35773A232300759760347CFFED4A24F" box="[304,369,1532,1555]" name="Japan" pageId="48" pageNumber="45">Japan</collectingCountry>
|
||
:
|
||
<collectingRegion id="4984FDD032300759764847C8FE41A24F" box="[379,484,1531,1555]" country="Japan" name="Hokkaido" pageId="48" pageNumber="45">Hokkaido</collectingRegion>
|
||
(Daisetsu Mts.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |