treatments-xml/data/03/E9/20/03E92035F96DFFA9FF5559065E7CA097.xml

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<mods:title id="A827E76147086307F155B194D8EBC8E2">Tarsal tetramery and extreme size reduction in Anillini (Coleoptera, Carabidae, Trechinae): the case of Typhlocharis Dieck, 1869; description of three new species and definition of a new intra-generic species group</mods:title>
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<mods:namePart id="3E84F1A940F3DFD906311A5B04443885">Pérez-González, Sergio</mods:namePart>
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<heading id="D0B7264FF96DFFADFF55590659F2A4AB" bold="true" box="[151,930,1398,1424]" fontSize="11" level="1" pageId="2" pageNumber="251" reason="1">
<emphasis id="B9344D31F96DFFADFF55590659F2A4AB" bold="true" box="[151,930,1398,1424]" pageId="2" pageNumber="251">
<taxonomicName id="4C40EAA0F96DFFADFF555906595BA4AB" ID-CoL="59VJ9" authority="Perez-Gonzalez &amp; Zaballos" authorityName="Perez-Gonzalez &amp; Zaballos" box="[151,779,1398,1424]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F96DFFADFF5559065BFEA4AB" bold="true" box="[151,430,1398,1424]" italics="true" pageId="2" pageNumber="251">Typhlocharis baeturica</emphasis>
Pérez-González &amp; Zaballos
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(
<figureCitation id="137B8DA6F96DFFADFF5D59E95B51A48A" box="[159,257,1433,1457]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1914,1937]" captionStartId-1="5.[151,250,952,975]" captionTargetBox-0="[177,1392,211,1861]" captionTargetBox-1="[207,1365,203,903]" captionTargetId-0="figure@4.[155,1432,195,1890]" captionTargetId-1="figure@5.[183,1393,177,927]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Typhlocharis baeturica sp. n. – habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." captionText-1="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri-0="https://zenodo.org/record/222600/files/figure.png" httpUri-1="https://zenodo.org/record/222601/files/figure.png" pageId="2" pageNumber="251">Figs 1–2</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BFF9123F96DFFADFF5559925BB9A7B2" blockId="2.[151,1437,1505,2033]" pageId="2" pageNumber="251">
<emphasis id="B9344D31F96DFFADFF5559925B72A4C1" bold="true" box="[151,290,1505,1530]" pageId="2" pageNumber="251">
<typeStatus id="54FB2F81F96DFFADFF5559925A82A4C1" box="[151,210,1506,1530]" pageId="2" pageNumber="251">Type</typeStatus>
series.
</emphasis>
<typeStatus id="54FB2F81F96DFFADFEEA59915BC3A4C2" box="[296,403,1505,1529]" pageId="2" pageNumber="251" type="holotype">Holotype</typeStatus>
, 3, 
<collectingCountry id="F357D1B3F96DFFADFE0259915843A4C2" box="[448,531,1505,1529]" name="Spain" pageId="2" pageNumber="251">SPAIN</collectingCountry>
: Huelva, Sanlúcar de Guadiana (
<quantity id="4CB83CC6F96DFFADFCBF5991599BA4C2" box="[893,971,1505,1529]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="251" unit="km" value="1.7">1.7 km</quantity>
SE), 
<date id="FFFEB7E3F96DFFADFBCB59915EDCA4C2" box="[1033,1164,1505,1529]" pageId="2" pageNumber="251" value="2011-02-11">11-02-2011</date>
, 61 m (
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, 
<geoCoordinate id="EE74F7E4F96DFFADFA9459925AEEA726" direction="west" orientation="longitude" pageId="2" pageNumber="251" precision="925" value="-7.45">W 07º 27’</geoCoordinate>
), J.P. Zaballos &amp; S. Pérez leg. (Coll. J.P. Zaballos, 
<collectionCode id="ED5109E6F96DFFADFCC45A75591BA726" LSID="urn:lsid:biocol.org:col:34854" box="[774,843,1541,1565]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="2" pageNumber="251">UCM</collectionCode>
). 
<typeStatus id="54FB2F81F96DFFADFC9D5A765980A725" box="[863,976,1542,1566]" pageId="2" pageNumber="251" type="paratype">Paratypes</typeStatus>
: 1 Ƥ same data as 
<typeStatus id="54FB2F81F96DFFADFB6A5A755F5EA726" box="[1192,1294,1541,1565]" pageId="2" pageNumber="251" type="holotype">holotype</typeStatus>
; 2 3 same locality as 
<typeStatus id="54FB2F81F96DFFADFECD5A595B24A77A" box="[271,372,1577,1601]" pageId="2" pageNumber="251" type="holotype">holotype</typeStatus>
, 
<date id="FFFEB7E3F96DFFADFEBD5A595853A77A" box="[383,515,1577,1601]" pageId="2" pageNumber="251" value="2012-11-21">21-11-2012</date>
, J.P. Zaballos, S. Pérez &amp; S. Ghannem leg.; 2 3, 5 ƤƤ 
<collectingCountry id="F357D1B3F96DFFADFBBD5A595E82A77A" box="[1151,1234,1577,1601]" name="Spain" pageId="2" pageNumber="251">SPAIN</collectingCountry>
: Huelva, Sanlúcar de Guadiana (
<quantity id="4CB83CC6F96DFFADFEF15A3D5B3DA75E" box="[307,365,1613,1637]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="251" unit="km" value="4.0">4 km</quantity>
SE), 
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21- 
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</date>
, 129 m (
<geoCoordinate id="EE74F7E4F96DFFADFD505A3D5952A75E" box="[658,770,1613,1637]" direction="north" orientation="latitude" pageId="2" pageNumber="251" precision="925" value="37.433334">N 37º 26’</geoCoordinate>
, 
<geoCoordinate id="EE74F7E4F96DFFADFCCD5A3E59D3A75E" box="[783,899,1613,1637]" direction="west" orientation="longitude" pageId="2" pageNumber="251" precision="925" value="-7.4333334">W 07º 26’</geoCoordinate>
), J.P. Zaballos, S. Pérez &amp; S. Ghannem leg. (10 ex. Coll. J.P. Zaballos, 
<collectionCode id="ED5109E6F96DFFADFE5A5A015B8DA7B2" LSID="urn:lsid:biocol.org:col:34854" box="[408,477,1649,1673]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="2" pageNumber="251">UCM</collectionCode>
).
</paragraph>
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<emphasis id="B9344D31F96DFFADFF055AE55B11A795" bold="true" box="[199,321,1685,1710]" pageId="2" pageNumber="251">Diagnosis.</emphasis>
Small endogean beetle, anophthalmous, with narrow and subrectangular body covered by microreticulate integument and scattered pubescence. Vertex with 
<emphasis id="B9344D31F96DFFADFC4D5ACA5E73A7EA" box="[911,1059,1721,1745]" italics="true" pageId="2" pageNumber="251">pars stridens</emphasis>
. Gula partially fused to cephalic capsule. Serration in lateral margins of elytra well developed, hooked in the first third. Apical margin with two pairs of denticles (a sutural pair and a pair associated to 7th stria), pointy and well defined. Umbilicate series with seven setae (4+3). Sternum II with a pair of foveae in females. Tarsal formula: 4-4-4. Aedeagus as in 
<figureCitation id="137B8DA6F96DFFADFADE5B555F0FA606" box="[1308,1375,1829,1853]" captionStart="FIGURE 2" captionStartId="5.[151,250,952,975]" captionTargetBox="[207,1365,203,903]" captionTargetId="figure@5.[183,1393,177,927]" captionTargetPageId="5" captionText="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="2" pageNumber="251">Fig. 2</figureCitation>
A, B, with two-pieced, &quot;scissor-shaped&quot;, endophallic sclerites. Female genitalia with tubular gonocoxites, as in 
<figureCitation id="137B8DA6F96DFFADFAF75B395F28A65A" box="[1333,1400,1865,1889]" captionStart="FIGURE 2" captionStartId="5.[151,250,952,975]" captionTargetBox="[207,1365,203,903]" captionTargetId="figure@5.[183,1393,177,927]" captionTargetPageId="5" captionText="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="2" pageNumber="251">Fig. 2</figureCitation>
C.
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F96DFFAAFF055B1D5BACA431" lastPageId="5" lastPageNumber="254" pageId="2" pageNumber="251" type="description">
<paragraph id="8BFF9123F96DFFADFF055B1D5B32A692" blockId="2.[151,1437,1505,2033]" pageId="2" pageNumber="251">
<emphasis id="B9344D31F96DFFADFF055B1D5B09A6BD" bold="true" box="[199,345,1901,1926]" pageId="2" pageNumber="251">Description.</emphasis>
Length of 
<typeStatus id="54FB2F81F96DFFADFE245B1D581CA6BE" box="[486,588,1901,1925]" pageId="2" pageNumber="251" type="holotype">holotype</typeStatus>
: 
<quantity id="4CB83CC6F96DFFADFD9F5B1D589AA6BE" box="[605,714,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="251" unit="mm" value="1.14">1.14 mm</quantity>
; length of 
<typeStatus id="54FB2F81F96DFFADFC965B1E5994A6BD" box="[852,964,1902,1926]" pageId="2" pageNumber="251" type="paratype">paratypes</typeStatus>
: 1.00– 
<quantity id="4CB83CC6F96DFFADFBD65B1D5E2DA6BE" box="[1044,1149,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="251" unit="mm" value="1.14">1.14 mm</quantity>
(males), 
<quantity id="4CB83CC6F96DFFADFB315B1D5FCCA6BE" box="[1267,1436,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.12" metricValueMin="1.06" pageId="2" pageNumber="251" unit="mm" value="1.09" valueMax="1.12" valueMin="1.06">1.06–1.12 mm</quantity>
(females) (
<figureCitation id="137B8DA6F96DFFADFED25BE15B01A692" box="[272,337,1937,1961]" captionStart="FIGURE 1" captionStartId="4.[151,250,1914,1937]" captionTargetBox="[177,1392,211,1861]" captionTargetId="figure@4.[155,1432,195,1890]" captionTargetPageId="4" captionText="FIGURE 1. Typhlocharis baeturica sp. n. – habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222600/files/figure.png" pageId="2" pageNumber="251">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BFF9123F96DFFACFF055BC55B7DA39B" blockId="2.[151,1437,1505,2033]" lastBlockId="3.[151,1437,151,2020]" lastPageId="3" lastPageNumber="252" pageId="2" pageNumber="251">
Head: wider (
<quantity id="4CB83CC6F96DFFADFEB05BC55872A6F6" box="[370,546,1973,1997]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="2" pageNumber="251" unit="mm" value="0.245" valueMax="0.26" valueMin="0.23">0.23–0.26 mm</quantity>
) than long (
<quantity id="4CB83CC6F96DFFADFD075BC55927A6F6" box="[709,887,1973,1997]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="2" pageNumber="251" unit="mm" value="0.205" valueMax="0.21" valueMin="0.2">0.20–0.21 mm</quantity>
), covered by subhexagonal microreticulation. Stridulatory organ (
<emphasis id="B9344D31F96DFFADFEB85BAA585FA6CA" box="[378,527,2009,2033]" italics="true" pageId="2" pageNumber="251">pars stridens</emphasis>
) on vertex, present in both sexes. Semilunar notch in posterolateral region of cephalic capsule. Sub-rectangular labrum, with a triangular area and middle button of thicker cuticle. Clypeus with slightly arcuate anterior margin, curved outwards. Moniliform antennae with 11 rounded antennomeres progressively more quadrate (
<emphasis id="B9344D31F96CFFACFE3E5CAF5839A1CC" bold="true" box="[508,617,222,247]" pageId="3" pageNumber="252">morph 1</emphasis>
, as defined in 
<bibRefCitation id="EFD1ECD2F96CFFACFCE85CAF5E96A1CC" author="Perez-Gonzalez" box="[810,1222,223,247]" pageId="3" pageNumber="252" refString="Perez-Gonzalez, S. &amp; Zaballos, J. P. (2013 b) Antennal morphology of the endogean carabid genus Typhlocharis (Coleoptera: Carabidae: Anillini): description of sensilla and taxonomic implications. Journal of Morphology, 274, 809 - 823 http: // dx. doi. org / 10.1002 / jmor. 20140." type="journal article" year="2013" yearSuffix="b">Pérez-González &amp; Zaballos 2013b</bibRefCitation>
), except last one, pyriform. The last antennomere has a pattern of three antero-dorsal and one postero-dorsal 
<emphasis id="B9344D31F96CFFACFB6D5D725FC4A021" box="[1199,1428,258,282]" italics="true" pageId="3" pageNumber="252">sensilla coeloconica</emphasis>
. Ventral 
<emphasis id="B9344D31F96CFFACFF315D565B8AA005" box="[243,474,294,318]" italics="true" pageId="3" pageNumber="252">sensilla coeloconica</emphasis>
in antennomeres 5th and 6th. Stem of third antennomere not elongate. Mandibles without terebral teeth. Labium with sharp, pointy epilobes, separated by a wide space, and blunt middle tooth. Ligula with convex middle lobe and long paraglossae. Gula partially fused to cephalic capsule (suture only visible in the proximal and distal ends, diffuse in the middle). Cephalic chaetotaxy: five or six pairs of labral setae (
<emphasis id="B9344D31F96CFFACFA925DFF5BC1A0F7" bold="true" pageId="3" pageNumber="252">s-(s)-lm-s-m / m-s-m-l-(s)-s</emphasis>
), two pairs of clypeal setae (
<emphasis id="B9344D31F96CFFACFD165DC45971A0F7" bold="true" box="[724,801,435,460]" pageId="3" pageNumber="252">l-s / s-l</emphasis>
), one pair of frontal setae, two pairs of supraocular setae (anterior and posterior), one pair of supra-antennal setae, one pair of vertical setae, three pairs of occipital setae and two pairs of genal setae, as well as scattered pubescence. Labium with a pair of setae near base of middle tooth, a pair of long setae near base of epilobes, a pair of very short setae near apex of epilobes and a pair of very short setae near posterior suture. Prebasilar with two pairs of setae near anterior margin (lateral pair much longer) and two pairs of setae (lateral pair much longer) in posterior region, with some degree of individual variation over this basic pattern.
</paragraph>
<paragraph id="8BFF9123F96CFFACFF055EDD59F6A571" blockId="3.[151,1437,151,2020]" pageId="3" pageNumber="252">
Pronotum: subquadrate, slightly longer (
<quantity id="4CB83CC6F96CFFACFD645EDC5902A3FF" box="[678,850,684,708]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="3.2" metricValueMin="2.8" pageId="3" pageNumber="252" unit="mm" value="0.3" valueMax="0.32" valueMin="0.28">0.28–0.32 mm</quantity>
) than wide (
<quantity id="4CB83CC6F96CFFACFC315EDC5EF2A3FF" box="[1011,1186,684,708]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="3" pageNumber="252" unit="mm" value="0.285" valueMax="0.3" valueMin="0.27">0.27–0.30 mm</quantity>
), subtly narrowed in posterior region. Anterior margin straight, slightly crenulate, with medial hiatus (approx. as wide as two adjacent intersetae spaces); posterior margin smoothly sinuate. Lateral margins with three-four faint to very faint denticles near posterior angles. Surface covered by subhexagonal microreticulation. Disc flattened, with a strongly developed medial line and a pair of faint lateral sulci. Chaetotaxy: one pair of long setae in first third of lateral margins, one pair of long setae near posterior angles, a row of seven to eight pairs of setae [
<emphasis id="B9344D31F96CFFACFB5C5F2D5A83A2A2" bold="true" pageId="3" pageNumber="252">l-(l)-l-l-l-l-l-l / l-l-l-l-ll-(l)-l</emphasis>
] parallel to anterior margin, four pairs of setae parallel to posterior margin 
<emphasis id="B9344D31F96CFFACFBFC5FF15F6FA2A2" bold="true" box="[1086,1343,896,921]" pageId="3" pageNumber="252">[m-m-l-m / m-l-m-m]</emphasis>
, row of small setae, regularly placed in anterior and posterior margins (bulb-shaped setae, thickened at base in anterior margin), row of short setae in lateral margins and approximately four pairs of longitudinal rows of short pubescence in disc, but irregularly distributed. Proepisternal suture marked. Prosternal apophysis rounded. Anterior margin of prosternum with a row of long, thin setae and six to eight pairs of short setae parallel to it. Prosternum covered in scattered pubescence, glabrous proepisternum.
</paragraph>
<paragraph id="8BFF9123F96CFFACFF0558255EACA4B2" blockId="3.[151,1437,151,2020]" pageId="3" pageNumber="252">
Elytra: approximately twice as long (
<quantity id="4CB83CC6F96CFFACFDB75825594FA556" box="[629,799,1109,1133]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="6.2" metricValueMin="5.2" pageId="3" pageNumber="252" unit="mm" value="0.57" valueMax="0.62" valueMin="0.52">0.52–0.62 mm</quantity>
) as wide (
<quantity id="4CB83CC6F96CFFACFC5858255E16A556" box="[922,1094,1109,1133]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="3" pageNumber="252" unit="mm" value="0.285" valueMax="0.3" valueMin="0.27">0.27–0.30 mm</quantity>
), subparallel. Lateral margins with 20–22 subtriangular denticles, well developed in all the length, &quot;hook-shaped&quot; in the first third. Apical margin with two pairs of large, pointy denticles (a pair associated to the end of 7th stria and a pair near the suture). Disc flattened, with longitudinal lateral carinae associated to 7th stria. Surface covered by irregular subhexagonal microreticulation. Slight elytral pores along 7th stria; scattered pores also in scutellar and discal region. Transverse scutellar organ (smoothly subtriangular or arquate to irregular) and pair of &quot;buttonholes&quot; present near base of elytra. Chaetotaxy: umbilicate series formed by anterior group of four setae and posterior group of three setae (
<figureCitation id="137B8DA6F96CFFACFAAC595A5AF4A45D" captionStart="FIGURE 1" captionStartId="4.[151,250,1914,1937]" captionTargetBox="[177,1392,211,1861]" captionTargetId="figure@4.[155,1432,195,1890]" captionTargetPageId="4" captionText="FIGURE 1. Typhlocharis baeturica sp. n. – habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222600/files/figure.png" pageId="3" pageNumber="252">Fig. 1</figureCitation>
). One pair of scutellar setae. No discal setae. Discal pubescence arranged in five or six pairs of longitudinal rows, those of the third row alternative long and short. Lateral margins with a short seta for every denticle.
</paragraph>
<paragraph id="8BFF9123F96CFFACFF0559E55B64A79E" blockId="3.[151,1437,151,2020]" pageId="3" pageNumber="252">
Legs: similar in both sexes. Wide intermetacoxal space. Metacoxae with a pointy triangular, dentiform &quot;flap&quot;. Rounded metatrochanters. Inner margin of femora with small scaly prominences. Slightly angular metafemora and metatibiae with dilated distal region. All tarsi with four tarsomeres (tarsal formula: 4-4-4), apparently from fusion of tarsomeres 4th and 5th. Abdomen: covered by irregular microreticulation, the last segment present a belt of scaly microsculpture, scales with finely serrated edge in both sexes. Females have a pair of foveae on first ventrite (sternum II); second ventrite (sternum III) with a slight pair of foveae. Males lack ventral foveae. Last ventrite with a pair of lateral teeth and six to eight pairs of setae in posterior margin, sexually dimorphic: 
<emphasis id="B9344D31F96CFFACFB555A195A8AA79E" bold="true" pageId="3" pageNumber="252">l-(s)-s-s-l-s-s / s-l-s-l-ss-(s)-l</emphasis>
(
<figureCitation id="137B8DA6F96CFFACFF285AFD5B77A79E" box="[234,295,1677,1701]" captionStart="FIGURE 1" captionStartId="4.[151,250,1914,1937]" captionTargetBox="[177,1392,211,1861]" captionTargetId="figure@4.[155,1432,195,1890]" captionTargetPageId="4" captionText="FIGURE 1. Typhlocharis baeturica sp. n. – habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex." httpUri="https://zenodo.org/record/222600/files/figure.png" pageId="3" pageNumber="252">Fig.1</figureCitation>
).
</paragraph>
<paragraph id="8BFF9123F96CFFACFF055AC059EBA634" blockId="3.[151,1437,151,2020]" pageId="3" pageNumber="252">
Aedeagus: sickle-shaped median lobe (length: 
<quantity id="4CB83CC6F96CFFACFD1F5AC05914A7F3" box="[733,836,1712,1736]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="252" unit="mm" value="0.17">0.17 mm</quantity>
) with subtriangular, smoothly rounded apex, bent to right (anatomically oriented) in dorsal view (
<figureCitation id="137B8DA6F96CFFACFD545AA4588BA7D7" box="[662,731,1748,1772]" captionStart="FIGURE 2" captionStartId="5.[151,250,952,975]" captionTargetBox="[207,1365,203,903]" captionTargetId="figure@5.[183,1393,177,927]" captionTargetPageId="5" captionText="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="3" pageNumber="252">Fig. 2</figureCitation>
A,B). Endophallus with two pieces, &quot;scissor-shaped&quot; sclerites. Parameres subtriangular, with two medium-sized apical setae (
<figureCitation id="137B8DA6F96CFFACFC915A8759C7A634" box="[851,919,1783,1807]" captionStart="FIGURE 2" captionStartId="5.[151,250,952,975]" captionTargetBox="[207,1365,203,903]" captionTargetId="figure@5.[183,1393,177,927]" captionTargetPageId="5" captionText="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="3" pageNumber="252">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8BFF9123F96CFFACFF055B6B5BD5A6DF" blockId="3.[151,1437,151,2020]" pageId="3" pageNumber="252">
Female genitalia: adjusts to the description of 
<bibRefCitation id="EFD1ECD2F96CFFACFD135B6B5985A608" author="Vigna-Taglianti" box="[721,981,1819,1843]" pageId="3" pageNumber="252" refString="Vigna-Taglianti, A. (1972) Un nuovo Typhlocharis di Spagna. (Col., Carab.). Bolletino della Societa Entomologica Italiana. Societa Entomologica Italiana, 104, 148 - 156." type="journal article" year="1972">Vigna-Taglianti (1972)</bibRefCitation>
. Tubular, slender, gonocoxites, with one apical setae. Two setae in distal region of the gonocoxite, a longer, ventral one (pointing outwards), and a shorter, dorsal one (pointing inwards). Lateral pores present in the gonocoxites. Acute gonosubcoxites. Moderately long gonoduct, with two regions: thinner, proximal (diameter: 
<quantity id="4CB83CC6F96CFFACFCF75BF559FDA6A6" box="[821,941,1925,1949]" metricMagnitude="-6" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="252" unit="mm" value="0.002">0.002 mm</quantity>
) and thicker, distal (diameter: 
<quantity id="4CB83CC6F96CFFACFAD45BF55FC0A6A6" box="[1302,1424,1925,1949]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="252" unit="mm" value="0.006">0.006 mm</quantity>
). Subsphaeric spermatheca (length: 
<quantity id="4CB83CC6F96CFFACFDE45BD958F2A6FA" box="[550,674,1961,1985]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="252" unit="mm" value="0.015">0.015 mm</quantity>
). Large, conical spermathecal gland (length: 
<quantity id="4CB83CC6F96CFFACFB755BD95F62A6FA" box="[1207,1330,1961,1985]" metricMagnitude="-5" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="252" unit="mm" value="0.029">0.029 mm</quantity>
), distally sclerotized (
<figureCitation id="137B8DA6F96CFFACFEDD5BBC5B33A6DF" box="[287,355,1996,2020]" captionStart="FIGURE 2" captionStartId="5.[151,250,952,975]" captionTargetBox="[207,1365,203,903]" captionTargetId="figure@5.[183,1393,177,927]" captionTargetPageId="5" captionText="FIGURE 2. Typhlocharis baeturica sp. n. (A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view)." httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="3" pageNumber="252">Fig. 2</figureCitation>
C).
</paragraph>
<caption id="DF3FC1ABF96BFFABFF555B0A5907A694" httpUri="https://zenodo.org/record/222600/files/figure.png" pageId="4" pageNumber="253" targetBox="[177,1392,211,1861]" targetPageId="4">
<paragraph id="8BFF9123F96BFFABFF555B0A5907A694" blockId="4.[151,1437,1914,1967]" pageId="4" pageNumber="253">
<emphasis id="B9344D31F96BFFABFF555B0A5B48A6AB" bold="true" box="[151,280,1914,1937]" pageId="4" pageNumber="253">FIGURE 1.</emphasis>
<taxonomicName id="4C40EAA0F96BFFABFEE15B0A5843A6AB" box="[291,531,1914,1936]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F96BFFABFEE15B0A5843A6AB" box="[291,531,1914,1936]" italics="true" pageId="4" pageNumber="253">Typhlocharis baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF96BFFABFDDC5B0B5801A6AB" box="[542,593,1915,1936]" pageId="4" pageNumber="253" rank="species">
<emphasis id="B9344D31F96BFFABFDDC5B0B5801A6AB" bold="true" box="[542,593,1915,1936]" pageId="4" pageNumber="253">sp.n.</emphasis>
</taxonomicNameLabel>
–habitus: ventral (left half: male, right half: female) and dorsal (male) views. Abdomen not shown in dorsal view for a clearer view of elytral apex.
</paragraph>
</caption>
<caption id="DF3FC1ABF96AFFAAFF555FC858DFA2D6" httpUri="https://zenodo.org/record/222601/files/figure.png" pageId="5" pageNumber="254" targetBox="[207,1365,203,903]" targetPageId="5">
<paragraph id="8BFF9123F96AFFAAFF555FC858DFA2D6" blockId="5.[151,1436,952,1006]" pageId="5" pageNumber="254">
<emphasis id="B9344D31F96AFFAAFF555FC85B46A2F5" bold="true" box="[151,278,952,975]" pageId="5" pageNumber="254">FIGURE 2.</emphasis>
<taxonomicName id="4C40EAA0F96AFFAAFEDD5FC8585DA2F5" box="[287,525,952,974]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F96AFFAAFEDD5FC8585DA2F5" box="[287,525,952,974]" italics="true" pageId="5" pageNumber="254">Typhlocharis baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF96AFFAAFDD45FC95819A2F5" box="[534,585,953,974]" pageId="5" pageNumber="254" rank="species">
<emphasis id="B9344D31F96AFFAAFDD45FC95819A2F5" bold="true" box="[534,585,953,974]" pageId="5" pageNumber="254">sp.n.</emphasis>
</taxonomicNameLabel>
(A, B) Aedeagus: A) median lobe (dorsal view); B) median lobe and parameres (lateral view). C) Female genitalia (ventral view).
</paragraph>
</caption>
<paragraph id="8BFF9123F96AFFAAFF0558695BACA431" blockId="5.[151,1436,1049,2011]" pageId="5" pageNumber="254">
<emphasis id="B9344D31F96AFFAAFF0558695B7AA509" bold="true" box="[199,298,1049,1074]" pageId="5" pageNumber="254">Habitat.</emphasis>
<taxonomicName id="4C40EAA0F96AFFAAFEF3586B5BE6A509" box="[305,438,1050,1074]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F96AFFAAFEF3586B5BE6A509" box="[305,438,1050,1074]" italics="true" pageId="5" pageNumber="254">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF96AFFAAFE7D586A5BA7A509" box="[447,503,1050,1074]" pageId="5" pageNumber="254" rank="species">
<emphasis id="B9344D31F96AFFAAFE7D586A5BA7A509" bold="true" box="[447,503,1050,1074]" pageId="5" pageNumber="254">sp.n.</emphasis>
</taxonomicNameLabel>
was collected in two localities, separated 
<quantity id="4CB83CC6F96AFFAAFC12586A5E72A509" box="[976,1058,1050,1074]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="254" unit="km" value="2.2">2.2 km</quantity>
: 1) stony slope and valley bottom in open holm oak forest (
<taxonomicName id="4C40EAA0F96AFFAAFE0D584F5832A56D" box="[463,610,1086,1110]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="254" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B9344D31F96AFFAAFE0D584F5832A56D" box="[463,610,1086,1110]" italics="true" pageId="5" pageNumber="254">Quercus ilex</emphasis>
</taxonomicName>
L.), with scarce laudanum shrubs (
<taxonomicName id="4C40EAA0F96AFFAAFBD2584F5E99A56D" box="[1040,1225,1086,1110]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="254" phylum="Tracheophyta" rank="species" species="ladanifer">
<emphasis id="B9344D31F96AFFAAFBD2584F5E99A56D" box="[1040,1225,1086,1110]" italics="true" pageId="5" pageNumber="254">Cistus ladanifer</emphasis>
</taxonomicName>
L.), olives (
<taxonomicName id="4C40EAA0F96AFFAAFAA7584E5B50A541" class="Gastropoda" family="Limapontiidae" genus="Olea" kingdom="Animalia" order="Sacoglossa" pageId="5" pageNumber="254" phylum="Mollusca" rank="species" species="europaea">
<emphasis id="B9344D31F96AFFAAFAA7584E5B50A541" italics="true" pageId="5" pageNumber="254">Olea europaea</emphasis>
</taxonomicName>
L.), almonds (
<taxonomicName id="4C40EAA0F96AFFAAFE6A58135971A541" authority="(Mill.) D.A. Webb" authorityName="(Mill.) D.A. Webb" box="[424,801,1122,1146]" class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="254" phylum="Tracheophyta" rank="species" species="dulcis">
<emphasis id="B9344D31F96AFFAAFE6A58135814A541" box="[424,580,1122,1146]" italics="true" pageId="5" pageNumber="254">Prunus dulcis</emphasis>
(Mill.) D.A. Webb
</taxonomicName>
), plantains (
<emphasis id="B9344D31F96AFFAAFC6958125E63A541" box="[939,1075,1122,1146]" italics="true" pageId="5" pageNumber="254">
<taxonomicName id="4C40EAA0F96AFFAAFC6958125E42A541" box="[939,1042,1122,1146]" class="Magnoliopsida" family="Plantaginaceae" genus="Plantago" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="254" phylum="Tracheophyta" rank="genus">Plantago</taxonomicName>
sp
</emphasis>
. L.) and asphodels (
<emphasis id="B9344D31F96AFFAAFAD958125AE7A5A5" italics="true" pageId="5" pageNumber="254">
<taxonomicName id="4C40EAA0F96AFFAAFAD958125FCCA541" box="[1307,1436,1122,1146]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Asphodelus" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="254" phylum="Tracheophyta" rank="genus">Asphodelus</taxonomicName>
sp.
</emphasis>
L.). Clayey soil with abundant and different sized boulders. 2) Top of hill in pasture-like terrain, with a recent holm oak plantation (scarce young trees aligned in rows), ploughed soil, boulders of shale (Upper Carboniferous) and with laudanum shrubs. It coexists in both localities with 
<taxonomicName id="4C40EAA0F96AFFAAFC9B58BE5F1CA5DD" authority="Serrano &amp; Aguiar, 2006" authorityName="Serrano &amp; Aguiar" authorityYear="2006" box="[857,1356,1230,1254]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="lunai">
<emphasis id="B9344D31F96AFFAAFC9B58BE5E7EA5DD" box="[857,1070,1230,1254]" italics="true" pageId="5" pageNumber="254">Typhlocharis lunai</emphasis>
Serrano &amp; Aguiar, 2006
</taxonomicName>
and 
<taxonomicName id="4C40EAA0F96AFFAAFA4558BF5BA8A431" authority="Serrano &amp; Aguiar, 2002" authorityName="Serrano &amp; Aguiar" authorityYear="2002" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="B9344D31F96AFFAAFA4558BF5AB0A431" italics="true" pageId="5" pageNumber="254">T. elenae</emphasis>
<bibRefCitation id="EFD1ECD2F96AFFAAFF2558825BA8A431" author="Serrano" box="[231,504,1266,1290]" pageId="5" pageNumber="254" refString="Serrano, A. R. M. &amp; Aguiar, C. A. S. (2002) The genus Typhlocharis Dieck, 1869 (Coleoptera: Carabidae) in Portugal: Description of two new species and faunistic notes. Boletim da Sociedade Portuguesa de Entomologia, 7, 181 - 197. http: // dx. doi. org / 10.3989 / graellsia. 2008. v 64. i 2.38" type="journal article" year="2002">Serrano &amp; Aguiar, 2002</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35AC2A8F96AFFA9FF0559665E7CA097" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254" type="etymology">
<paragraph id="8BFF9123F96AFFAAFF055966580BA44D" blockId="5.[151,1436,1049,2011]" pageId="5" pageNumber="254">
<emphasis id="B9344D31F96AFFAAFF0559665B1EA415" bold="true" box="[199,334,1302,1326]" pageId="5" pageNumber="254">Etymology.</emphasis>
The Roman 
<taxonomicName id="4C40EAA0F96AFFAAFE215967589BA415" authority="Celticorum" authorityName="Celticorum" box="[483,715,1302,1326]" class="Insecta" family="Cicadidae" genus="Baeturia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B9344D31F96AFFAAFE215967589BA415" box="[483,715,1302,1326]" italics="true" pageId="5" pageNumber="254">Baeturia Celticorum</emphasis>
</taxonomicName>
was defined by Caius Plinius Secundus (
<emphasis id="B9344D31F96AFFAAFB6059665F20A414" bold="true" box="[1186,1392,1302,1327]" italics="true" pageId="5" pageNumber="254">Naturalis historia</emphasis>
III: pp. 13–14) as the territory between rivers 
<taxonomicName id="4C40EAA0F96AFFAAFD59594B58B1A469" box="[667,737,1339,1362]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B9344D31F96AFFAAFD59594B58B1A469" box="[667,737,1339,1362]" italics="true" pageId="5" pageNumber="254">Baetis</emphasis>
</taxonomicName>
–now Guadalquivir- and 
<taxonomicName id="4C40EAA0F96AFFAAFBD9594A5E03A469" box="[1051,1107,1338,1362]" class="Aves" family="Anatidae" genus="Anas" kingdom="Animalia" order="Anseriformes" pageId="5" pageNumber="254" phylum="Chordata" rank="genus">
<emphasis id="B9344D31F96AFFAAFBD9594A5E03A469" box="[1051,1107,1338,1362]" italics="true" pageId="5" pageNumber="254">Anas</emphasis>
</taxonomicName>
–now Guadiana- bordering 
<collectingCountry id="F357D1B3F96AFFAAFF55592E5B52A44D" box="[151,258,1374,1398]" name="Portugal" pageId="5" pageNumber="254">Lusitania</collectingCountry>
, where this species was found.
</paragraph>
<paragraph id="8BFF9123F96AFFAAFF0559F15B2DA7AD" blockId="5.[151,1436,1049,2011]" pageId="5" pageNumber="254">
<emphasis id="B9344D31F96AFFAAFF0559F15B1DA4A1" bold="true" box="[199,333,1409,1434]" pageId="5" pageNumber="254">Variability.</emphasis>
There is certain degree of common intraspecific variation affecting some characters, such as chaetotaxy of labium and basilar, number and marking of posterolateral denticles in pronotum (from faintly marked to almost absent), marking of the hook-shaped lateral denticles of elytra, marking of sutural denticles of elytra or chaetotaxy of the last ventrite. The transverse scutellar organ varies between subtriangular and pointed, smoothly and slightly curved and roughly and irregularly curved. It is also common that some individuals show short and long setae irregularly mixed in the anterior row of setae in pronotum. The apodemal ring of male genitalia shows more variation than observed for other species, with an anterior projection ranging from narrow and prominent to more broad and low.
</paragraph>
<paragraph id="8BFF9123F96AFFAAFF055AD25882A7E5" blockId="5.[151,1436,1049,2011]" pageId="5" pageNumber="254">
One of the female 
<typeStatus id="54FB2F81F96AFFAAFE625AD3585BA780" box="[416,523,1699,1723]" pageId="5" pageNumber="254" type="paratype">paratypes</typeStatus>
lacks the hiatus in the anterior margin of pronotum. One of the male 
<typeStatus id="54FB2F81F96AFFAAFAF35AD35FCCA780" box="[1329,1436,1699,1723]" pageId="5" pageNumber="254" type="paratype">paratypes</typeStatus>
exhibits a malformed, abnormally narrow, clypeus.
</paragraph>
<paragraph id="8BFF9123F96AFFA9FF055A995EA4A1EF" blockId="5.[151,1436,1049,2011]" lastBlockId="6.[151,1436,151,428]" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254">
<emphasis id="B9344D31F96AFFAAFF055A995B6CA639" bold="true" box="[199,316,1769,1794]" pageId="5" pageNumber="254">Affinities.</emphasis>
The new species have an umbilicate series of 4+3 setae, which places it in the 
<taxonomicName id="4C40EAA0F96AFFAAFB0A5A9A5F5BA639" box="[1224,1291,1770,1794]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="diecki">
<emphasis id="B9344D31F96AFFAAFB0A5A9A5F5BA639" box="[1224,1291,1770,1794]" italics="true" pageId="5" pageNumber="254">diecki</emphasis>
</taxonomicName>
group 
<emphasis id="B9344D31F96AFFAAFA9C5A9B5FCCA639" box="[1374,1436,1771,1794]" italics="true" pageId="5" pageNumber="254">sensu</emphasis>
<bibRefCitation id="EFD1ECD2F96AFFAAFF555B7E584CA61D" author="Zaballos" box="[151,540,1806,1830]" pageId="5" pageNumber="254" refString="Zaballos, J. P. &amp; Ruiz-Tapiador, I. (1997) Nuevos Typhlocharis Dieck (Coleoptera: Caraboidea: Trechidae) de Espana. Graellsia, 52, 95 - 106. http: // dx. doi. org / 10.3989 / graellsia. 1996. v 52. i 0.381" type="journal article" year="1997">Zaballos &amp; Ruiz-Tapiador (1997)</bibRefCitation>
. Nevertheless, the possession of a diffuse gula and four tarsomeres is a rare condition only known, but not described, in other species of the genus: 
<taxonomicName id="4C40EAA0F96AFFAAFC115B425A83A655" authority="Serrano &amp; Aguiar, 2008" authorityName="Serrano &amp; Aguiar" authorityYear="2008" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F96AFFAAFC115B425E90A671" box="[979,1216,1842,1866]" italics="true" pageId="5" pageNumber="254">Typhlocharis crespoi</emphasis>
<bibRefCitation id="EFD1ECD2F96AFFAAFB0A5B425A83A655" author="Serrano" pageId="5" pageNumber="254" refString="Serrano, A. R. M. &amp; Aguiar C. A. S. (2008) Two new species of Typhlocharis Dieck, 1869 (Coleoptera, Carabidae) from Portugal: Description and notes on related species. G raellsia, 64, 281 - 293. http: // dx. doi. org / 10.3989 / graellsia. 2008. v 64. i 2.38" type="journal article" year="2008">Serrano &amp; Aguiar, 2008</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="4C40EAA0F96AFFAAFF205B275831A655" authority="Coiffait, 1969" authorityName="Coiffait" authorityYear="1969" box="[226,609,1878,1902]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F96AFFAAFF205B275BF0A655" box="[226,416,1878,1902]" italics="true" pageId="5" pageNumber="254">T. quadridentata</emphasis>
(
<bibRefCitation id="EFD1ECD2F96AFFAAFE765B265808A655" author="Coiffait" box="[436,600,1878,1902]" pageId="5" pageNumber="254" refString="Coiffait, H. (1969) Nouveaux Anillini du Maroc et Sud de la peninsule iberique. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 48, 55 - 56." type="journal article" year="1969">Coiffait, 1969</bibRefCitation>
)
</taxonomicName>
also possess a diffuse gula, but five tarsomeres. Both species are the morphologically closest to 
<taxonomicName id="4C40EAA0F96AFFAAFE055B0B581BA6A9" box="[455,587,1914,1938]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="baeturica" status="sp. nov.">
<emphasis id="B9344D31F96AFFAAFE055B0B581BA6A9" box="[455,587,1914,1938]" italics="true" pageId="5" pageNumber="254">T. baeturica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A207F04AF96AFFAAFD975B0A58DCA6A9" box="[597,652,1914,1938]" pageId="5" pageNumber="254" rank="species">
<emphasis id="B9344D31F96AFFAAFD975B0A58DCA6A9" bold="true" box="[597,652,1914,1938]" pageId="5" pageNumber="254">sp.n.</emphasis>
</taxonomicNameLabel>
, sharing a very small body size, shape of elytral apex, construction of aedeagus and parameres and particular shape of the labium (with prominent and pointy epilobes separated by a wide gap with a low medial tooth). However, the new species is well differentiated from both, by the 4+3 umbilicate series instead of 4+2, presence of stridulatory organ (absent in 
<taxonomicName id="4C40EAA0F969FFA9FC345CE95EE3A18B" box="[1014,1203,152,176]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F969FFA9FC345CE95EE3A18B" box="[1014,1203,152,176]" italics="true" pageId="6" pageNumber="255">T. quadridentata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C40EAA0F969FFA9FB325CE95F0EA18B" box="[1264,1374,153,176]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F969FFA9FB325CE95F0EA18B" box="[1264,1374,153,176]" italics="true" pageId="6" pageNumber="255">T. crespoi</emphasis>
</taxonomicName>
) and slender, tubular gonocoxites, unguiform-like in 
<taxonomicName id="4C40EAA0F969FFA9FD6F5CCD5938A1EF" box="[685,872,188,212]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B9344D31F969FFA9FD6F5CCD5938A1EF" box="[685,872,188,212]" italics="true" pageId="6" pageNumber="255">T. quadridentata</emphasis>
</taxonomicName>
and tubular, but short in 
<taxonomicName id="4C40EAA0F969FFA9FB435CCD5EBDA1EF" box="[1153,1261,189,212]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="crespoi">
<emphasis id="B9344D31F969FFA9FB435CCD5EBDA1EF" box="[1153,1261,189,212]" italics="true" pageId="6" pageNumber="255">T. crespoi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFF9123F969FFA9FF055C905E7CA097" blockId="6.[151,1436,151,428]" pageId="6" pageNumber="255">
Apart from the diffuse gula, the new species has many features that resembles 
<taxonomicName id="4C40EAA0F969FFA9FBB15C915F27A1CC" authority="Ehlers, 1883" authorityName="Ehlers" authorityYear="1883" box="[1139,1399,223,248]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="diecki">
<emphasis id="B9344D31F969FFA9FBB15C915E86A1C3" box="[1139,1238,224,248]" italics="true" pageId="6" pageNumber="255">T. diecki</emphasis>
Ehlers, 1883
</taxonomicName>
; 
<taxonomicName id="4C40EAA0F969FFA9FA455C915BF0A027" authority="Novoa, 1978" authorityName="Novoa" authorityYear="1978" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="outereloi">
<emphasis id="B9344D31F969FFA9FA455C915AABA027" italics="true" pageId="6" pageNumber="255">T. outereloi</emphasis>
Novoa, 1978
</taxonomicName>
; 
<taxonomicName id="4C40EAA0F969FFA9FE735D755911A027" authority="Serrano &amp; Aguiar, 2006" authorityName="Serrano &amp; Aguiar" authorityYear="2006" box="[433,833,260,284]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="bivari">
<emphasis id="B9344D31F969FFA9FE735D755843A027" box="[433,531,260,284]" italics="true" pageId="6" pageNumber="255">T. bivari</emphasis>
Serrano &amp; Aguiar, 2006
</taxonomicName>
; 
<taxonomicName id="4C40EAA0F969FFA9FC905D755F57A027" authority="Zaballos &amp; Wrase, 1998" authorityName="Zaballos &amp; Wrase" authorityYear="1998" box="[850,1287,260,284]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="navarica">
<emphasis id="B9344D31F969FFA9FC905D755987A027" box="[850,983,261,284]" italics="true" pageId="6" pageNumber="255">T. navarica</emphasis>
<bibRefCitation id="EFD1ECD2F969FFA9FC215D745F57A027" author="Zaballos" box="[995,1287,260,284]" pageId="6" pageNumber="255" refString="Zaballos, J. P. &amp; Wrase, D. (1998) Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia, 54, 43 - 52." type="journal article" year="1998">Zaballos &amp; Wrase, 1998</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C40EAA0F969FFA9FAD55D755B1AA004" authority="Normand, 1915" authorityName="Normand" authorityYear="1915" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="sanstchii">
<emphasis id="B9344D31F969FFA9FAD55D755FCCA027" box="[1303,1436,260,284]" italics="true" pageId="6" pageNumber="255">T. sanstchii</emphasis>
Normand, 1915
</taxonomicName>
or 
<taxonomicName id="4C40EAA0F969FFA9FEB75D595978A004" authority="Serrano &amp; Aguiar, 2008" authorityName="Serrano &amp; Aguiar" authorityYear="2008" box="[373,808,295,320]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="rochapitei">
<emphasis id="B9344D31F969FFA9FEB75D595855A07B" box="[373,517,296,320]" italics="true" pageId="6" pageNumber="255">T. rochapitei</emphasis>
<bibRefCitation id="EFD1ECD2F969FFA9FDCD5D575978A004" author="Serrano" box="[527,808,295,319]" pageId="6" pageNumber="255" refString="Serrano, A. R. M. &amp; Aguiar C. A. S. (2008) Two new species of Typhlocharis Dieck, 1869 (Coleoptera, Carabidae) from Portugal: Description and notes on related species. G raellsia, 64, 281 - 293. http: // dx. doi. org / 10.3989 / graellsia. 2008. v 64. i 2.38" type="journal article" year="2008">Serrano &amp; Aguiar, 2008</bibRefCitation>
</taxonomicName>
; such as the presence of a wide intermetacoxal space, shape of antennomeres, pattern of basilar chaetotaxy, presence of stridulatory organ, very faint or absent elytral pores, lateral carinae of elytra associated to apical denticles or morphology and chaetotaxy of last ventrite. Within these, 
<taxonomicName id="4C40EAA0F969FFA9FF1C5DE55B6BA097" box="[222,315,404,428]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="bivari">
<emphasis id="B9344D31F969FFA9FF1C5DE55B6BA097" box="[222,315,404,428]" italics="true" pageId="6" pageNumber="255">T. bivari</emphasis>
</taxonomicName>
also show a similar shaped aedeagus, with two endophallic pieces.
</paragraph>
</subSubSection>
</treatment>
</document>