534 lines
69 KiB
XML
534 lines
69 KiB
XML
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<mods:title id="456539DDD41D049B509E35B07F0A29B6">A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): systematic and biogeographical implications</mods:title>
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<mods:namePart id="7BA43F0F7D8405A791D7D158F4DCE11D">Geniez, Philippe</mods:namePart>
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<mods:namePart id="BB5EF9B7E1D8853A0B7864196841BF2A">Ineich, Ivan</mods:namePart>
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<mods:date id="3A11B37E33A2A0AF6FC90E691EDB75F2">2003</mods:date>
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<treatment id="03C1AF1C831CFFF2FCCC3D071265FE1F" ID-DOI="http://doi.org/10.5281/zenodo.5490908" ID-GBIF-Taxon="185447195" ID-Zenodo-Dep="5490908" LSID="urn:lsid:plazi:treatment:03C1AF1C831CFFF2FCCC3D071265FE1F" httpUri="http://treatment.plazi.org/id/03C1AF1C831CFFF2FCCC3D071265FE1F" lastPageId="24" lastPageNumber="141" pageId="20" pageNumber="137">
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<subSubSection id="C3724D81831CFFFEFCCC3D071394F9EA" box="[816,1412,1565,1589]" pageId="20" pageNumber="137" type="nomenclature">
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<paragraph id="8BD71E0A831CFFFEFCCC3D071394F9EA" blockId="20.[816,1412,1565,1589]" box="[816,1412,1565,1589]" pageId="20" pageNumber="137">
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<heading id="D09FA966831CFFFEFCCC3D071394F9EA" box="[816,1412,1565,1589]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="137" reason="2">
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<taxonomicName id="4C686589831CFFFEFCCC3D071394F9EA" authority="(AUDOUIN, 1827)" baseAuthorityName="Audouin" baseAuthorityYear="1827" box="[816,1412,1565,1589]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="137" phylum="Chordata" rank="species" species="scutellatus">
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<emphasis id="B91CC218831CFFFEFCCC3D0712A8F9EB" box="[816,1208,1565,1589]" italics="true" pageId="20" pageNumber="137">ACANTHODACTYLUS SCUTELLATUS</emphasis>
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(
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<bibRefCitation id="EFF963FB831CFFFEFB343D07136DF9EA" author="Audouin V" box="[1224,1405,1565,1589]" pageId="20" pageNumber="137" pagination="1 - 350" refId="ref21438" refString="Audouin V. 1827. Reptiles (suppl.). Explication sommaire des planches de reptiles (supplement) publiees par Jules-Cesar Savigny. In: Description de l'Egypte ou Recueil des Observations et des Recherches qui ont ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise, Publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand. Histoire Naturelle. Tome Premier, 1 ere Partie, et Atlas. Paris: Imprimerie imperiale / Imprimerie royale, 1 - 350." type="book chapter" year="1827">AUDOUIN, 1827</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3724D81831CFFFFFCDA3D5D14C9FEB5" lastPageId="21" lastPageNumber="138" pageId="20" pageNumber="137" type="reference_group">
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<paragraph id="8BD71E0A831CFFFFFCDA3D5D17E0FF27" blockId="20.[806,1423,1606,1904]" lastBlockId="21.[155,771,196,248]" lastPageId="21" lastPageNumber="138" pageId="20" pageNumber="137">
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<treatmentCitationGroup id="AB783924831CFFFFFCDA3D5D17E0FF27" lastPageId="21" lastPageNumber="138" pageId="20" pageNumber="137">
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<treatmentCitation id="0AC9381B831CFFFEFCDA3D5D12E6F983" author="Audouin V" box="[806,1270,1606,1628]" page="172" pageId="20" pageNumber="137" year="1827">
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<taxonomicName id="4C686589831CFFFEFCDA3D5D12E6F983" authority="Audouin, 1827: 172" authorityName="Audouin" authorityPageNumber="172" authorityYear="1827" box="[806,1270,1606,1628]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="137" phylum="Chordata" rank="species" species="scutellata">
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<emphasis id="B91CC218831CFFFEFCDA3D5D15EAF984" box="[806,1018,1606,1628]" italics="true" pageId="20" pageNumber="137">Lacerta scutellata</emphasis>
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<bibRefCitation id="EFF963FB831CFFFEFBF53D5C12E6F983" author="Audouin V" box="[1033,1270,1606,1628]" pageId="20" pageNumber="137" pagination="1 - 350" refId="ref21438" refString="Audouin V. 1827. Reptiles (suppl.). Explication sommaire des planches de reptiles (supplement) publiees par Jules-Cesar Savigny. In: Description de l'Egypte ou Recueil des Observations et des Recherches qui ont ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise, Publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand. Histoire Naturelle. Tome Premier, 1 ere Partie, et Atlas. Paris: Imprimerie imperiale / Imprimerie royale, 1 - 350." type="book chapter" year="1827">Audouin, 1827: 172</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. i, fig. 7. Name-bearing type: the original description was clearly based on a single specimen (figured on the plate), which is thus the
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<typeStatus id="54D3A0A8831CFFFEFBC73DB8128FF967" box="[1083,1183,1698,1720]" pageId="20" pageNumber="137" type="holotype">holotype</typeStatus>
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. This specimen is not in the collections of the MNHN, and was perhaps never given to the MNHN collections (
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<treatmentCitation id="0AC9381B831CFFFEFB123DFA157EF8CB" author="Brygoo ER" page="44" pageId="20" pageNumber="137" year="1988">
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<bibRefCitation id="EFF963FB831CFFFEFB123DFA157EF8CB" author="Brygoo ER" pageId="20" pageNumber="137" pagination="3 - 57" refId="ref21773" refString="Brygoo ER. 1988. Les types de Lacertides (Reptiles, Sauriens) du Museum national d'Histoire naturelle. Catalogue critique. Bulletin du Museum National d'Histoire Naturelle, Section A, Zoologie, Biologie et Ecologie Animales (Supplement) 10: 3 - 57." type="journal article" year="1988">Brygoo, 1988: 44–45</bibRefCitation>
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</treatmentCitation>
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). It should be considered as lost. A
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<typeStatus id="54D3A0A8831CFFFEFAC93DE5139EF8CB" box="[1333,1422,1791,1812]" pageId="20" pageNumber="137" type="neotype">neotype</typeStatus>
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should be designated to stabilize the nomenclatural status of this taxon. We refrain from doing so here pending a more complete analysis of the populations from the eastern part of the species range, including
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<collectingCountry id="F37F5E9A831DFFFFFF673BF916F2FF27" box="[155,226,227,248]" name="Egypt" pageId="21" pageNumber="138">Egypt</collectingCountry>
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. Type locality: ‘Egypte’.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD71E0A831DFFFFFF483A0C14C9FEB5" blockId="21.[155,771,278,362]" pageId="21" pageNumber="138">
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<emphasis id="B91CC218831DFFFFFF483A0C172EFEF4" box="[180,318,278,299]" italics="true" pageId="21" pageNumber="138">Chresonyms</emphasis>
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.
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<taxonomicName id="4C686589831DFFFFFEB03A0C16CDFE94" authority="(Audouin, 1827)" baseAuthorityName="Audouin" baseAuthorityYear="1827" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="scutellatus">
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<emphasis id="B91CC218831DFFFFFEB03A0C149AFEF4" box="[332,650,278,299]" italics="true" pageId="21" pageNumber="138">Acanthodactylus scutellatus</emphasis>
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(
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<bibRefCitation id="EFF963FB831DFFFFFD663A0C16C5FE94" author="Audouin V" pageId="21" pageNumber="138" pagination="1 - 350" refId="ref21438" refString="Audouin V. 1827. Reptiles (suppl.). Explication sommaire des planches de reptiles (supplement) publiees par Jules-Cesar Savigny. In: Description de l'Egypte ou Recueil des Observations et des Recherches qui ont ete Faites en Egypte Pendant l'Expedition de l'Armee Francaise, Publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand. Histoire Naturelle. Tome Premier, 1 ere Partie, et Atlas. Paris: Imprimerie imperiale / Imprimerie royale, 1 - 350." type="book chapter" year="1827">Audouin, 1827</bibRefCitation>
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)
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||
</taxonomicName>
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:
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<bibRefCitation id="EFF963FB831DFFFFFF163A2C141AFE95" author="Bons J & Girot B" box="[234,522,309,331]" pageId="21" pageNumber="138" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964: 319</bibRefCitation>
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;
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<collectingCountry id="F37F5E9A831DFFFFFDE43A2F146DFE94" box="[536,637,309,331]" name="El Salvador" pageId="21" pageNumber="138">Salvador</collectingCountry>
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, 1982: 113;
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<bibRefCitation id="EFF963FB831DFFFFFF673A4E1776FEB6" author="Arnold EN" box="[155,358,340,362]" pageId="21" pageNumber="138" pagination="291 - 339" refId="ref21308" refString="Arnold EN. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History) Zoology 44: 291 - 339." type="journal article" year="1983">Arnold, 1983: 322</bibRefCitation>
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(part);
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<bibRefCitation id="EFF963FB831DFFFFFE413A4E1498FEB5" author="Arnold EN" box="[445,648,340,362]" pageId="21" pageNumber="138" pagination="385 - 435" refId="ref21343" refString="Arnold EN. 1986. A Key and annotated check list to the lizards and Amphisbaenians of Arabia. In: Buttiker W, Krupp F, eds. Fauna of Saudi Arabia, Vol. 8. Jeddah: Meteorology and Environmental Protection Administration, 385 - 435." type="book chapter" year="1986">Arnold, 1986: 425</bibRefCitation>
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(part).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3724D81831DFFF2FF483A9C17D1FE32" lastPageId="24" lastPageNumber="141" pageId="21" pageNumber="138" type="distribution">
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<paragraph id="8BD71E0A831DFFFFFF483A9C1784FD31" blockId="21.[155,771,390,750]" pageId="21" pageNumber="138">
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<emphasis id="B91CC218831DFFFFFF483A9C172FFE44" box="[180,319,390,411]" italics="true" pageId="21" pageNumber="138">Distribution</emphasis>
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(
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<figureCitation id="1353028F831DFFFFFEAF3A9C17B2FE42" box="[339,418,390,413]" captionStart="Figure 13" captionStartId="21.[155,233,1826,1845]" captionTargetBox="[157,1429,981,1786]" captionTargetId="figure-442@21.[156,1433,973,1793]" captionTargetPageId="21" captionText="Figure 13. Geographic distribution of Acanthodactylus scutellatus in the Sahara (thus excluding part of the species range, see text). Dotted lines indicate the approximate limits of the distribution of A. dumerili. Data from Salvador (1982), Baha El Din (1994), Joger & Lambert (1996), Nouïra (1996), this study." figureDoi="http://doi.org/10.5281/zenodo.5437181" httpUri="https://zenodo.org/record/5437181/files/figure.png" pageId="21" pageNumber="138">Fig. 13</figureCitation>
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). In the Sahara, south-eastern
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<collectingCountry id="F37F5E9A831DFFFFFF673ABF16E1FE64" box="[155,241,421,443]" name="Algeria" pageId="21" pageNumber="138">Algeria</collectingCountry>
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, north-eastern
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<collectingCountry id="F37F5E9A831DFFFFFE5D3ABF17C8FE64" box="[417,472,421,443]" name="Serbia" pageId="21" pageNumber="138">Mali</collectingCountry>
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, northern Niger, northern
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<collectingCountry id="F37F5E9A831DFFFFFF673ADE16CAFE05" box="[155,218,452,474]" name="Chad" pageId="21" pageNumber="138">Chad</collectingCountry>
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, northern
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<collectingCountry id="F37F5E9A831DFFFFFEA13ADE17B9FE05" box="[349,425,452,474]" name="Sudan" pageId="21" pageNumber="138">Sudan</collectingCountry>
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,
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<collectingCountry id="F37F5E9A831DFFFFFE463ADE1410FE06" box="[442,512,452,473]" name="Egypt" pageId="21" pageNumber="138">Egypt</collectingCountry>
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,
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<collectingCountry id="F37F5E9A831DFFFFFDED3ADE1444FE05" box="[529,596,452,474]" name="Libya" pageId="21" pageNumber="138">Libya</collectingCountry>
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, and southern
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<collectingCountry id="F37F5E9A831DFFFFFF673AF816E2FE27" box="[155,242,482,504]" name="Tunisia" pageId="21" pageNumber="138">Tunisia</collectingCountry>
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(
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<collectingCountry id="F37F5E9A831DFFFFFEF93AF8177BFE27" box="[261,363,482,504]" name="El Salvador" pageId="21" pageNumber="138">Salvador</collectingCountry>
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, 1982;
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<bibRefCitation id="EFF963FB831DFFFFFE3E3AF9144CFE27" author="Nouira S" box="[450,604,482,504]" pageId="21" pageNumber="138" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra, 1996</bibRefCitation>
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; Ch. P. Blanc, pers. com.; this study). The characters given above and a careful examination of the pictures published by
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<bibRefCitation id="EFF963FB831DFFFFFF6739251790FD8B" author="Bons J & Girot B" box="[155,384,574,596]" pageId="21" pageNumber="138" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot (1964)</bibRefCitation>
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lead us to reject the occurrence of this species in
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<collectingCountry id="F37F5E9A831DFFFFFEB7394417B8FDAC" box="[331,424,606,627]" name="Morocco" pageId="21" pageNumber="138">Morocco</collectingCountry>
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(cf.
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<bibRefCitation id="EFF963FB831DFFFFFE22394414C6FDAC" author="Bons J & Geniez Ph" box="[478,726,605,627]" pageId="21" pageNumber="138" refId="ref21662" refString="Bons J, Geniez Ph. 1996. Amphibiens et Reptiles du Maroc, Sahara Occidental compris. Atlas biogeographique. Barcelona: Asociacion Herpetologica Espanola." type="book" year="1996">Bons & Geniez, 1996</bibRefCitation>
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), in opposition to the opinions of
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<bibRefCitation id="EFF963FB831DFFFFFDFF396614EEFD4D" author="Bons J & Girot B" box="[515,766,636,658]" pageId="21" pageNumber="138" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot (1964)</bibRefCitation>
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,
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<bibRefCitation id="EFF963FB831DFFFFFF67398017D7FD6E" author="Mellado J & Olmedo G" box="[155,455,666,689]" pageId="21" pageNumber="138" pagination="131 - 146" refId="ref22578" refString="Mellado J, Olmedo G. 1990. El genero Acanthodactylus en Marruecos: problemas de identification en los grupos de especies A. pardalis y A. scutellatus. Amphibia-Reptilia 11: 131 - 146." type="journal article" year="1990">Mellado & Olmedo (1990)</bibRefCitation>
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and Pérez Mellado (1992). The species is also encountered in
|
||
<collectingCountry id="F37F5E9A831DFFFFFDCC39A31465FD10" box="[560,629,697,719]" name="Israel" pageId="21" pageNumber="138">Israel</collectingCountry>
|
||
, Arabia and
|
||
<collectingCountry id="F37F5E9A831DFFFFFF6739C216DBFD32" box="[155,203,728,749]" name="Iraq" pageId="21" pageNumber="138">Iraq</collectingCountry>
|
||
(
|
||
<collectingCountry id="F37F5E9A831DFFFFFF2739C21751FD31" box="[219,321,728,750]" name="El Salvador" pageId="21" pageNumber="138">Salvador</collectingCountry>
|
||
, 1982).
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A831DFFFDFF48381115A0F908" blockId="21.[155,772,778,892]" lastBlockId="23.[819,1435,1269,1751]" lastPageId="23" lastPageNumber="140" pageId="21" pageNumber="138">
|
||
<emphasis id="B91CC218831DFFFFFF4838111734FCFF" box="[180,292,779,800]" italics="true" pageId="21" pageNumber="138">Diagnosis</emphasis>
|
||
. (Remarks: the following diagnosis is based on specimens of
|
||
<taxonomicName id="4C686589831DFFFFFE4038331712FC82" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC218831DFFFFFE4038331712FC82" italics="true" pageId="21" pageNumber="138">Acanthodactylus scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
only). The largest species of the group (reaching
|
||
<quantity id="4C90B3EF831DFFFFFEF2387C1761FCA3" box="[270,369,870,892]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" pageId="21" pageNumber="138" unit="mm" value="72.5">72.5 mm</quantity>
|
||
snout-vent length, mean = 63.9; cf.
|
||
<tableCitation id="C6EA2BB1831DFFFFFCCF3BDE1598FF06" box="[819,904,196,218]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="21" pageNumber="138">Table 2</tableCitation>
|
||
,
|
||
<figureCitation id="1353028F831DFFFFFC6E3BDE15F5FF06" box="[914,997,196,218]" captionStart="Figure 14" captionStartId="22.[142,220,909,928]" captionTargetBox="[367,1196,198,873]" captionTargetId="figure-46@22.[366,1198,196,877]" captionTargetPageId="22" captionText="Figure 14. Snout-vent length of males in our samples of continental Sahara Acanthodactylus dumerili and A. scutellatus audouini." figureDoi="http://doi.org/10.5281/zenodo.5437183" httpUri="https://zenodo.org/record/5437183/files/figure.png" pageId="21" pageNumber="138">Figs 14</figureCitation>
|
||
and
|
||
<figureCitation id="1353028F831DFFFFFBE03BDE1229FF05" box="[1052,1081,196,218]" captionStart="Figure 15" captionStartId="22.[142,220,1854,1873]" captionTargetBox="[368,1195,1148,1819]" captionTargetId="figure-68@22.[366,1198,1146,1822]" captionTargetPageId="22" captionText="Figure 15. Snout-vent length of females in our samples of continental Sahara Acanthodactylus dumerili and A. scutellatus audouini." figureDoi="http://doi.org/10.5281/zenodo.5437185" httpUri="https://zenodo.org/record/5437185/files/figure.png" pageId="21" pageNumber="138">15</figureCitation>
|
||
). The subocular in contact with three or (rarely) four supralabials in 99% of the individuals (
|
||
<tableCitation id="C6EA2BB1831DFFFFFC6A3A1B15FAFEC8" box="[918,1002,257,279]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="21" pageNumber="138">Table 3</tableCitation>
|
||
) distinguishes
|
||
<emphasis id="B91CC218831DFFFFFB613A1B132BFEC9" box="[1181,1339,257,279]" italics="true" pageId="21" pageNumber="138">A. s. audouini</emphasis>
|
||
from
|
||
<taxonomicName id="4C686589831DFFFFFA7F3A1B1590FEE9" authorityName="Gunther" authorityYear="1903" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="aureus">
|
||
<emphasis id="B91CC218831DFFFFFA7F3A1B1590FEE9" italics="true" pageId="21" pageNumber="138">A. aureus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C686589831DFFFFFC463A3A1245FEEA" authorityName="Geniez & Foucart" authorityYear="1995" box="[954,1109,288,309]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="taghitensis">
|
||
<emphasis id="B91CC218831DFFFFFC463A3A1245FEEA" box="[954,1109,288,309]" italics="true" pageId="21" pageNumber="138">A. taghitensis</emphasis>
|
||
</taxonomicName>
|
||
. Dorsal scales usually small and rather numerous (in the Sahara, range: 39–69, mean: 53.4;
|
||
<tableCitation id="C6EA2BB1831DFFFFFC393A47120AFEAD" box="[965,1050,349,371]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="21" pageNumber="138">Table 2</tableCitation>
|
||
, see remarks below under ‘Geographical variation’), nearly equal in size between the dorsum and the flanks, and moderately to strongly carinate (codes 4 or
|
||
<quantity id="4C90B3EF831DFFFFFBDD3AA3125FFE10" box="[1057,1103,441,463]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="138" unit="in" value="5.0">5 in</quantity>
|
||
86% of our sample, no individual reaches code 6;
|
||
<tableCitation id="C6EA2BB1831DFFFFFBAD3AC212B5FE32" box="[1105,1189,472,494]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="21" pageNumber="138">Table 3</tableCitation>
|
||
, see
|
||
<figureCitation id="1353028F831DFFFFFB173AC2132AFE31" box="[1259,1338,472,494]" captionStart="Figure 16" captionStartId="23.[155,233,1091,1110]" captionTargetBox="[154,1434,199,1058]" captionTargetId="figure-466@23.[153,1435,199,1058]" captionTargetPageId="23" captionText="Figure 16. (a) Acanthodactylus scutellatus audouini, male, paralectotype of Acanthodactylus scutellatus var. audouini, BMNH 97.10.28.316, Wadi Halfa (Egypt), close-up of dorsal scales. (b) A. senegalensis, male, EPHE Mch1, coast north of Dakar (Senegal), close-up of dorsal scales. (c) A. dumerili, male, MNHN 1997.3764, Blaouakh (Mauritania), close-up of dorsal scales. (d) A. longipes, male, EPHE AF4, Akreïdil, 50 km NE of Nouakchott (Mauritania), close-up of dorsal scales." figureDoi="http://doi.org/10.5281/zenodo.5437187" httpUri="https://zenodo.org/record/5437187/files/figure.png" pageId="21" pageNumber="138">Fig. 16</figureCitation>
|
||
). Fewer than two rows of supraciliary granules in 70% of the specimens, two rows in 28%, more than two rows in only 2% (
|
||
<tableCitation id="C6EA2BB1831DFFFFFC54392E15EDFD95" box="[936,1021,564,586]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="21" pageNumber="138">Table 3</tableCitation>
|
||
). Generally 13 or 14 longitudinal rows of ventral scales (in 83% of the specimens, range: 12–16, mean: 13.9;
|
||
<tableCitation id="C6EA2BB1831DFFFFFBDB396B126BFD58" box="[1063,1147,625,647]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="21" pageNumber="138">Table 2</tableCitation>
|
||
). Femoral pores rather numerous (range: 16–26, mean: 21.1;
|
||
<tableCitation id="C6EA2BB1831DFFFFFB13398A1354FD79" box="[1263,1348,656,678]" captionStart="Table 2" captionStartId="5.[155,219,196,215]" captionText="Table 2. Means, standard deviations (in parentheses) and ranges of the quantitative variables used in this study. See Table 1 for variable definitions" pageId="21" pageNumber="138">Table 2</tableCitation>
|
||
). Adult males with a black, highly contrasting, dorsal reticulation. Females with isolated black spots at regular spaces on the back. Pale dorsal spots often lacking in adults, which have a dorsal coloration made of only two colours; most individuals have black spots on their pileus. For separation of
|
||
<emphasis id="B91CC218831DFFFFFB793852133CFC82" box="[1157,1324,840,862]" italics="true" pageId="21" pageNumber="138">A. s. audouini</emphasis>
|
||
from
|
||
<taxonomicName id="4C686589831DFFFFFA7F38521580FCA4" authorityName=", Mellado & Olmedo" authorityYear="1990" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="B91CC218831DFFFFFA7F38521580FCA4" italics="true" pageId="21" pageNumber="138">A. longipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C686589831DFFFFFC12387D128EFCA4" authorityName="Chabanaud" authorityYear="1918" box="[1006,1182,870,892]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="senegalensis">
|
||
<emphasis id="B91CC218831DFFFFFC12387D128EFCA4" box="[1006,1182,870,892]" italics="true" pageId="21" pageNumber="138">A. senegalensis</emphasis>
|
||
</taxonomicName>
|
||
, see these species.
|
||
<taxonomicName id="4C686589831FFFFDFF673FEF1442FAD5" box="[155,594,1269,1290]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC218831FFFFDFF673FEF1442FAD5" box="[155,594,1269,1290]" italics="true" pageId="23" pageNumber="140">Acanthodactylus scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the closely similar
|
||
<taxonomicName id="4C686589831FFFFDFDD73E0E14A2FAF6" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[555,690,1300,1321]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFDD73E0E14A2FAF6" box="[555,690,1300,1321]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
by its larger maximum size and by a different pileus coloration. In
|
||
<taxonomicName id="4C686589831FFFFDFF0F3E4B1763FAB9" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[243,371,1361,1382]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFF0F3E4B1763FAB9" box="[243,371,1361,1382]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
, as in the remaining species within the
|
||
<taxonomicName id="4C686589831FFFFDFF373E6A1755FA5A" baseAuthorityName="Audouin" baseAuthorityYear="1827" box="[203,325,1392,1413]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC218831FFFFDFF373E6A1755FA5A" box="[203,325,1392,1413]" italics="true" pageId="23" pageNumber="140">scutellatus</emphasis>
|
||
</taxonomicName>
|
||
group, the pileus is either uniformly pale or marked with darker vermiculations or small dots, but not with distinct and well-individualized large black spots as in
|
||
<emphasis id="B91CC218831FFFFDFE4E3ED61441FA3E" box="[434,593,1484,1505]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
. In addition, a higher proportion of adults
|
||
<emphasis id="B91CC218831FFFFDFE273EF11469FA20" box="[475,633,1514,1536]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
have a dorsal coloration consisting of a dark pattern on a uniform background (two colours only on the dorsum, COUL code 4, in 27% of the specimens;
|
||
<tableCitation id="C6EA2BB1831FFFFDFDA53D5C14BDF983" box="[601,685,1606,1628]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="23" pageNumber="140">Table 3</tableCitation>
|
||
). When other species within the scutellatus group have only two colours on the dorsum, it is usually light spots on a darker ground colour (code 2,
|
||
<tableCitation id="C6EA2BB1831FFFFDFDF63DB8144EF967" box="[522,606,1698,1720]" captionStart="Table 3" captionStartId="6.[142,206,196,215]" captionText="Table 3. Frequency of occurrence of each class of the qualitative and semiquantitative variables except SO1, SO4 and GPRO. Variable definitions and coding as in Table 1. Bottom line indicates sample size for each character" pageId="23" pageNumber="140">Table 3</tableCitation>
|
||
). According to
|
||
<bibRefCitation id="EFF963FB831FFFFDFF673DDB1725F908" author="Nouira S" box="[155,309,1729,1751]" pageId="23" pageNumber="140" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra (1996</bibRefCitation>
|
||
; p. 246),
|
||
<emphasis id="B91CC218831FFFFDFE5E3DDB1450F909" box="[418,576,1729,1751]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
and
|
||
<taxonomicName id="4C686589831FFFFDFD833DDB1513F909" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[639,771,1729,1750]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFD833DDB1513F909" box="[639,771,1729,1750]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
also differ in the fragmentation of the cephalic plates,
|
||
<emphasis id="B91CC218831FFFFDFF673DE51727F8CC" box="[155,311,1790,1812]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
having the first supraocular (SO1) usually separated from the second supraocular (SO2) by a row of proximal granules and a strongly fragmented fourth supraocular (SO4). We checked the validity of these proposed differences on 116
|
||
<emphasis id="B91CC218831FFFFDFB443FEF1344FAD5" box="[1208,1364,1269,1291]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
(most of them from
|
||
<collectingCountry id="F37F5E9A831FFFFDFC253E0E1221FAF5" box="[985,1073,1300,1322]" name="Tunisia" pageId="23" pageNumber="140">Tunisia</collectingCountry>
|
||
) and 298
|
||
<taxonomicName id="4C686589831FFFFDFB513E0E1323FAF6" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[1197,1331,1300,1321]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFB513E0E1323FAF6" box="[1197,1331,1300,1321]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<emphasis id="B91CC218831FFFFDFA943E0E158AFA98" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
, 48% of the individuals have SO1 and SO2 partly or totally separated (including 18% with completely separated SO1 and SO4;
|
||
<tableCitation id="C6EA2BB1831FFFFDFB4D3E6A1316FA59" box="[1201,1286,1392,1414]" captionStart="Table 4" captionStartId="7.[155,219,196,215]" captionText="Table 4. Frequency of occurrence of each class of the variables SO1, SO4 and GPRO in a subsample of Acanthodactylus dumerili (mainly Tunisia) and A. scutellatus and sample size of each species for each variable (n)" pageId="23" pageNumber="140">Table 4</tableCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C686589831FFFFDFCCF3E9515A7FA7C" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[819,951,1422,1444]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFCCF3E9515A7FA7C" box="[819,951,1422,1444]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
only 7% of the specimens have SO1 and SO2 partly separated (none having SO1 and SO4 completely separated;
|
||
<tableCitation id="C6EA2BB1831FFFFDFBF73ED61271FA3D" box="[1035,1121,1484,1506]" captionStart="Table 4" captionStartId="7.[155,219,196,215]" captionText="Table 4. Frequency of occurrence of each class of the variables SO1, SO4 and GPRO in a subsample of Acanthodactylus dumerili (mainly Tunisia) and A. scutellatus and sample size of each species for each variable (n)" pageId="23" pageNumber="140">Table 4</tableCitation>
|
||
). In
|
||
<emphasis id="B91CC218831FFFFDFB663ED6132CFA3E" box="[1178,1340,1484,1505]" italics="true" pageId="23" pageNumber="140">A. s. audouini</emphasis>
|
||
, 30% of the specimens have an entire SO4, 33% have a partially fragmented SO4 and 37% have a completely fragmented SO4. In
|
||
<taxonomicName id="4C686589831FFFFDFBE63D32128BF9E2" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[1050,1179,1576,1597]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFBE63D32128BF9E2" box="[1050,1179,1576,1597]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
, 49% of the specimens have an entire SO4, 38% a partially fragmented one, and only 14% a completely fragmented SO4 (
|
||
<tableCitation id="C6EA2BB1831FFFFDFACA3D7F139EF9A4" box="[1334,1422,1637,1659]" captionStart="Table 4" captionStartId="7.[155,219,196,215]" captionText="Table 4. Frequency of occurrence of each class of the variables SO1, SO4 and GPRO in a subsample of Acanthodactylus dumerili (mainly Tunisia) and A. scutellatus and sample size of each species for each variable (n)" pageId="23" pageNumber="140">Table 4</tableCitation>
|
||
). The differences described by
|
||
<bibRefCitation id="EFF963FB831FFFFDFB703D9E133EF945" author="Nouira S" box="[1164,1326,1668,1690]" pageId="23" pageNumber="140" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra (1996)</bibRefCitation>
|
||
are thus real but of limited use when trying to identify single specimens.
|
||
</paragraph>
|
||
<caption id="DF174E82831DFFFFFF673C381541F8B0" ID-DOI="http://doi.org/10.5281/zenodo.5437181" ID-Zenodo-Dep="5437181" httpUri="https://zenodo.org/record/5437181/files/figure.png" pageId="21" pageNumber="138" startId="21.[155,233,1826,1845]" targetBox="[157,1429,981,1786]" targetPageId="21">
|
||
<paragraph id="8BD71E0A831DFFFFFF673C381541F8B0" blockId="21.[155,1435,1826,1904]" pageId="21" pageNumber="138">
|
||
<emphasis id="B91CC218831DFFFFFF673C381700F8EA" bold="true" box="[155,272,1826,1845]" pageId="21" pageNumber="138">Figure 13.</emphasis>
|
||
Geographic distribution of
|
||
<taxonomicName id="4C686589831DFFFFFDCF3C381546F8EA" baseAuthorityName="Audouin" baseAuthorityYear="1827" box="[563,854,1826,1845]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC218831DFFFFFDCF3C381546F8EA" box="[563,854,1826,1845]" italics="true" pageId="21" pageNumber="138">Acanthodactylus scutellatus</emphasis>
|
||
</taxonomicName>
|
||
in the Sahara (thus excluding part of the species range, see text). Dotted lines indicate the approximate limits of the distribution of
|
||
<taxonomicName id="4C686589831DFFFFFC473C251222F88D" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[955,1074,1855,1874]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="138" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831DFFFFFC473C251222F88D" box="[955,1074,1855,1874]" italics="true" pageId="21" pageNumber="138">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
. Data from Salvador (1982),
|
||
<bibRefCitation id="EFF963FB831DFFFFFA9F3C25173BF8B0" author="Baha El Din SM" pageId="21" pageNumber="138" pagination="18 - 27" refId="ref21524" refString="Baha El Din SM. 1994. A contribution to the herpetology of Sinai. British Herpetological Society Bulletin 48: 18 - 27." type="journal article" year="1994">Baha El Din (1994)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF963FB831DFFFFFECB3C471427F8B0" author="Joger U & Lambert MRK" box="[311,567,1884,1904]" pageId="21" pageNumber="138" pagination="21 - 51" refId="ref22405" refString="Joger U, Lambert MRK. 1996. Analysis of the herpetofauna of the Republic of Mali, I. Annotated inventory, with description of a new Uromastyx (Sauria: Agamidae). Journal of African Zoology 110: 21 - 51." type="journal article" year="1996">Joger & Lambert (1996)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF963FB831DFFFFFDBF3C4714C7F8B0" author="Nouira S" box="[579,727,1884,1904]" pageId="21" pageNumber="138" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra (1996)</bibRefCitation>
|
||
, this study.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF174E82831EFFFCFF72389716E3FC61" ID-DOI="http://doi.org/10.5281/zenodo.5437183" ID-Zenodo-Dep="5437183" httpUri="https://zenodo.org/record/5437183/files/figure.png" pageId="22" pageNumber="139" startId="22.[142,220,909,928]" targetBox="[367,1196,198,873]" targetPageId="22">
|
||
<paragraph id="8BD71E0A831EFFFCFF72389716E3FC61" blockId="22.[142,1422,909,958]" pageId="22" pageNumber="139">
|
||
<emphasis id="B91CC218831EFFFCFF7238971713FC7F" bold="true" box="[142,259,909,928]" pageId="22" pageNumber="139">Figure 14.</emphasis>
|
||
Snout-vent length of males in our samples of continental Sahara
|
||
<taxonomicName id="4C686589831EFFFCFC42389712DDFC7F" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[958,1229,909,928]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="139" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831EFFFCFC42389712DDFC7F" box="[958,1229,909,928]" italics="true" pageId="22" pageNumber="139">Acanthodactylus dumerili</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C686589831EFFFCFAFE389416FDFC61" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="139" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC218831EFFFCFAFE389416FDFC61" italics="true" pageId="22" pageNumber="139">A. scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF174E82831EFFFCFF723C2416E3F8B0" ID-DOI="http://doi.org/10.5281/zenodo.5437185" ID-Zenodo-Dep="5437185" httpUri="https://zenodo.org/record/5437185/files/figure.png" pageId="22" pageNumber="139" startId="22.[142,220,1854,1873]" targetBox="[368,1195,1148,1819]" targetPageId="22">
|
||
<paragraph id="8BD71E0A831EFFFCFF723C2416E3F8B0" blockId="22.[142,1422,1854,1903]" pageId="22" pageNumber="139">
|
||
<emphasis id="B91CC218831EFFFCFF723C241713F88E" bold="true" box="[142,259,1854,1873]" pageId="22" pageNumber="139">Figure 15.</emphasis>
|
||
Snout-vent length of females in our samples of continental Sahara
|
||
<taxonomicName id="4C686589831EFFFCFC3F3C2512C1F88D" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[963,1233,1855,1874]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="139" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831EFFFCFC3F3C2512C1F88D" box="[963,1233,1855,1874]" italics="true" pageId="22" pageNumber="139">Acanthodactylus dumerili</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C686589831EFFFCFAFF3C2516FDF8B0" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="139" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC218831EFFFCFAFF3C2516FDF8B0" italics="true" pageId="22" pageNumber="139">A. scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF174E82831FFFFDFF673F59136CFB71" ID-DOI="http://doi.org/10.5281/zenodo.5437187" ID-Zenodo-Dep="5437187" httpUri="https://zenodo.org/record/5437187/files/figure.png" pageId="23" pageNumber="140" startId="23.[155,233,1091,1110]" targetBox="[154,1434,199,1058]" targetPageId="23">
|
||
<paragraph id="8BD71E0A831FFFFDFF673F59136CFB71" blockId="23.[155,1435,1091,1198]" pageId="23" pageNumber="140">
|
||
<emphasis id="B91CC218831FFFFDFF673F591700FB89" bold="true" box="[155,272,1091,1110]" pageId="23" pageNumber="140">Figure 16.</emphasis>
|
||
(a)
|
||
<taxonomicName id="4C686589831FFFFDFEBD3F5914C0FB89" box="[321,720,1091,1110]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC218831FFFFDFEBD3F5914C0FB89" box="[321,720,1091,1110]" italics="true" pageId="23" pageNumber="140">Acanthodactylus scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
, male, paralectotype of
|
||
<taxonomicName id="4C686589831FFFFDFC293F5916F6FBAB" authority=", BMNH" authorityName="BMNH" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="variety" species="scutellatus" variety="audouini">
|
||
<emphasis id="B91CC218831FFFFDFC293F5912ECFB89" box="[981,1276,1091,1110]" italics="true" pageId="23" pageNumber="140">Acanthodactylus scutellatus</emphasis>
|
||
var.
|
||
<emphasis id="B91CC218831FFFFDFACA3F591385FB89" box="[1334,1429,1091,1110]" italics="true" pageId="23" pageNumber="140">audouini</emphasis>
|
||
, BMNH
|
||
</taxonomicName>
|
||
97.10.28.316, Wadi Halfa (Egypt), close-up of dorsal scales. (b)
|
||
<taxonomicName id="4C686589831FFFFDFC763F7A1238FBAC" authorityName="Chabanaud" authorityYear="1918" box="[906,1064,1120,1139]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="senegalensis">
|
||
<emphasis id="B91CC218831FFFFDFC763F7A1238FBAC" box="[906,1064,1120,1139]" italics="true" pageId="23" pageNumber="140">A. senegalensis</emphasis>
|
||
</taxonomicName>
|
||
, male, EPHE Mch1, coast north of Dakar (Senegal), close-up of dorsal scales. (c)
|
||
<taxonomicName id="4C686589831FFFFDFD883F6414FBFB4F" baseAuthorityName="Milne Edwards" baseAuthorityYear="1829" box="[628,747,1149,1169]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="dumerili">
|
||
<emphasis id="B91CC218831FFFFDFD883F6414FBFB4F" box="[628,747,1149,1169]" italics="true" pageId="23" pageNumber="140">A. dumerili</emphasis>
|
||
</taxonomicName>
|
||
, male, MNHN 1997.3764, Blaouakh (Mauritania), close-up of dorsal scales. (d)
|
||
<taxonomicName id="4C686589831FFFFDFED33F8117B2FB71" authorityName=", Mellado & Olmedo" authorityYear="1990" box="[303,418,1179,1198]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="B91CC218831FFFFDFED33F8117B2FB71" box="[303,418,1179,1198]" italics="true" pageId="23" pageNumber="140">A. longipes</emphasis>
|
||
</taxonomicName>
|
||
, male, EPHE AF4, Akreïdil, 50 km NE of Nouakchott (Mauritania), close-up of dorsal scales.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD71E0A831FFFF2FCB03DE41403FE4D" blockId="23.[819,1435,1790,1904]" lastBlockId="24.[142,758,196,402]" lastPageId="24" lastPageNumber="141" pageId="23" pageNumber="140">
|
||
<emphasis id="B91CC218831FFFFDFCB03DE41244F8CB" box="[844,1108,1790,1812]" italics="true" pageId="23" pageNumber="140">Geographical variation</emphasis>
|
||
. Three subspecies have been recognized. The subspecies
|
||
<taxonomicName id="4C686589831FFFFDFB883C0712D0F8ED" authorityName="Haas" authorityYear="1957" box="[1140,1216,1821,1842]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="140" phylum="Chordata" rank="species" species="hardyi">
|
||
<emphasis id="B91CC218831FFFFDFB883C0712D0F8ED" box="[1140,1216,1821,1842]" italics="true" pageId="23" pageNumber="140">hardyi</emphasis>
|
||
</taxonomicName>
|
||
is recognized as a valid taxon in most of the recent works on the genus (e.g.
|
||
<bibRefCitation id="EFF963FB831FFFFDFC9B3C411253F8AF" author="Bons J & Girot B" box="[871,1091,1882,1904]" pageId="23" pageNumber="140" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964</bibRefCitation>
|
||
;
|
||
<collectingCountry id="F37F5E9A831FFFFDFBB23C4012A3F8AF" box="[1102,1203,1882,1904]" name="El Salvador" pageId="23" pageNumber="140">Salvador</collectingCountry>
|
||
, 1982;
|
||
<bibRefCitation id="EFF963FB831FFFFDFAFE3C401387F8AF" author="Arnold EN" box="[1282,1431,1882,1904]" pageId="23" pageNumber="140" pagination="385 - 435" refId="ref21343" refString="Arnold EN. 1986. A Key and annotated check list to the lizards and Amphisbaenians of Arabia. In: Buttiker W, Krupp F, eds. Fauna of Saudi Arabia, Vol. 8. Jeddah: Meteorology and Environmental Protection Administration, 385 - 435." type="book chapter" year="1986">Arnold, 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF963FB8310FFF2FF723BDE1735FF05" author="Nouira S" box="[142,293,196,218]" pageId="24" pageNumber="141" refId="ref22685" refString="Nouira S. 1996. Systematique, ecologie et biogeographie evolutive des Lacertidae (Reptilia, Sauria). Importance dans l'herpetofaune tunisienne. DPhil Thesis, University of Tunis." type="book" year="1996">Nouïra, 1996</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFF963FB8310FFF2FECB3BDE142FFF05" author="Harris DJ & Arnold EN" box="[311,575,196,218]" pageId="24" pageNumber="141" pagination="351 - 362" refId="ref22188" refString="Harris DJ, Arnold EN. 2000. Elucidation of the relationships of spiny-footed lizards, Acanthodactylus spp. (Reptilia: Lacertidae) using mitochondrial DNA sequence, with comments on their biogeography and evolution. Journal of Zoology, London 252: 351 - 362." type="journal article" year="2000">Harris & Arnold (2000)</bibRefCitation>
|
||
even propose to give specific status to this taxon, although without justification. In addition,
|
||
<bibRefCitation id="EFF963FB8310FFF2FE733A181466FEC8" author="Bons J & Girot B" box="[399,630,257,279]" pageId="24" pageNumber="141" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot (1964)</bibRefCitation>
|
||
recognized the subspecies
|
||
<emphasis id="B91CC2188310FFF2FEC63A3A17B1FEEA" box="[314,417,288,309]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
, based on an analysis of large series of specimens. We follow their opinion here, based on a preliminary examination of a number of specimens from
|
||
<collectingCountry id="F37F5E9A8310FFF2FEB63A66179EFE4E" box="[330,398,380,401]" name="Egypt" pageId="24" pageNumber="141">Egypt</collectingCountry>
|
||
and
|
||
<collectingCountry id="F37F5E9A8310FFF2FE353A66141FFE4D" box="[457,527,380,402]" name="Israel" pageId="24" pageNumber="141">Israel</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8310FFF2FF5B3AA317D1FE32" blockId="24.[142,757,441,494]" pageId="24" pageNumber="141">
|
||
<emphasis id="B91CC2188310FFF2FF5B3AA31731FE11" box="[167,289,441,462]" italics="true" pageId="24" pageNumber="141">Subspecies</emphasis>
|
||
.
|
||
<taxonomicName id="4C6865898310FFF2FECD3AA314E5FE11" authority="scutellatus" box="[305,757,441,462]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="scutellatus">
|
||
<emphasis id="B91CC2188310FFF2FECD3AA314E5FE11" box="[305,757,441,462]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus scutellatus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F37F5E9A8310FFF2FF513AC216E1FE31" box="[173,241,472,494]" name="Israel" pageId="24" pageNumber="141">Israel</collectingCountry>
|
||
,
|
||
<collectingRegion id="49ACD0E88310FFF2FF023AC2172AFE31" box="[254,314,472,494]" country="Egypt" name="Shamal Sina'" pageId="24" pageNumber="141">Sinai</collectingRegion>
|
||
and
|
||
<collectingCountry id="F37F5E9A8310FFF2FE893AC217ACFE32" box="[373,444,472,493]" name="Egypt" pageId="24" pageNumber="141">Egypt</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3724D818310FFF2FF723929146CFB70" pageId="24" pageNumber="141" type="reference_group">
|
||
<paragraph id="8BD71E0A8310FFF2FF72392916FDFBAD" blockId="24.[142,759,563,1199]" pageId="24" pageNumber="141">
|
||
<taxonomicName id="4C6865898310FFF2FF72392914FBFD96" authority="Boulenger, 1918" authorityName="Boulenger" authorityYear="1918" box="[142,747,563,585]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC2188310FFF2FF7239291421FD97" box="[142,561,563,584]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus audouini</emphasis>
|
||
<bibRefCitation id="EFF963FB8310FFF2FDC5392914FBFD96" author="Boulenger GA" box="[569,747,563,585]" pageId="24" pageNumber="141" pagination="143 - 155" refId="ref21726" refString="Boulenger GA. 1918. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Societe Zoologique de France 43: 143 - 155." type="journal article" year="1918">Boulenger, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C6865898310FFF2FF72394817DFFDB8" baseAuthorityName="Audouin" baseAuthorityYear="1827" box="[142,463,594,615]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC2188310FFF2FF72394817DFFDB8" box="[142,463,594,615]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus</emphasis>
|
||
</taxonomicName>
|
||
var. A
|
||
<emphasis id="B91CC2188310FFF2FDE239481467FDB8" box="[542,631,594,615]" italics="true" pageId="24" pageNumber="141">udouini</emphasis>
|
||
<bibRefCitation id="EFF963FB8310FFF2FD7D394816C8FD58" author="Boulenger GA" pageId="24" pageNumber="141" pagination="143 - 155" refId="ref21726" refString="Boulenger GA. 1918. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Societe Zoologique de France 43: 143 - 155." type="journal article" year="1918">Boulenger (1918)</bibRefCitation>
|
||
: 154. Name-bearing type: Boulenger did not refer to any precise specimen in his original description, but stated that he used specimens in the British Museum, especially specimens collected by F. Lataste. These specimens came from ‘
|
||
<collectingCountry id="F37F5E9A8310FFF2FE1639F6142DFCDE" box="[490,573,748,769]" name="Egypt" pageId="24" pageNumber="141">Egypte</collectingCountry>
|
||
, Nubie, Tripoli, sud de la Tunisie’. The following specimens are probably all
|
||
<typeStatus id="54D3A0A88310FFF2FED738301780FCE0" box="[299,400,810,831]" pageId="24" pageNumber="141" type="syntype">syntypes</typeStatus>
|
||
(see
|
||
<collectingCountry id="F37F5E9A8310FFF2FE253833142EFCE0" box="[473,574,809,831]" name="El Salvador" pageId="24" pageNumber="141">Salvador</collectingCountry>
|
||
, 1982): BMNH 97.10.28.315-319 (Wadi Halfa,
|
||
<collectingCountry id="F37F5E9A8310FFF2FDD33852146BFC81" box="[559,635,840,862]" name="Sudan" pageId="24" pageNumber="141">Sudan</collectingCountry>
|
||
), BMNH 1913.12.30.6-10 (Homs, Tripoli), BMNH 91.5.4.85-91 (Duirat, southern
|
||
<collectingCountry id="F37F5E9A8310FFF2FE8C389F17DAFC44" box="[368,458,901,923]" name="Tunisia" pageId="24" pageNumber="141">Tunisia</collectingCountry>
|
||
), BMNH 1920.1.20.3006 (Wed El Kreil,
|
||
<collectingCountry id="F37F5E9A8310FFF2FEC938BE179FFC65" box="[309,399,932,954]" name="Tunisia" pageId="24" pageNumber="141">Tunisia</collectingCountry>
|
||
). Considering the uncertainties over the limits of the range of
|
||
<emphasis id="B91CC2188310FFF2FE0B38D8144EFC08" box="[503,606,962,983]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
, we feel it is important to select a
|
||
<typeStatus id="54D3A0A88310FFF2FE6838FB17EAFC28" box="[404,506,993,1015]" pageId="24" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
from an area where
|
||
<emphasis id="B91CC2188310FFF2FF723F1A16E5FBCA" box="[142,245,1024,1045]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
(as generally understood) is certainly present. We select the specimen BMNH 1920. 1.20.3006 as
|
||
<typeStatus id="54D3A0A88310FFF2FED23F271787FB8C" box="[302,407,1085,1107]" pageId="24" pageNumber="141" type="lectotype">lectotype</typeStatus>
|
||
. Type locality: Wed El Kreil,
|
||
<collectingCountry id="F37F5E9A8310FFF2FF723F4616F9FBAD" box="[142,233,1116,1138]" name="Tunisia" pageId="24" pageNumber="141">Tunisia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8310FFF2FF5B3F60146CFB70" blockId="24.[142,759,563,1199]" pageId="24" pageNumber="141">
|
||
Distributed in the west of the species range, east to north-eastern
|
||
<collectingCountry id="F37F5E9A8310FFF2FECF3F83176DFB70" box="[307,381,1177,1199]" name="Sudan" pageId="24" pageNumber="141">Sudan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFF963FB8310FFF2FE713F831461FB70" author="Bons J & Girot B" box="[397,625,1177,1199]" pageId="24" pageNumber="141" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3724D818310FFF2FF723FEF1781FAD4" box="[142,401,1269,1291]" pageId="24" pageNumber="141" type="description">
|
||
<paragraph id="8BD71E0A8310FFF2FF723FEF1781FAD4" blockId="24.[142,759,1269,1904]" box="[142,401,1269,1291]" pageId="24" pageNumber="141">
|
||
<emphasis id="B91CC2188310FFF2FF723FEF16ABFAD5" box="[142,187,1269,1290]" italics="true" pageId="24" pageNumber="141">A. s.</emphasis>
|
||
<taxonomicName id="4C6865898310FFF2FF3F3FEF1781FAD4" authority="Haas, 1957" authorityName="Haas" authorityYear="1957" box="[195,401,1269,1291]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="hardyi">
|
||
<emphasis id="B91CC2188310FFF2FF3F3FEF171DFAD5" box="[195,269,1269,1290]" italics="true" pageId="24" pageNumber="141">hardyi</emphasis>
|
||
<bibRefCitation id="EFF963FB8310FFF2FEE93FEF1781FAD4" author="Haas G" box="[277,401,1269,1291]" pageId="24" pageNumber="141" pagination="47 - 86" refId="ref22160" refString="Haas G. 1957. Some amphibians and reptiles from Arabia. Proceedings of the California Academy of Sciences, Fourth Series 29: 47 - 86." type="book chapter" year="1957">Haas, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3724D818310FFF2FF723E0E1797FA1C" pageId="24" pageNumber="141" type="reference_group">
|
||
<paragraph id="8BD71E0A8310FFF2FF723E0E17E8FA59" blockId="24.[142,759,1269,1904]" pageId="24" pageNumber="141">
|
||
<treatmentCitationGroup id="AB7839248310FFF2FF723E0E17E8FA59" pageId="24" pageNumber="141">
|
||
<treatmentCitation id="0AC9381B8310FFF2FF723E0E14E3FAF6" author="Haas G" box="[142,755,1300,1322]" page="72" pageId="24" pageNumber="141" year="1957">
|
||
<taxonomicName id="4C6865898310FFF2FF723E0E14E3FAF6" authority="Haas, 1957: 72" authorityName="Haas" authorityPageNumber="72" authorityYear="1957" box="[142,755,1300,1322]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="hardyi">
|
||
<emphasis id="B91CC2188310FFF2FF723E0E143CFAF6" box="[142,556,1300,1321]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus hardyi</emphasis>
|
||
<materialsCitation id="3B0014578310FFF2FDC73E0E14E3FAF6" ID-GBIF-Occurrence="3341581301" box="[571,755,1300,1322]" county="Hebrew University" location="Saudi Arabia'" municipality="Hirmas Station" pageId="24" pageNumber="141" specimenCount="1" stateProvince="Jerusalem" typeStatus="holotype">
|
||
<bibRefCitation id="EFF963FB8310FFF2FDC73E0E14E3FAF6" author="Haas G" box="[571,755,1300,1322]" pageId="24" pageNumber="141" pagination="47 - 86" refId="ref22160" refString="Haas G. 1957. Some amphibians and reptiles from Arabia. Proceedings of the California Academy of Sciences, Fourth Series 29: 47 - 86." type="book chapter" year="1957">Haas, 1957: 72</bibRefCitation>
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Name-bearing type:
|
||
<typeStatus id="54D3A0A88310FFF2FE873E2817CCFA97" box="[379,476,1330,1352]" pageId="24" pageNumber="141" type="holotype">holotype</typeStatus>
|
||
by original designation:
|
||
<collectingCounty id="62B666868310FFF2FF723E4B177BFAB8" box="[142,363,1361,1383]" pageId="24" pageNumber="141">Hebrew University</collectingCounty>
|
||
of
|
||
<collectingRegion id="49ACD0E88310FFF2FE6F3E4B141AFAB8" box="[403,522,1361,1383]" country="Israel" name="Jerusalem" pageId="24" pageNumber="141">Jerusalem</collectingRegion>
|
||
2682. Type locality: ‘
|
||
<collectingMunicipality id="6BB384708310FFF2FF6F3E6A1758FA59" box="[147,328,1392,1414]" pageId="24" pageNumber="141">Hirmas Station</collectingMunicipality>
|
||
,
|
||
<location id="8EB748D18310FFF2FEA83E6A17E4FA59" LSID="urn:lsid:plazi:treatment:03C1AF1C831CFFF2FCCC3D071265FE1F:8EB748D18310FFF2FEA83E6A17E4FA59" box="[340,500,1392,1414]" county="Hebrew University" municipality="Hirmas Station" name="Saudi Arabia'" pageId="24" pageNumber="141" stateProvince="Jerusalem">Saudi Arabia’</location>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8310FFF2FF5B3E941797FA1C" blockId="24.[142,759,1269,1904]" pageId="24" pageNumber="141">
|
||
Distributed in Northern
|
||
<collectingCountry id="F37F5E9A8310FFF2FE3F3E94144DFA7B" box="[451,605,1422,1444]" name="Saudi Arabia" pageId="24" pageNumber="141">Saudi Arabia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F37F5E9A8310FFF2FD953E9414AFFA7B" box="[617,703,1422,1444]" name="Kuwait" pageId="24" pageNumber="141">Kuwait</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F37F5E9A8310FFF2FF723EB416AEFA1C" box="[142,190,1454,1475]" name="Iraq" pageId="24" pageNumber="141">Iraq</collectingCountry>
|
||
(
|
||
<collectingCountry id="F37F5E9A8310FFF2FF333EB71725FA1C" box="[207,309,1453,1475]" name="El Salvador" pageId="24" pageNumber="141">Salvador</collectingCountry>
|
||
, 1982).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3724D818310FFF2FF5B3ED61265FE1F" pageId="24" pageNumber="141" type="description">
|
||
<paragraph id="8BD71E0A8310FFF2FF5B3ED6176FF8CB" blockId="24.[142,759,1269,1904]" pageId="24" pageNumber="141">
|
||
We retain for the time being the subspecies
|
||
<emphasis id="B91CC2188310FFF2FF723EF016E5FA20" box="[142,245,1514,1535]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
for the Saharan specimens, characterized by a lower number of more strongly keeled dorsal scales (73 on average in
|
||
<taxonomicName id="4C6865898310FFF2FE8E3D3214E7F9E1" authority="(Bons & Girot, 1964)" baseAuthorityName="Bons & Girot" baseAuthorityYear="1964" box="[370,759,1576,1598]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC2188310FFF2FE8E3D3217FCF9E2" box="[370,492,1576,1597]" italics="true" pageId="24" pageNumber="141">scutellatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF963FB8310FFF2FDFD3D3214FFF9E1" author="Bons J & Girot B" box="[513,751,1576,1598]" pageId="24" pageNumber="141" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
against 53 on average, and not more than
|
||
<quantity id="4C90B3EF8310FFF2FD483D5C14E6F983" box="[692,758,1606,1628]" metricMagnitude="0" metricUnit="m" metricValue="1.7271999999999998" pageId="24" pageNumber="141" unit="in" value="68.0">68 in</quantity>
|
||
<emphasis id="B91CC2188310FFF2FF723D7F16E5F9A5" box="[142,245,1637,1658]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
[own data]) and a smaller size (
|
||
<bibRefCitation id="EFF963FB8310FFF2FD873D7F16D7F946" author="Boulenger GA" pageId="24" pageNumber="141" refId="ref21752" refString="Boulenger GA. 1921. Monograph of the Lacertidae, II. London: British Museum (Natural History)." type="book" year="1921">Boulenger, 1921</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF963FB8310FFF2FF283D9E17A7F945" author="Bons J & Girot B" box="[212,439,1668,1690]" pageId="24" pageNumber="141" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot, 1964</bibRefCitation>
|
||
). A revision of the material included in
|
||
<taxonomicName id="4C6865898310FFF2FED43DB81460F968" baseAuthorityName="Bons & Girot" baseAuthorityYear="1964" box="[296,624,1698,1719]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC2188310FFF2FED43DB81460F968" box="[296,624,1698,1719]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus</emphasis>
|
||
</taxonomicName>
|
||
is needed before any firm conclusion can be drawn. The limits of the range of
|
||
<emphasis id="B91CC2188310FFF2FEE33DFA1728F92A" box="[287,312,1760,1781]" italics="true" pageId="24" pageNumber="141">A.</emphasis>
|
||
s.
|
||
<taxonomicName id="4C6865898310FFF2FEA43DFA17C2F92A" baseAuthorityName="Bons & Girot" baseAuthorityYear="1964" box="[344,466,1760,1781]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="scutellatus">
|
||
<emphasis id="B91CC2188310FFF2FEA43DFA17C2F92A" box="[344,466,1760,1781]" italics="true" pageId="24" pageNumber="141">scutellatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91CC2188310FFF2FDF73DFA1434F92A" box="[523,548,1760,1781]" italics="true" pageId="24" pageNumber="141">A.</emphasis>
|
||
s.
|
||
<emphasis id="B91CC2188310FFF2FDB83DFA14BBF92A" box="[580,683,1760,1781]" italics="true" pageId="24" pageNumber="141">audouini</emphasis>
|
||
follow
|
||
<bibRefCitation id="EFF963FB8310FFF2FF723DE5176BF8CB" author="Bons J & Girot B" box="[142,379,1790,1812]" pageId="24" pageNumber="141" pagination="311 - 336" refId="ref21689" refString="Bons J, Girot B. 1964. Revision de l'espece Acanthodactylus scutellatus (Lacertide-saurien). Bulletin de la Societe des Sciences naturelles et physique du Maroc 42 (1962): 311 - 336." type="journal article" year="1964">Bons & Girot (1964)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8310FFF2FF5B3C071369FEC8" blockId="24.[142,759,1269,1904]" lastBlockId="24.[806,1422,196,279]" pageId="24" pageNumber="141">
|
||
There is a slight morphological variation among populations of
|
||
<taxonomicName id="4C6865898310FFF2FEBC3C2614E5F88E" authorityName="Boulenger" authorityYear="1918" box="[320,757,1852,1873]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="subSpecies" species="scutellatus" subSpecies="audouini">
|
||
<emphasis id="B91CC2188310FFF2FEBC3C2614E5F88E" box="[320,757,1852,1873]" italics="true" pageId="24" pageNumber="141">Acanthodactylus scutellatus audouini</emphasis>
|
||
</taxonomicName>
|
||
in the Sahara. In
|
||
<collectingCountry id="F37F5E9A8310FFF2FEAF3C4017BCF8AF" box="[339,428,1882,1904]" name="Tunisia" pageId="24" pageNumber="141">Tunisia</collectingCountry>
|
||
, most of the individuals have a grey throat and a reddish tail, whereas this coloration is exhibited, more or less markedly, by only a small number of individuals from southern Sahara.
|
||
</paragraph>
|
||
<paragraph id="8BD71E0A8310FFF2FCC33A351265FE1F" blockId="24.[806,1422,303,448]" pageId="24" pageNumber="141">
|
||
<emphasis id="B91CC2188310FFF2FCC33A3515B4FE9B" box="[831,932,303,324]" italics="true" pageId="24" pageNumber="141">Remarks</emphasis>
|
||
. Victor Audouin published the description of
|
||
<taxonomicName id="4C6865898310FFF2FCB83A551201FEBC" authorityName="Audouin" authorityYear="1827" box="[836,1041,334,356]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="141" phylum="Chordata" rank="species" species="scutellata">
|
||
<emphasis id="B91CC2188310FFF2FCB83A551201FEBC" box="[836,1041,334,356]" italics="true" pageId="24" pageNumber="141">Lacerta scutellata</emphasis>
|
||
</taxonomicName>
|
||
in 1827 and not in 1809 (
|
||
<collectingCountry id="F37F5E9A8310FFF2FAB93A54155BFE5D" name="El Salvador" pageId="24" pageNumber="141">Salvador</collectingCountry>
|
||
, 1982; according to his birth date, V. Audouin was only 12 years old in 1809 [R. Bour, pers. com.]) or 1829 (
|
||
<bibRefCitation id="EFF963FB8310FFF2FCD23AB015D4FE60" author="Arnold EN" box="[814,964,426,448]" pageId="24" pageNumber="141" pagination="291 - 339" refId="ref21308" refString="Arnold EN. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum (Natural History) Zoology 44: 291 - 339." type="journal article" year="1983">Arnold, 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF963FB8310FFF2FC2C3AB01278FE1F" author="Brygoo ER" box="[976,1128,426,448]" pageId="24" pageNumber="141" pagination="3 - 57" refId="ref21773" refString="Brygoo ER. 1988. Les types de Lacertides (Reptiles, Sauriens) du Museum national d'Histoire naturelle. Catalogue critique. Bulletin du Museum National d'Histoire Naturelle, Section A, Zoologie, Biologie et Ecologie Animales (Supplement) 10: 3 - 57." type="journal article" year="1988">Brygoo, 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |