189 lines
25 KiB
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189 lines
25 KiB
XML
<document id="AB05910D042DC9FB07BA2CF475B3F6F8" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. & Herbert, D. G." docDate="2013" docId="03BC1F3CE539D90507F8F9DB4489B596" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Clavator griffithsjonesi Emberton 1999" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="16" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CD2AC564584D02555868D1079B3968BD">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:name id="05240484F5E07A3DD9663A10579E3FA3" type="personal">
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<mods:namePart id="E15BF1700967E069970CC53E57C832A4">Griffiths, O. L.</mods:namePart>
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<mods:affiliation id="FFA8C70A130358A8E898FE4BD593E6E5">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
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<mods:nameIdentifier id="B9B942BE075471F4141188B96EC0C87D" type="email">owen@bcm.intnet.mu</mods:nameIdentifier>
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<mods:namePart id="397EE59EE6B63075371E133406709107">Herbert, D. G.</mods:namePart>
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<mods:affiliation id="9B7AF99A65762542A6DF721165CECECA">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
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<mods:nameIdentifier id="7E1FB974AFAB090F9FA84BBD960263FC" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
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<mods:title id="828DA535712B4999680988544DF90417">African Invertebrates</mods:title>
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<mods:part id="1C29C4E6A3A767FE471BE496330E7AE4">
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<mods:date id="8F88CADD20E01982EA84DCFA621F479E">2013</mods:date>
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<mods:number id="8AE75603A800505B3C51A21041880741">2013-02-13</mods:number>
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<mods:number id="B664509B48DC80FB106E03D270377897">54</mods:number>
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<treatment id="03BC1F3CE539D90507F8F9DB4489B596" ID-DOI="http://doi.org/10.5281/zenodo.7670256" ID-GBIF-Taxon="209341521" ID-Zenodo-Dep="7670256" LSID="urn:lsid:plazi:treatment:03BC1F3CE539D90507F8F9DB4489B596" httpUri="http://treatment.plazi.org/id/03BC1F3CE539D90507F8F9DB4489B596" lastPageId="18" lastPageNumber="18" pageId="16" pageNumber="16">
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<subSubSection id="C30FFDA1E539D90707F8F9DB4554B127" box="[437,885,1610,1637]" pageId="16" pageNumber="16" type="nomenclature">
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<paragraph id="8BAAAE2AE539D90707F8F9DB4554B127" blockId="16.[434,889,1578,1637]" box="[437,885,1610,1637]" pageId="16" pageNumber="16">
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<heading id="D0E21946E539D90707F8F9DB4554B127" box="[437,885,1610,1637]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="16" reason="3">
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<taxonomicName id="4C15D5A9E539D90707F8F9DB4554B127" ID-CoL="VV25" ID-ENA="1642598" authority="Emberton, 1999" authorityName="Emberton" authorityYear="1999" box="[437,885,1610,1637]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="16" phylum="Mollusca" rank="species" species="griffithsjonesi">
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<emphasis id="B9617238E539D90707F8F9DB449BB126" box="[437,698,1610,1637]" italics="true" pageId="16" pageNumber="16">Clavator griffithsjonesi</emphasis>
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Emberton, 1999
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FFDA1E539D9070400F9E444FCB1CF" box="[589,733,1650,1676]" pageId="16" pageNumber="16" type="description">
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<paragraph id="8BAAAE2AE539D9070400F9E444FCB1CF" blockId="16.[589,733,1650,1676]" box="[589,733,1650,1676]" pageId="16" pageNumber="16">
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<figureCitation id="132EB2AFE539D9070400F9E44480B1CF" box="[589,673,1650,1676]" captionStart="Fig" captionStartId="17.[144,177,1534,1555]" captionTargetBox="[151,1091,1084,1508]" captionTargetId="figure-323@17.[144,1104,1070,1516]" captionTargetPageId="17" captionText="Fig. 11. Clavator griffithsjonesi Emberton, 1999, genitalia, NMSA L7236: (A) entire reproductive tract (minus ovotestis); (B) penis opened longitudinally to show internal folds and pore at insertion of epiphallus.Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, epp – epiphallus pore, f – flagellum, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670126" httpUri="https://zenodo.org/record/7670126/files/figure.png" pageId="16" pageNumber="16">Figs 11</figureCitation>
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,
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<figureCitation id="132EB2AFE539D90704E0F9E444FCB1CF" box="[685,733,1650,1676]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette & Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette & Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="16" pageNumber="16">25D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FFDA1E538D90606DDFF5B4532B7BA" pageId="17" pageNumber="17" type="reference_group">
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<paragraph id="8BAAAE2AE538D90606DDFF5B4532B7BA" blockId="17.[144,1105,204,1035]" pageId="17" pageNumber="17">
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<taxonomicName id="4C15D5A9E538D90606DDFF5B440AB7A2" ID-CoL="VV25" authority=": Emberton 1999: 93" authorityName=": Emberton" authorityPageNumber="93" authorityYear="1999" box="[144,555,204,225]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="17" phylum="Mollusca" rank="species" species="griffithsjonesi">
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<emphasis id="B9617238E538D90606DDFF5B4747B7A3" box="[144,358,204,225]" italics="true" pageId="17" pageNumber="17">Clavator griffithsjonesi</emphasis>
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:
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<materialsCitation id="3B7DA477E538D9060739FF5A440AB7A2" box="[372,555,204,225]" collectorName="N Bemaraha Reserve" country="Madagascar" latitude="-18.016666" location="Bemaraha Reserve" longLatPrecision="1277" longitude="44.516666" pageId="17" pageNumber="17" specimenCount="1">Emberton 1999: 93</materialsCitation>
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</taxonomicName>
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, figs 13, 14.
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<typeStatus id="54AE1088E538D90604E5FF5A44F4B7A2" box="[680,725,204,225]" pageId="17" pageNumber="17">Type</typeStatus>
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loc.:
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<collectingCountry id="F302EEBAE538D9060543FF5A455EB7A2" box="[782,895,204,225]" name="Madagascar" pageId="17" pageNumber="17">Madagascar</collectingCountry>
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:
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<collectorName id="26E0CBFCE538D90605C7FF5A4271B7A2" box="[906,1104,204,225]" pageId="17" pageNumber="17">
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N
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<location id="8ECAF8F1E538D90605EFFF5A4271B7A2" LSID="urn:lsid:plazi:treatment:03BC1F3CE539D90507F8F9DB4489B596:8ECAF8F1E538D90605EFFF5A4271B7A2" box="[930,1104,204,225]" country="Madagascar" latitude="-18.016666" longLatPrecision="1277" longitude="44.516666" name="Bemaraha Reserve" pageId="17" pageNumber="17">Bemaraha Reserve</location>
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</collectorName>
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(
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<geoCoordinate id="EE21C8EDE538D90606BAFF724712B7BA" box="[247,307,228,249]" degrees="18" direction="south" minutes="1" orientation="latitude" pageId="17" pageNumber="17" precision="925" value="-18.016666">18°1'S</geoCoordinate>
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<geoCoordinate id="EE21C8EDE538D9060774FF7247A0B7BA" box="[313,385,228,249]" degrees="44" direction="east" minutes="31" orientation="longitude" pageId="17" pageNumber="17" precision="925" value="44.516666">44°31'E</geoCoordinate>
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) [= the area now known as Tsingy Beanka].
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FFDA1E538D90606DDFE9E4746B348" pageId="17" pageNumber="17" type="description">
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<paragraph id="8BAAAE2AE538D90606DDFE9E47A0B661" blockId="17.[144,1105,204,1035]" box="[144,385,264,290]" pageId="17" pageNumber="17">Morphological notes:</paragraph>
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<paragraph id="8BAAAE2AE538D90606DDFEA447E1B6C9" blockId="17.[144,1105,204,1035]" pageId="17" pageNumber="17">
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<emphasis id="B9617238E538D90606DDFEA4476FB609" box="[144,334,306,330]" italics="true" pageId="17" pageNumber="17">External features</emphasis>
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(
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<figureCitation id="132EB2AFE538D9060711FEA647E3B609" box="[348,450,304,330]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette & Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette & Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="17" pageNumber="17">Fig. 25D</figureCitation>
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): Head-foot mostly greyish cream with pedal margin tinged orange-brown; dorso-lateral areas of neck yellowish, mid-dorsal region greyish; distal half of optic tentacles grey.
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</paragraph>
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<paragraph id="8BAAAE2AE538D90606DDFE0C47F5B491" blockId="17.[144,1105,204,1035]" pageId="17" pageNumber="17">
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<emphasis id="B9617238E538D90606DDFE0C471EB6F1" box="[144,319,410,434]" italics="true" pageId="17" pageNumber="17">Distal genitalia</emphasis>
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(
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<figureCitation id="132EB2AFE538D9060700FE0E47BEB6F1" box="[333,415,408,434]" captionStart="Fig" captionStartId="17.[144,177,1534,1555]" captionTargetBox="[151,1091,1084,1508]" captionTargetId="figure-323@17.[144,1104,1070,1516]" captionTargetPageId="17" captionText="Fig. 11. Clavator griffithsjonesi Emberton, 1999, genitalia, NMSA L7236: (A) entire reproductive tract (minus ovotestis); (B) penis opened longitudinally to show internal folds and pore at insertion of epiphallus.Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, epp – epiphallus pore, f – flagellum, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670126" httpUri="https://zenodo.org/record/7670126/files/figure.png" pageId="17" pageNumber="17">Fig. 11</figureCitation>
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): Penis slender and cylindrical, with relatively muscular wall; apical
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<emphasis id="B9617238E538D9060691FE2E46D0B691" bold="true" box="[220,241,440,466]" pageId="17" pageNumber="17">⅔</emphasis>
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comprising a blind ending caecum, above insertion of epiphallus; retractor muscle inserted at apex of caecum; interior of penis with approx. 6 well-developed, longitudinal pilasters; epiphallus somewhat recurved at its insertion, opening into a deep cleft in one pilaster, near to base of penis; pilasters dividing and anastomosing somewhat in caecum. Epiphallus extremely long and convoluted, its convolutions and those of vas deferens enveloped in membranous covering of connective tissue and attached to lower spermoviduct and free oviduct; a short flagellum present at junction of vas deferens and epiphallus; interior wall of epiphallus with one large longitudinal pilaster and 6 or 7 smaller longitudinal ridges, these continue into flagellum, evanescing near its tip; vas deferens discharges into epiphallus via a pore situated on large pilaster; vas deferens passing over and down lower sperm-oviduct to its origin at base of prostate; lumen of vas deferens for the most part simple, but broadening just prior to junction with epiphallus, and inner wall with 3 longitudinal folds each with side branches. Genital atrium short and simple; bursa copulatrix duct arising after a short vagina; duct very long, of more or less even width, its wall relatively sturdy; bursa copulatrix situated just below level of albumen gland, bean-shaped with a slightly pointed apex; bursa and its duct brownish in colour. Free oviduct short.
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</paragraph>
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<paragraph id="8BAAAE2AE538D90606DDFC484746B348" blockId="17.[144,1105,204,1035]" pageId="17" pageNumber="17">
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Locality data:
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<taxonomicName id="4C15D5A9E538D906075CFC7647B4B4B0" authorityName=", Antsingimavo" authorityYear="2013" box="[273,405,992,1011]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="17" phylum="Mollusca" rank="species" species="beanka">
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<emphasis id="B9617238E538D906075CFC7647B4B4B0" box="[273,405,992,1011]" italics="true" pageId="17" pageNumber="17">Tsingy Beanka</emphasis>
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</taxonomicName>
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: st’ns 11/06, 12/06, 15/06 (subfossil), 16/06, 18/06, 06/09, 07/09, 08/09 (subfossil and fresh), 09/09.
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</paragraph>
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</subSubSection>
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||
<caption id="DF6AFEA2E538D90606DDFA684786B1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670126" ID-Zenodo-Dep="7670126" httpUri="https://zenodo.org/record/7670126/files/figure.png" pageId="17" pageNumber="17" startId="17.[144,177,1534,1555]" targetBox="[151,1091,1084,1508]" targetPageId="17">
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<paragraph id="8BAAAE2AE538D90606DDFA684786B1C8" blockId="17.[144,1104,1534,1675]" pageId="17" pageNumber="17">
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Fig. 11.
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<taxonomicName id="4C15D5A9E538D9060692F9964473B150" authority="Emberton, 1999" authorityName=": Emberton" authorityYear="1999" box="[223,594,1534,1555]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="17" phylum="Mollusca" rank="species" species="griffithsjonesi">
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<emphasis id="B9617238E538D9060692F9964795B150" box="[223,436,1534,1555]" italics="true" pageId="17" pageNumber="17">Clavator griffithsjonesi</emphasis>
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Emberton, 1999
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</taxonomicName>
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, genitalia, NMSA L7236: (A) entire reproductive tract (minus ovotestis); (B) penis opened longitudinally to show internal folds and pore at insertion of epiphallus. Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, epp – epiphallus pore, f – flagellum, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens.
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</paragraph>
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</caption>
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<subSubSection id="C30FFDA1E53BD90506F8FF5D4250B646" pageId="18" pageNumber="18" type="distribution">
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<paragraph id="8BAAAE2AE53BD90506F8FF5D4250B646" blockId="18.[181,1142,203,726]" pageId="18" pageNumber="18">
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Distribution: Evidently a narrow-range endemic; not found in either the Tsingy de Bemaraha or at Antsingimavo, and currently known to occur only in the
|
||
<taxonomicName id="4C15D5A9E53BD905058EFF7D424CB646" authorityName=", Antsingimavo" authorityYear="2013" box="[963,1133,235,261]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="18" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FFDA1E53BD90506F8FE85478CB62D" pageId="18" pageNumber="18" type="biology_ecology">
|
||
<paragraph id="8BAAAE2AE53BD90506F8FE85478CB62D" blockId="18.[181,1142,203,726]" pageId="18" pageNumber="18">
|
||
Habitat: Mostly found in tall deciduous forest growing on limestone in the southern areas of
|
||
<taxonomicName id="4C15D5A9E53BD905075DFEA54796B60E" authorityName=", Antsingimavo" authorityYear="2013" box="[272,439,307,333]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="18" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
; less common as one moves north. Buries shallowly in leaf-litter during the dry season.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FFDA1E53BD90506F8FEED4489B596" pageId="18" pageNumber="18" type="discussion">
|
||
<paragraph id="8BAAAE2AE53BD90506F8FEED455AB6F6" blockId="18.[181,1142,203,726]" pageId="18" pageNumber="18">
|
||
Remarks: A characteristic element of the
|
||
<taxonomicName id="4C15D5A9E53BD90504DAFEED4560B6D6" authorityName=", Antsingimavo" authorityYear="2013" box="[663,833,379,405]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="18" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
malacofauna and the only species of
|
||
<taxonomicName id="4C15D5A9E53BD9050761FE0847B1B6F5" box="[300,400,414,438]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9617238E53BD9050761FE0847B1B6F5" box="[300,400,414,438]" italics="true" pageId="18" pageNumber="18">Clavator</emphasis>
|
||
</taxonomicName>
|
||
recorded from central western
|
||
<collectingCountry id="F302EEBAE53BD90504A3FE0D4556B6F6" box="[750,887,411,437]" name="Madagascar" pageId="18" pageNumber="18">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE53BD905069DFE2D4489B596" blockId="18.[181,1142,203,726]" pageId="18" pageNumber="18">
|
||
The distal genitalia closely resemble those figured by
|
||
<bibRefCitation id="EF84D3DBE53BD9050554FE2D45F9B696" author="SCHILEYKO, A. A." box="[793,984,443,469]" pageId="18" pageNumber="18" pagination="436 - 564" refId="ref18859" refString="SCHILEYKO, A. A. 1999. Treatise on Recent terrestrial pulmonate molluscs. Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae [sic]. Ruthenica Supplement 2: 436 - 564." type="journal article" year="1999">Schileyko (1999)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C15D5A9E53BD905024FFE284780B6B6" authority="(Shuttleworth, 1852)" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="species" species="eximius">
|
||
<emphasis id="B9617238E53BD905024FFE284254B695" box="[1026,1141,446,470]" italics="true" pageId="18" pageNumber="18">C. eximius</emphasis>
|
||
(Shuttleworth, 1852)
|
||
</taxonomicName>
|
||
. Fischer-Piette
|
||
<emphasis id="B9617238E53BD9050406FE48445AB6B5" box="[587,635,478,502]" italics="true" pageId="18" pageNumber="18">et al</emphasis>
|
||
. (1975) illustrated the genitalia of two further
|
||
<taxonomicName id="4C15D5A9E53BD90506F8FE684756B556" box="[181,375,507,534]" pageId="18" pageNumber="18">
|
||
<emphasis id="B9617238E53BD90506F8FE684738B555" box="[181,281,510,534]" italics="true" pageId="18" pageNumber="18">Clavator</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, of these
|
||
<taxonomicName id="4C15D5A9E53BD90507A1FE68459DB556" authority="(Petit de la Saussaye, 1844)" baseAuthorityName="Petit de la Saussaye" baseAuthorityYear="1844" box="[492,956,507,534]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="species" species="clavator">
|
||
<emphasis id="B9617238E53BD90507A1FE68444CB555" box="[492,621,510,534]" italics="true" pageId="18" pageNumber="18">C. clavator</emphasis>
|
||
(Petit de la Saussaye, 1844)
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C15D5A9E53BD9050211FE684771B575" authorityName=": Emberton" authorityYear="1999" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="species" species="griffithsjonesi">
|
||
<emphasis id="B9617238E53BD9050211FE684771B575" italics="true" pageId="18" pageNumber="18">C. griffithsjonesi</emphasis>
|
||
</taxonomicName>
|
||
, but it has a more slender bursa copulatrix duct. These authors did not differentiate between an epiphallus and vas deferens in
|
||
<taxonomicName id="4C15D5A9E53BD905057FFDA84593B515" baseAuthorityName="Petit de la Saussaye" baseAuthorityYear="1844" box="[818,946,574,598]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="species" species="clavator">
|
||
<emphasis id="B9617238E53BD905057FFDA84593B515" box="[818,946,574,598]" italics="true" pageId="18" pageNumber="18">C. clavator</emphasis>
|
||
</taxonomicName>
|
||
, neither did they report a flagellum at the junction of the epiphallus and vas deferens, but this may have been overlooked.
|
||
<taxonomicName id="4C15D5A9E53BD9050733FDE844E7B5D6" authority="(Deshayes, 1851)" baseAuthorityName="Deshayes" baseAuthorityYear="1851" box="[382,710,635,662]" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="18" phylum="Mollusca" rank="species" species="moreleti">
|
||
<emphasis id="B9617238E53BD9050733FDE847DAB5D5" box="[382,507,638,662]" italics="true" pageId="18" pageNumber="18">C. moreleti</emphasis>
|
||
(Deshayes, 1851)
|
||
</taxonomicName>
|
||
differs considerably in having a much shorter epiphallus/vas deferens that inserts just below the apex of the penis. It also has a shorter, very slender bursa copulatrix duct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |