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<document id="6C95CED161278471ECB42BDE0E6A4D0C" ID-DOI="10.5281/zenodo.214512" ID-GBIF-Dataset="829967ed-0590-49ee-9582-e409743d0502" ID-ISSN="1175-5326" ID-Zenodo-Dep="214512" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460476430745" checkinUser="plazi" docAuthor="Rodríguez, Fernando &amp; Heads, Sam W." docDate="2012" docId="03ABF808277BFFFEFF72CAB1FE26CB64" docLanguage="en" docName="zt03282p068.pdf" docOrigin="Zootaxa 3282" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Scapteriscus cerberus Rodriguez &amp; Heads, sp. nov." docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FF928070277AFFFAFFE5CE2BFFC8C973" masterDocTitle="New mole crickets of the genus Scapteriscus Scudder from Colombia (Orthoptera: Gryllotalpidae; Scapteriscinae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698306805647" updateUser="plazi">
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<mods:title id="A2367AD6A0CF9A0AC29DF4DA1616EA03">New mole crickets of the genus Scapteriscus Scudder from Colombia (Orthoptera: Gryllotalpidae; Scapteriscinae)</mods:title>
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<mods:namePart id="5545231D08AC8B5CDE385643F22EABB7">Rodríguez, Fernando</mods:namePart>
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<mods:namePart id="084417C0617CF7F435F709B3D85272BE">Heads, Sam W.</mods:namePart>
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<mods:date id="1B7C15AAD5FDE9A7A8BD474FAE005256">2012</mods:date>
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<treatment id="03ABF808277BFFFEFF72CAB1FE26CB64" ID-DOI="http://doi.org/10.5281/zenodo.5680706" ID-GBIF-Taxon="119516427" ID-Zenodo-Dep="5680706" LSID="urn:lsid:plazi:treatment:03ABF808277BFFFEFF72CAB1FE26CB64" httpUri="http://treatment.plazi.org/id/03ABF808277BFFFEFF72CAB1FE26CB64" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="C3181A95277BFFFBFF72CAB1FEF9CDA6" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8BBD491E277BFFFBFF72CAB1FCC0CDC7" blockId="1.[151,776,1178,1237]" box="[151,776,1178,1204]" pageId="1" pageNumber="62">
<heading id="D0F5FE72277BFFFBFF72CAB1FCC0CDC7" bold="true" box="[151,776,1178,1204]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="B976950C277BFFFBFF72CAB1FCC0CDC7" bold="true" box="[151,776,1178,1204]" pageId="1" pageNumber="62">
<taxonomicName id="4C02329D277BFFFBFF72CAB1FD56CDC7" authority="Rodriguez &amp; Heads" authorityName="Rodriguez &amp; Heads" box="[151,670,1178,1204]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B976950C277BFFFBFF72CAB1FE55CDC7" bold="true" box="[151,413,1178,1204]" italics="true" pageId="1" pageNumber="62">Scapteriscus cerberus</emphasis>
Rodríguez &amp; Heads
</taxonomicName>
,
<taxonomicNameLabel id="A2452877277BFFFBFD4ECAB1FCC0CDC7" box="[683,776,1178,1204]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD491E277BFFFBFF72CA96FEF9CDA6" blockId="1.[151,776,1178,1237]" box="[151,305,1213,1237]" pageId="1" pageNumber="62">
<figureCitation id="1339559B277BFFFBFF72CA96FEE5CDA6" box="[151,301,1213,1237]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1616,1639]" captionStartId-1="3.[151,250,1797,1820]" captionTargetBox-0="[205,1394,199,1587]" captionTargetBox-1="[151,1435,193,1755]" captionTargetId-0="figure@2.[184,1403,195,1610]" captionTargetId-1="figure@3.[151,1436,193,1776]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." captionText-1="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri-0="https://zenodo.org/record/214513/files/figure.png" httpUri-1="https://zenodo.org/record/214514/files/figure.png" pageId="1" pageNumber="62">Figures 1 2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3181A95277BFFFBFF72CB2EFC6FCCA2" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="8BBD491E277BFFFBFF72CB2EFC6FCCA2" blockId="1.[151,1437,1284,2032]" pageId="1" pageNumber="62">
<emphasis id="B976950C277BFFFBFF72CB2EFE8ECC6E" bold="true" box="[151,326,1284,1309]" pageId="1" pageNumber="62">
<typeStatus id="54B9F7BC277BFFFBFF72CB2EFF18CC6E" box="[151,208,1285,1309]" pageId="1" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B9F7BC277BFFFBFEAACB2EFE7FCC6E" box="[335,439,1285,1309]" pageId="1" pageNumber="62" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F315098E277BFFFBFDEFCB2EFD6CCC6E" box="[522,676,1285,1309]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Cundinamarca, Puente Quetame.
<geoCoordinate id="EE362FD9277BFFFBFBD0CB2EFB6FCC6E" box="[1077,1191,1285,1309]" direction="north" orientation="latitude" pageId="1" pageNumber="62" precision="15" value="4.3330555">4º19'59''N</geoCoordinate>
<geoCoordinate id="EE362FD9277BFFFBFB57CB2EFAF1CC6E" box="[1202,1337,1285,1309]" direction="west" orientation="longitude" pageId="1" pageNumber="62" precision="15" value="-75.863884">75º51'50''W</geoCoordinate>
.
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,
<date id="FFBC6FDE277BFFFBFF72CB02FECCCC32" box="[151,260,1321,1345]" pageId="1" pageNumber="62" value="2009-10">Oct-2009</date>
, Balvuena S. [
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3008].
<typeStatus id="54B9F7BC277BFFFBFD40CB01FCCFCC31" box="[677,775,1322,1346]" pageId="1" pageNumber="62" type="paratype">Paratype</typeStatus>
male
<collectingCountry id="F315098E277BFFFBFCBECB02FC3DCC32" box="[859,1013,1321,1345]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Santander, Barbosa,
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,
<date id="FFBC6FDE277BFFFBFA8DCB02FF07CC16" pageId="1" pageNumber="62" value="2009-10">Oct- 2009</date>
[
<collectionCode id="ED13D1DB277BFFFBFF39CB66FEE7CC16" box="[220,303,1357,1381]" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="1" pageNumber="62">UNAB</collectionCode>
3577].
<typeStatus id="54B9F7BC277BFFFBFE62CB65FE21CC15" box="[391,489,1358,1382]" pageId="1" pageNumber="62" type="paratype">Paratype</typeStatus>
female
<collectingCountry id="F315098E277BFFFBFDA3CB66FD28CC16" box="[582,736,1357,1381]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Antioquia Bello,
<date id="FFBC6FDE277BFFFBFC55CB66FBE8CC16" box="[944,1056,1357,1381]" pageId="1" pageNumber="62" value="1995-04">Abr-1995</date>
. Cardona E. [
<collectionCode id="ED13D1DB277BFFFBFB5ECB66FADBCC16" box="[1211,1299,1357,1381]" pageId="1" pageNumber="62">CCOM</collectionCode>
5100].
<typeStatus id="54B9F7BC277BFFFBFA84CB65FF0FCCF9" pageId="1" pageNumber="62" type="paratype">Paratype</typeStatus>
male
<collectingCountry id="F315098E277BFFFBFEF6CB5AFE65CCFA" box="[275,429,1393,1417]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Antioquia Bello,
<date id="FFBC6FDE277BFFFBFD66CB5AFD3BCCFA" box="[643,755,1393,1417]" pageId="1" pageNumber="62" value="1995-04">Abr-1995</date>
. Cardona E. [
<collectionCode id="ED13D1DB277BFFFBFC76CB5AFC25CCFA" box="[915,1005,1393,1417]" pageId="1" pageNumber="62">CCOM</collectionCode>
5101].
<typeStatus id="54B9F7BC277BFFFBFBD9CB59FB56CCF9" box="[1084,1182,1394,1418]" pageId="1" pageNumber="62" type="paratype">Paratype</typeStatus>
female
<collectingCountry id="F315098E277BFFFBFB1ACB5AFA51CCFA" box="[1279,1433,1393,1417]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Antioquia, San Luis, Rio claro.
<specimenCount id="9D048297277BFFFBFDE0CBBEFD9ACCDE" box="[517,594,1429,1453]" pageId="1" pageNumber="62" type="generic">
<quantity id="4CFAE4FB277BFFFBFDE0CBBEFD86CCDE" box="[517,590,1429,1453]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="62" unit="m" value="440.0">440m</quantity>
.
</specimenCount>
s.n.m,
<date id="FFBC6FDE277BFFFBFD7CCBBEFCC0CCDE" box="[665,776,1429,1453]" pageId="1" pageNumber="62" value="1994-03">mar-1994</date>
, Amat G. [
<collectionCode id="ED13D1DB277BFFFBFC61CBBEFC0ACCDE" box="[900,962,1429,1453]" httpUri="http://grbio.org/cool/3cgd-ve52" name="Museo Javeriano de Historia Natural, Laboratoriao de Entomologia" pageId="1" pageNumber="62">MUJ</collectionCode>
386].
<typeStatus id="54B9F7BC277BFFFBFBE7CBBDFBACCCDD" box="[1026,1124,1430,1454]" pageId="1" pageNumber="62" type="paratype">Paratype</typeStatus>
male
<collectingCountry id="F315098E277BFFFBFB48CBBEFA8FCCDE" box="[1197,1351,1429,1453]" name="Colombia" pageId="1" pageNumber="62">COLOMBIA</collectingCountry>
: Antioquia, San Luis, Rio claro.
<specimenCount id="9D048297277BFFFBFE5ECB92FDC0CCA2" box="[443,520,1465,1489]" pageId="1" pageNumber="62" type="generic">
<quantity id="4CFAE4FB277BFFFBFE5ECB92FDCCCCA2" box="[443,516,1465,1489]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="62" unit="m" value="440.0">440m</quantity>
.
</specimenCount>
s.n.m,
<date id="FFBC6FDE277BFFFBFDA9CB92FD74CCA2" box="[588,700,1465,1489]" pageId="1" pageNumber="62" value="1994-03">mar-1994</date>
. Amat G. [
<collectionCode id="ED13D1DB277BFFFBFCD7CB92FCB8CCA2" box="[818,880,1465,1489]" httpUri="http://grbio.org/cool/3cgd-ve52" name="Museo Javeriano de Historia Natural, Laboratoriao de Entomologia" pageId="1" pageNumber="62">MUJ</collectionCode>
395].
</paragraph>
</subSubSection>
<subSubSection id="C3181A95277BFFFBFF22CBF6FDA0CF33" pageId="1" pageNumber="62" type="etymology">
<paragraph id="8BBD491E277BFFFBFF22CBF6FDA0CF33" blockId="1.[151,1437,1284,2032]" pageId="1" pageNumber="62">
<emphasis id="B976950C277BFFFBFF22CBF6FE86CC86" bold="true" box="[199,334,1501,1525]" pageId="1" pageNumber="62">Etymology.</emphasis>
The specific epithet is a noun in apposition derived from
<taxonomicName id="4C02329D277BFFFBFBC3CBF6FB46CC86" box="[1062,1166,1501,1525]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="67" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B976950C277BFFFBFBC3CBF6FB46CC86" box="[1062,1166,1501,1525]" italics="true" pageId="1" pageNumber="62">Cerberus</emphasis>
</taxonomicName>
(a Latinization of the Greek Kέρβερος or 'kerberos'), the fearsome dog of Greek and Roman mythology that guards the gates of Hades, the underworld. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3181A95277BFFFBFF22C867FAF4CF87" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="8BBD491E277BFFFBFF22C867FAF4CF87" blockId="1.[151,1437,1284,2032]" pageId="1" pageNumber="62">
<emphasis id="B976950C277BFFFBFF22C867FE89CF16" bold="true" box="[199,321,1612,1637]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
Brachypterous species with a large and robust pronotum; protibial dactyls evenly spaced (interdactylar distance approximately
<quantity id="4CFAE4FB277BFFFBFE3BC85BFD8ECFFB" box="[478,582,1648,1672]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="1" pageNumber="62" unit="mm" value="0.36">0.36 mm</quantity>
); tympana almost completely exposed; protrochanteral blade approximately
<quantity id="4CFAE4FB277BFFFBFF72C8BFFF25CFDF" box="[151,237,1684,1708]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="62" unit="mm" value="1.6">1.6 mm</quantity>
long.
<taxonomicName id="4C02329D277BFFFBFED1C8BEFDE3CFDE" box="[308,555,1685,1709]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="B976950C277BFFFBFED1C8BEFDE3CFDE" box="[308,555,1685,1709]" italics="true" pageId="1" pageNumber="62">Scapteriscus cerberus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C02329D277BFFFBFD19C8BEFC46CFDE" box="[764,910,1685,1709]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B976950C277BFFFBFD19C8BEFC46CFDE" box="[764,910,1685,1709]" italics="true" pageId="1" pageNumber="62">S. variegatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C02329D277BFFFBFC20C8BEFBACCFDE" box="[965,1124,1685,1709]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="abbreviatus">
<emphasis id="B976950C277BFFFBFC20C8BEFBACCFDE" box="[965,1124,1685,1709]" italics="true" pageId="1" pageNumber="62">S. abbreviatus</emphasis>
</taxonomicName>
in that it displays a marked reduction of the hind wings (brachyptery) and ocelli. Nevertheless, it is readily separated from these species by its large and robust pronotum, pigmented tegmina, reduced ocelli and markedly longer protrochanteral blade.
</paragraph>
</subSubSection>
<subSubSection id="C3181A95277BFFFEFF22C92BFE26CB64" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62" type="description">
<paragraph id="8BBD491E277BFFFBFF22C92BFAADCE4F" blockId="1.[151,1437,1284,2032]" pageId="1" pageNumber="62">
<emphasis id="B976950C277BFFFBFF22C92BFE91CE6A" bold="true" box="[199,345,1792,1817]" pageId="1" pageNumber="62">Description.</emphasis>
<emphasis id="B976950C277BFFFBFEBAC92AFE6DCE6A" box="[351,421,1793,1817]" italics="true" pageId="1" pageNumber="62">Head:</emphasis>
small ocelli, subcircular and relatively separated from compound eyes which are also reduced. General coloration: brown with 2 paired dark bands behind the compound eyes and ocelli. (
<figureCitation id="1339559B277BFFFBFB1BC90FFA89CE4F" box="[1278,1345,1828,1852]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8BBD491E277BFFFBFF22C962FE77CE83" blockId="1.[151,1437,1284,2032]" pageId="1" pageNumber="62">
<emphasis id="B976950C277BFFFBFF22C962FEE9CE12" box="[199,289,1865,1889]" italics="true" pageId="1" pageNumber="62">Thorax:</emphasis>
Pronotum robust and trapezoidal; two brown bands line the middle of the thorax; Two large anterior brown patches positioned surrounding the middle bands tips, whereas, two small posterior patches are positioned near the center of these bands. Another dark area can be seen in the anterior margin of the thorax, surrounding a bone-shaped clear area in the upper middle; two additional small brown patches present in the central region of the posterior margin (
<figureCitation id="1339559B277BFFFBFEBAC9F3FE6CCE83" box="[351,420,2008,2032]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
c).
</paragraph>
<caption id="DF7D19962778FFF8FF72C87BFB9ACFF7" httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="2" pageNumber="63" targetBox="[205,1394,199,1587]" targetPageId="2">
<paragraph id="8BBD491E2778FFF8FF72C87BFB9ACFF7" blockId="2.[151,1435,1616,1669]" pageId="2" pageNumber="63">
<emphasis id="B976950C2778FFF8FF72C87BFEDBCF16" bold="true" box="[151,275,1616,1639]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
<taxonomicName id="4C02329D2778FFF8FEFCC87AFE31CF15" box="[281,505,1616,1638]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="cerberus" status="sp. nov.">
<emphasis id="B976950C2778FFF8FEFCC87AFE31CF15" box="[281,505,1616,1638]" italics="true" pageId="2" pageNumber="63">Scapteriscus cerberus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24528772778FFF8FE1AC87AFD98CF15" box="[511,592,1617,1638]" pageId="2" pageNumber="63" rank="species">
<emphasis id="B976950C2778FFF8FE1AC87AFD98CF15" bold="true" box="[511,592,1617,1638]" pageId="2" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype male
<emphasis id="B976950C2778FFF8FD0BC87BFCC1CF14" bold="true" box="[750,777,1616,1639]" pageId="2" pageNumber="63">A)</emphasis>
lateral habitus,
<emphasis id="B976950C2778FFF8FC49C87BFC0DCF14" bold="true" box="[940,965,1616,1639]" pageId="2" pageNumber="63">B)</emphasis>
Fore tibiae dactyls
<emphasis id="B976950C2778FFF8FB68C87BFB60CF14" bold="true" box="[1165,1192,1616,1639]" pageId="2" pageNumber="63">C)</emphasis>
Dorsal habitus
<emphasis id="B976950C2778FFF8FAACC87BFAABCF14" bold="true" box="[1353,1379,1616,1639]" pageId="2" pageNumber="63">D)</emphasis>
Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1mm.
</paragraph>
</caption>
<paragraph id="8BBD491E2778FFF8FF22C887FC5ECE97" blockId="2.[151,1436,1708,2020]" pageId="2" pageNumber="63">
Lateral lobes of pronotum slightly pigmented (
<figureCitation id="1339559B2778FFF8FD3EC887FCD0CFB7" box="[731,792,1708,1732]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="2" pageNumber="63">Fig.2</figureCitation>
b); dactyls in the fore tibia rather thin and widely spaced and divergent (
<figureCitation id="1339559B2778FFF8FEDBC8FBFEB1CF9B" box="[318,377,1744,1768]" captionStart="FIGURE 1" captionStartId="2.[151,250,1616,1639]" captionTargetBox="[205,1394,199,1587]" captionTargetId="figure@2.[184,1403,195,1610]" captionTargetPageId="2" captionText="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="2" pageNumber="63">Fig.1</figureCitation>
b, 2e). The internal area of the fore tibiae is almost totally covered by granulations with a small smooth area in the basal zone (
<figureCitation id="1339559B2778FFF8FE13C8DFFDF3CE7F" box="[502,571,1780,1804]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
h). Tympana almost totally exposed in dorsal view (
<figureCitation id="1339559B2778FFF8FB63C8DFFB77CE7F" box="[1158,1215,1780,1804]" captionStart="FIGURE 1" captionStartId="2.[151,250,1616,1639]" captionTargetBox="[205,1394,199,1587]" captionTargetId="figure@2.[184,1403,195,1610]" captionTargetPageId="2" captionText="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="2" pageNumber="63">Fig 1</figureCitation>
d, 2d); the fore trochanter blade long, extending more than 2/3 the length of trochanter process (
<figureCitation id="1339559B2778FFF8FBCEC933FBAACE43" box="[1067,1122,1816,1840]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="2" pageNumber="63">Fig2</figureCitation>
b).
<collectingCountry id="F315098E2778FFF8FB6CC933FB09CE43" box="[1161,1217,1816,1840]" name="India" pageId="2" pageNumber="63">Hind</collectingCountry>
femora pigmented around its posterior middle with dark brown patches forming two well defined lines which become clearer as they move closer to the basal region.
<collectingCountry id="F315098E2778FFF8FE1AC94BFDFFCE0B" box="[511,567,1888,1912]" name="India" pageId="2" pageNumber="63">Hind</collectingCountry>
tibia with a dark band close to the femora articulation (
<figureCitation id="1339559B2778FFF8FB44C94BFB15CE0B" box="[1185,1245,1888,1912]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="2" pageNumber="63">Fig 2</figureCitation>
k); dorsal margin of the hind tibia with 4 spines, with six are located in its the apical border (
<figureCitation id="1339559B2778FFF8FC3EC9AFFBD7CEEF" box="[987,1055,1924,1948]" captionStart="FIGURE 2" captionStartId="3.[151,250,1797,1820]" captionTargetBox="[151,1435,193,1755]" captionTargetId="figure@3.[151,1436,193,1776]" captionTargetPageId="3" captionText="FIGURE 2. Scapteriscus cerverus sp. nov. A) Head: anterior view. B) Head and pronotum: lateral view. C) Pronotum: dorsal view. D) Timpanum: dorsal view. E) External view of fore tibiae showing disposition of dactyls. F) Fore throchanter blade. G) Male tegmina. H) Inner view of fore tibiae showing granulation. I) Inner view of hind tibiae showing spines. J) External view of hind tibiae, showing spines. K) Hind leg, showing pigmentation of femur. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="2" pageNumber="63">
Fig.
<date id="FFBC6FDE2778FFF8FBEAC9AFFBD7CEEF" box="[1039,1055,1924,1948]" pageId="2" pageNumber="63">2</date>
</figureCitation>
i, 2j). Wings short, covering about 2/3 of the abdomen; tegmina slightly pigmented, not transparent, and covering about 2/3 of the abdomen (
<figureCitation id="1339559B2778FFF8FAA3C983FA4FCEB3" box="[1350,1415,1960,1984]" captionStart="FIGURE 1" captionStartId="2.[151,250,1616,1639]" captionTargetBox="[205,1394,199,1587]" captionTargetId="figure@2.[184,1403,195,1610]" captionTargetPageId="2" captionText="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
c,
<quantity id="4CFAE4FB2778FFF8FF72C9E7FF7DCE97" box="[151,181,1996,2020]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="63" unit="g" value="2.0">2g</quantity>
); tegmina and wings approximately of the same lengths (
<figureCitation id="1339559B2778FFF8FCD3C9E7FCBFCE97" box="[822,887,1996,2020]" captionStart="FIGURE 1" captionStartId="2.[151,250,1616,1639]" captionTargetBox="[205,1394,199,1587]" captionTargetId="figure@2.[184,1403,195,1610]" captionTargetPageId="2" captionText="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
c).
</paragraph>
<caption id="DF7D19962779FFF9FF72C92EFB7DCE04" httpUri="https://zenodo.org/record/214514/files/figure.png" pageId="3" pageNumber="64" targetBox="[151,1435,193,1755]" targetPageId="3">
<paragraph id="8BBD491E2779FFF9FF72C92EFB7DCE04" blockId="3.[151,1437,1797,1912]" pageId="3" pageNumber="64">
<emphasis id="B976950C2779FFF9FF72C92EFEDBCE69" bold="true" box="[151,275,1797,1820]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
<taxonomicName id="4C02329D2779FFF9FEFFC92DFE30CE68" box="[282,504,1798,1819]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="cerverus" status="sp. nov.">
<emphasis id="B976950C2779FFF9FEFFC92DFE30CE68" box="[282,504,1798,1819]" italics="true" pageId="3" pageNumber="64">Scapteriscus cerverus</emphasis>
</taxonomicName>
<emphasis id="B976950C2779FFF9FE1AC92DFDBACE6F" bold="true" box="[511,626,1797,1820]" pageId="3" pageNumber="64">
<taxonomicNameLabel id="A24528772779FFF9FE1AC92DFD98CE68" box="[511,592,1798,1819]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
A)
</emphasis>
Head: anterior view.
<emphasis id="B976950C2779FFF9FCAAC92EFCA1CE6F" bold="true" box="[847,873,1797,1820]" pageId="3" pageNumber="64">B)</emphasis>
Head and pronotum: lateral view.
<emphasis id="B976950C2779FFF9FB2EC92EFB2DCE6F" bold="true" box="[1227,1253,1797,1820]" pageId="3" pageNumber="64">C)</emphasis>
Pronotum: dorsal view.
<emphasis id="B976950C2779FFF9FF36C908FF25CE49" bold="true" box="[211,237,1827,1850]" pageId="3" pageNumber="64">D)</emphasis>
Timpanum: dorsal view.
<emphasis id="B976950C2779FFF9FE14C908FDC1CE49" bold="true" box="[497,521,1827,1850]" pageId="3" pageNumber="64">E)</emphasis>
External view of fore tibiae showing disposition of dactyls.
<emphasis id="B976950C2779FFF9FB8AC908FB4FCE49" bold="true" box="[1135,1159,1827,1850]" pageId="3" pageNumber="64">F)</emphasis>
Fore throchanter blade.
<emphasis id="B976950C2779FFF9FA65C908FA55CE49" bold="true" box="[1408,1437,1827,1850]" pageId="3" pageNumber="64">G)</emphasis>
Male tegmina.
<emphasis id="B976950C2779FFF9FED5C969FE84CE2A" bold="true" box="[304,332,1858,1881]" pageId="3" pageNumber="64">H)</emphasis>
Inner view of fore tibiae showing granulation.
<emphasis id="B976950C2779FFF9FCCBC969FC89CE2A" bold="true" box="[814,833,1858,1881]" pageId="3" pageNumber="64">I)</emphasis>
Inner view of hind tibiae showing spines.
<emphasis id="B976950C2779FFF9FB17C969FACFCE2A" bold="true" box="[1266,1287,1858,1881]" pageId="3" pageNumber="64">J)</emphasis>
External view of hind tibiae, showing spines.
<emphasis id="B976950C2779FFF9FE36C94AFE26CE0B" bold="true" box="[467,494,1889,1912]" pageId="3" pageNumber="64">K)</emphasis>
Hind leg, showing pigmentation of femur. Length of scale line: 1mm.
</paragraph>
</caption>
<paragraph id="8BBD491E2779FFFEFF22C98FFDADC9A7" blockId="3.[151,1436,1956,2016]" lastBlockId="4.[151,1437,151,535]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">
<emphasis id="B976950C2779FFF9FF22C98FFEF2CECF" box="[199,314,1956,1980]" italics="true" pageId="3" pageNumber="64">Abdomen:</emphasis>
General color yellowish, with dark brown marks on posterior margin of each sclerite. Other dark bands are located along the dorsal and dorsolateral lines, and some small marks located in the lateral margin of each sclerite. Cerci short and the subgenital plate is not produced as it is in some species of the genus (e.g.
<emphasis id="B976950C277EFFFEFA62CEB3FE25C9A7" italics="true" pageId="4" pageNumber="65">
<taxonomicName id="4C02329D277EFFFEFA62CEB3FE98C9A7" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="cuadripunctatus">S. cuadripunctatus</taxonomicName>
,
<taxonomicName id="4C02329D277EFFFEFEBECE96FE25C9A7" box="[347,493,189,212]" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="mexicanus">S. mexicanus</taxonomicName>
</emphasis>
) (
<figureCitation id="1339559B277EFFFEFDE0CE97FD8EC9A7" box="[517,582,188,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1616,1639]" captionTargetBox="[205,1394,199,1587]" captionTargetId="figure@2.[184,1403,195,1610]" captionTargetPageId="2" captionText="FIGURE 1. Scapteriscus cerberus sp. nov. holotype male A) lateral habitus, B) Fore tibiae dactyls C) Dorsal habitus D) Timpanum. Notice the robust appearance that characterize this species. Length of scale line: 1 mm." httpUri="https://zenodo.org/record/214513/files/figure.png" pageId="4" pageNumber="65">Fig. 1</figureCitation>
c).
</paragraph>
<paragraph id="8BBD491E277EFFFEFF22CECBFE26CB64" blockId="4.[151,1437,151,535]" pageId="4" pageNumber="65">
<emphasis id="B976950C277EFFFEFF22CECBFE04C98B" bold="true" box="[199,460,223,248]" pageId="4" pageNumber="65">Measurements (mm):</emphasis>
<specimenCount id="9D048297277EFFFEFE3CCEF4FD82C984" box="[473,586,223,247]" pageId="4" pageNumber="65" type="female">2 females</specimenCount>
: Ocellar lenght: ~0.3 0.270.32; Interocellar distance: ~1.07 0.971.17; Interocular distance: ~1.87 1.672.07; Ocular-ocellar distance: ~0.34 0.300.37; Length of pronotum: ~8.50 7.509.50; Width of pronotum: ~7.53 7.267.78; Trochanter blade length: ~1.26 1.251.27; Interdactyl distance: ~0.25 0.20.3; Total length: ~26.7 24.9028.50.
<specimenCount id="9D048297277EFFFEFD2DCF61FCE1C817" box="[712,809,330,356]" pageId="4" pageNumber="65" type="male">4 males</specimenCount>
: Ocellar length ~0.34 0.220.40; Interocellar distance: ~1.11 0.801.31; Interocular distance: ~1.81 1.352.17; Ocular-ocellar distance: ~0.28 0.250.35; Length of pronotum: ~9.85 7.9611.96; Width of pronotum: ~7.15 6.028.54; Trochanter blade length: ~1.73 1.002.53; Interdactyl distance: ~0.41 0.220.61; Total length: ~28.46 24.0034.52. Ratios of morphometric measurements of
<taxonomicName id="4C02329D277EFFFEFA62CF93FF31C887" class="Insecta" family="Lathiceridae" genus="Scapteriscus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="B976950C277EFFFEFA62CF93FF31C887" italics="true" pageId="4" pageNumber="65">S. cerberus</emphasis>
</taxonomicName>
are shown in the table 1. One of the most representative is pronotal length/ total length, which reflects the large pronotum in this species.
</paragraph>
</subSubSection>
</treatment>
</document>