347 lines
37 KiB
XML
347 lines
37 KiB
XML
<document id="4A8DD60BF971A86AED72CBCD46A33064" ID-DOI="10.5281/zenodo.5171342" ID-ISSN="1942-1354" ID-Zenodo-Dep="5171342" ID-ZooBank="511C0AF6-E9BF-4D75-B9ED-A19FCC30C721" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628272289452" checkinUser="felipe" docAuthor="Otto, Robert L." docDate="2017" docId="03AB8D4DFFFA890DEFFB685C8299F88C" docLanguage="en" docName="InsectaMundi.0558.1-19.pdf" docOrigin="Insecta Mundi 2017 (558)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Miruantennus cuneiformis , Otto 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="12" masterDocId="FF92F535FFF68900EF3B6C1E8375FFF5" masterDocTitle="Descriptions of six new species of false click beetles (Coleoptera: Eucnemidae: Macraulacinae), with new identification keys for one tribe and two genera" masterLastPageNumber="19" masterPageNumber="1" pageNumber="11" updateTime="1699038813006" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="A2C78F60C95E395AF6BED3491698FFC6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E2E420DA1A925D49AE6E25CDC5758EDC">
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<mods:title id="6030AA479E422A59534571CFDDC143C9">Descriptions of six new species of false click beetles (Coleoptera: Eucnemidae: Macraulacinae), with new identification keys for one tribe and two genera</mods:title>
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<mods:name id="83CE5329901CFBADDB2789CB5FAF88E1" type="personal">
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<mods:roleTerm id="7419249C9F88382E50270F1BC53C6380">Author</mods:roleTerm>
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<mods:namePart id="B6304BA8D06217E01409EA5B65A3F733">Otto, Robert L.</mods:namePart>
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<mods:typeOfResource id="042A201122AF2078190D1CECF4635D7A">text</mods:typeOfResource>
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<mods:title id="40E4DF4D29921E166900A3A115D6CEDF">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="BB027764244FEDCB3A40EBD4C25ECAFA">
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<mods:date id="9E6A18576D9DDD51463CBC3E2FAE327E">2017</mods:date>
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<mods:detail id="674DC702FC71C081A14D88E31CB8AA3B" type="pubDate">
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<mods:number id="877286FB9B8750D118B77A02B638D662">2017-06-30</mods:number>
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<mods:detail id="5DA1ECB0A9C25AB50C256FD4C614D55F" type="volume">
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<mods:number id="AF35857A6D44A2031A6973A9DC5DFF32">2017</mods:number>
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</mods:detail>
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<mods:detail id="1AD1C0FF306B4F7D3F1D73C6C0B98A8F" type="issue">
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<mods:number id="12866D4A690529BB4A17B192D826ED28">558</mods:number>
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<mods:start id="F46788AC83AA4E8935456A8A0F05FB8A">1</mods:start>
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<mods:end id="A12E8E6AD305B91790B56E6C795B7468">19</mods:end>
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<mods:classification id="75DBD166737BEEDE5C96BBF123B3C420">journal article</mods:classification>
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<mods:identifier id="5C8DFB03DB98639A5E7FFD8220AEFB9B" type="DOI">10.5281/zenodo.5171342</mods:identifier>
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<mods:identifier id="FDE60D9D703DCFE7B7A9FE4D729B4FAF" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="7070EB32562C201DCBAD3BB9D8182524" type="Zenodo-Dep">5171342</mods:identifier>
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<treatment id="03AB8D4DFFFA890DEFFB685C8299F88C" ID-DOI="http://doi.org/10.5281/zenodo.5185334" ID-Zenodo-Dep="5185334" LSID="urn:lsid:plazi:treatment:03AB8D4DFFFA890DEFFB685C8299F88C" httpUri="http://treatment.plazi.org/id/03AB8D4DFFFA890DEFFB685C8299F88C" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
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<subSubSection id="C3186FD0FFFA890CEFFB685C81EAFBAF" box="[192,671,1089,1114]" pageId="12" pageNumber="11" type="nomenclature">
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<paragraph id="8BBD3C5BFFFA890CEFFB685C81EAFBAF" blockId="12.[192,671,1089,1146]" box="[192,671,1089,1114]" pageId="12" pageNumber="11">
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<heading id="D0F58B37FFFA890CEFFB685C81EAFBAF" bold="true" box="[192,671,1089,1114]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
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<emphasis id="B976E049FFFA890CEFFB685C81EAFBAF" bold="true" box="[192,671,1089,1114]" pageId="12" pageNumber="11">
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<taxonomicName id="4C0247D8FFFA890CEFFB685C815AFBAF" authority="Otto, 2017" authorityName=", Otto" authorityYear="2017" box="[192,559,1089,1114]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="cuneiformis" status="sp. nov.">
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<emphasis id="B976E049FFFA890CEFFB685C815AFBAF" bold="true" box="[192,559,1089,1114]" italics="true" pageId="12" pageNumber="11">Miruantennus cuneiformis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2455D32FFFA890CED0D685C81EAFBAF" box="[566,671,1090,1114]" pageId="12" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3186FD0FFFA890CEFFB687C8234FB8F" box="[192,321,1122,1146]" pageId="12" pageNumber="11" type="description">
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<paragraph id="8BBD3C5BFFFA890CEFFB687C8234FB8F" blockId="12.[192,671,1089,1146]" box="[192,321,1122,1146]" pageId="12" pageNumber="11">
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<figureCitation id="133920DEFFFA890CEFFB687C8234FB8F" box="[192,321,1122,1146]" captionStart="Figures 23–27" captionStartId="20.[193,289,1629,1651]" captionTargetBox="[318,1316,192,1591]" captionTargetId="figure-61@20.[318,1317,192,1613]" captionTargetPageId="20" captionText="Figures 23–27. Miruantennus cuneiformis sp.nov. 23) Male holotype (BMNH), dorsal habitus. 24) Male holotype (BMNH), lateral habitus. 25) Male holotype (BMNH), antenna, dorsal view. 26) Female paratype (GERP), dorsal habitus. 27) Female paratype (GERP), antenna, dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5171356" httpUri="https://zenodo.org/record/5171356/files/figure.png" pageId="12" pageNumber="11">Fig. 23–27</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3186FD0FFFA890CEFFB68BF8280FB2C" pageId="12" pageNumber="11" type="diagnosis">
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<paragraph id="8BBD3C5BFFFA890CEFFB68BF8280FB2C" blockId="12.[192,1440,1185,1242]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFFB68BF8225FB4C" bold="true" box="[192,336,1185,1209]" pageId="12" pageNumber="11">Diagnosis.</emphasis>
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Black coloration and strongly cuneiform habitus will distinguish the new species from
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<taxonomicName id="4C0247D8FFFA890CEA4468BC8262FB2F" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="basalis">
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<emphasis id="B976E049FFFA890CEA4468BC8262FB2F" italics="true" pageId="12" pageNumber="11">M. basalis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C0247D8FFFA890CEE6F68DC829BFB2F" authorityName=", Otto" authorityYear="2017" box="[340,494,1218,1242]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="chinensis">
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<emphasis id="B976E049FFFA890CEE6F68DC829BFB2F" box="[340,494,1218,1242]" italics="true" pageId="12" pageNumber="11">M. chinensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3186FD0FFFA890DEFFB691F86D5FCEC" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="description">
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<paragraph id="8BBD3C5BFFFA890CEFFB691F87F9FA6F" blockId="12.[192,1440,1281,1850]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFFB691F8141FAEF" bold="true" box="[192,564,1281,1306]" pageId="12" pageNumber="11">
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Description. Male
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<typeStatus id="54B982F9FFFA890CEE8E691C815BFAEF" box="[437,558,1282,1306]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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Length, 7.0 mm. Width,
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<quantity id="4CFA91BEFFFA890CEC60691F80B3FAEC" box="[859,966,1281,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="11" unit="mm" value="1.25">1.25 mm</quantity>
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. Body subcylindrical, elongate, strongly cuneiform toward elytral apex; uniformly black; scape and antennomeres VI–XI black, pedicel and antennomeres III–V dark brown; femur and tibiae dark brown, tarsi dark brown; head, apical 2/3 of pronotum and much of elytra clothed with short, recumbent whitish setae; basal 1/3 of pronotum, elytral humeri and sutural areas with elongate, recumbent whitish setae (
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<figureCitation id="133920DEFFFA890CECC1699C870AFA6F" box="[1018,1151,1410,1434]" captionStart="Figures 23–27" captionStartId="20.[193,289,1629,1651]" captionTargetBox="[318,1316,192,1591]" captionTargetId="figure-61@20.[318,1317,192,1613]" captionTargetPageId="20" captionText="Figures 23–27. Miruantennus cuneiformis sp.nov. 23) Male holotype (BMNH), dorsal habitus. 24) Male holotype (BMNH), lateral habitus. 25) Male holotype (BMNH), antenna, dorsal view. 26) Female paratype (GERP), dorsal habitus. 27) Female paratype (GERP), antenna, dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5171356" httpUri="https://zenodo.org/record/5171356/files/figure.png" pageId="12" pageNumber="11">Fig. 23–24</figureCitation>
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).
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</paragraph>
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<paragraph id="8BBD3C5BFFFA890CEFCB69BC8764FA0C" blockId="12.[192,1440,1281,1850]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFCB69BC8235FA4F" bold="true" box="[240,320,1442,1466]" pageId="12" pageNumber="11">Head:</emphasis>
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Densely punctate, almost rugose, subspherical; frons convex, without median carina or fovea above frontoclypeal region; surfaces shiny; apical margin of frontoclypeal region rounded, about 2.0 times wider than base; mandibles stout, bidentate, densely punctate.
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</paragraph>
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<paragraph id="8BBD3C5BFFFA890CEFCB6A1F8612F98F" blockId="12.[192,1440,1281,1850]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFCB6A1F8213F9EC" bold="true" box="[240,358,1537,1561]" pageId="12" pageNumber="11">Antenna</emphasis>
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(
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<figureCitation id="133920DEFFFA890CEE4C6A1C82B8F9EC" box="[375,461,1537,1562]" captionStart="Figures 23–27" captionStartId="20.[193,289,1629,1651]" captionTargetBox="[318,1316,192,1591]" captionTargetId="figure-61@20.[318,1317,192,1613]" captionTargetPageId="20" captionText="Figures 23–27. Miruantennus cuneiformis sp.nov. 23) Male holotype (BMNH), dorsal habitus. 24) Male holotype (BMNH), lateral habitus. 25) Male holotype (BMNH), antenna, dorsal view. 26) Female paratype (GERP), dorsal habitus. 27) Female paratype (GERP), antenna, dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5171356" httpUri="https://zenodo.org/record/5171356/files/figure.png" pageId="12" pageNumber="11">Fig. 25</figureCitation>
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): Capitate from antennomeres VIII–XI, attaining elytral humeri; antennomere III short, as long as II; antennomere IV slightly shorter than III, as long as V; combined lengths of antennomeres IV and V longer than III; antennomeres VI and VII quadrate, both larger than V; antennomeres VIII–XI each much longer than wide, as long as the combined lengths of III–V together.
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</paragraph>
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<paragraph id="8BBD3C5BFFFA890CEFCB6A9C86D5F92F" blockId="12.[192,1440,1281,1850]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFCB6A9C82F6F96F" bold="true" box="[240,387,1666,1690]" pageId="12" pageNumber="11">Pronotum:</emphasis>
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Surfaces dull, densely rugose to granulose; longer than wide, with moderate, sharp hind angles; lateral sides arcuate, slightly constricted basally above hind angles; disc convex with shallow median groove, without circular fovea; base sinuous, slightly depressed at either side above scutellum.
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</paragraph>
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<paragraph id="8BBD3C5BFFFA890CEFCB6AFF87FCF90F" blockId="12.[192,1440,1281,1850]" box="[240,1161,1761,1786]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFCB6AFF82F1F90C" bold="true" box="[240,388,1761,1785]" pageId="12" pageNumber="11">Scutellum:</emphasis>
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Short, sub-triangular, medially elevated and distally rounded.
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</paragraph>
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<paragraph id="8BBD3C5BFFFA890CEFCB6B1C81FCF8CC" blockId="12.[192,1440,1281,1850]" pageId="12" pageNumber="11">
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<emphasis id="B976E049FFFA890CEFCB6B1C823AF8EF" bold="true" box="[240,335,1794,1818]" pageId="12" pageNumber="11">Elytra:</emphasis>
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Indistinctly striate, except along elytral humeri and sutural areas; interstices flattened; surfaces shiny with dense punctures.
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</paragraph>
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<paragraph id="8BBD3C5BFFFB890DEFCB6CDC878CFEEF" blockId="13.[192,1440,194,378]" pageId="13" pageNumber="12">
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<emphasis id="B976E049FFFB890DEFCB6CDC824DFF2F" bold="true" box="[240,312,194,218]" pageId="13" pageNumber="12">Legs:</emphasis>
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First tarsomere as long as the combined lengths of the remaining four on mesothoracic and metathoracic tarsi; tibiae rounded in cross section; metathoracic tarsomeres I–III simple; metathoracic tarsomere IV excavated; metathoracic tarsomere V elongate with basally swollen claws.
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</paragraph>
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<paragraph id="8BBD3C5BFFFB890DEFCB6D3C817FFE8F" blockId="13.[192,1440,194,378]" pageId="13" pageNumber="12">
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<emphasis id="B976E049FFFB890DEFCB6D3C8223FECF" bold="true" box="[240,342,290,314]" pageId="13" pageNumber="12">Venter:</emphasis>
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Closely punctate, with elongate, recumbent whitish setae; hypomeron simple, without lateral antennal grooves; metathoracic episternum apically wide; metathoracic coxal plates medially 3 times wider than laterally.
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</paragraph>
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<paragraph id="8BBD3C5BFFFB890DEFFB6DBF86D5FCEC" blockId="13.[192,1440,417,794]" pageId="13" pageNumber="12">
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<emphasis id="B976E049FFFB890DEFFB6DBF8239FE4C" bold="true" box="[192,332,417,441]" pageId="13" pageNumber="12">Variation.</emphasis>
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One male and
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<specimenCount id="9D04F7D2FFFB890DED3E6DBC81FCFE4F" box="[517,649,418,442]" pageId="13" pageNumber="12" type="female">one female</specimenCount>
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<typeStatus id="54B982F9FFFB890DEDAA6DBC818AFE4F" box="[657,767,418,442]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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were examined. Male
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<typeStatus id="54B982F9FFFB890DEB2F6DBC87F7FE4F" box="[1044,1154,418,442]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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measured
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<quantity id="4CFA91BEFFFB890DEA326DBF8611FE4C" box="[1289,1380,417,441]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="13" pageNumber="12" unit="mm" value="6.5">6.5 mm</quantity>
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long and 1.0 mm wide, slightly shorter and narrower than the
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<typeStatus id="54B982F9FFFB890DEC486DDC80AAFE2F" box="[883,991,450,474]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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. Female
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<typeStatus id="54B982F9FFFB890DEB726DDC87C2FE2F" box="[1097,1207,450,474]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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(
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<figureCitation id="133920DEFFFB890DEBFD6DDC866CFE2C" box="[1222,1305,449,474]" captionStart="Figures 23–27" captionStartId="20.[193,289,1629,1651]" captionTargetBox="[318,1316,192,1591]" captionTargetId="figure-61@20.[318,1317,192,1613]" captionTargetPageId="20" captionText="Figures 23–27. Miruantennus cuneiformis sp.nov. 23) Male holotype (BMNH), dorsal habitus. 24) Male holotype (BMNH), lateral habitus. 25) Male holotype (BMNH), antenna, dorsal view. 26) Female paratype (GERP), dorsal habitus. 27) Female paratype (GERP), antenna, dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5171356" httpUri="https://zenodo.org/record/5171356/files/figure.png" pageId="13" pageNumber="12">Fig. 26</figureCitation>
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) measured
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<quantity id="4CFA91BEFFFB890DEFFB6DFF826BFE0C" box="[192,286,481,505]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="12" unit="mm" value="4.5">4.5 mm</quantity>
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long and 1.0 mm wide, also shorter and narrower than the
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<typeStatus id="54B982F9FFFB890DEB236DFC87F0FE0F" box="[1048,1157,482,506]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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. Male
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<typeStatus id="54B982F9FFFB890DEBE26DFC8632FE0F" box="[1241,1351,482,506]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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differs from the
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<typeStatus id="54B982F9FFFB890DEE096E1C82EEFDEF" box="[306,411,514,538]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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on a number of features. Firstly, arrangement of elongate setae on the basal 1/3 of the pronotum as well as elytral humeri and sutural areas are absent in the male
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<typeStatus id="54B982F9FFFB890DEB956E3C8655FDCF" box="[1198,1312,546,570]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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. Secondly, median groove on the pronotal disc is also absent in the
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<typeStatus id="54B982F9FFFB890DEC446E5C8084FDAF" box="[895,1009,578,602]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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. Lastly, pedicel and antennomeres III–V are darker than the
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<typeStatus id="54B982F9FFFB890DED386E7C8105FD8F" box="[515,624,610,634]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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. Female
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<typeStatus id="54B982F9FFFB890DEDE66E7C803EFD8F" box="[733,843,610,634]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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differs from the male
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<typeStatus id="54B982F9FFFB890DEB676E7C87B0FD8F" box="[1116,1221,610,634]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
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based on two features. First, the last abdominal segment is hidden beneath the elytra, exposed beyond elytral apices in males. Second, female antennal structures (
|
||
<figureCitation id="133920DEFFFB890DEC396EBC802DFD4F" box="[770,856,674,698]" captionStart="Figures 23–27" captionStartId="20.[193,289,1629,1651]" captionTargetBox="[318,1316,192,1591]" captionTargetId="figure-61@20.[318,1317,192,1613]" captionTargetPageId="20" captionText="Figures 23–27. Miruantennus cuneiformis sp.nov. 23) Male holotype (BMNH), dorsal habitus. 24) Male holotype (BMNH), lateral habitus. 25) Male holotype (BMNH), antenna, dorsal view. 26) Female paratype (GERP), dorsal habitus. 27) Female paratype (GERP), antenna, dorsal view. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5171356" httpUri="https://zenodo.org/record/5171356/files/figure.png" pageId="13" pageNumber="12">Fig. 27</figureCitation>
|
||
) are markedly different than the
|
||
<typeStatus id="54B982F9FFFB890DEBC66EBC861FFD4F" box="[1277,1386,674,698]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
. For female
|
||
<typeStatus id="54B982F9FFFB890DEE236EDC82FCFD2F" box="[280,393,706,730]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
|
||
, antennomere III much longer than IV; combined lengths of antennomeres IV and V as long as III; antennomere VI as long as V; antennomere VII slightly longer and wider than VI, shorter than VIII; antennomeres VIII–X each wider than VII, quadrate; antennomere XI slightly longer than X.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186FD0FFFB890DEFFB6F5C81DEFB0F" pageId="13" pageNumber="12" type="materials_examined">
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB6F5C8054FC2C" blockId="13.[192,1440,833,986]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB6F5C816EFCAF" bold="true" box="[192,539,833,858]" pageId="13" pageNumber="12">
|
||
Type Material.
|
||
<typeStatus id="54B982F9FFFB890DEEA76F5C816EFCAF" box="[412,539,834,858]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<specimenCount id="9D04F7D2FFFB890DED156F5C8137FCAC" box="[558,578,834,857]" pageId="13" pageNumber="12" type="male">♂</specimenCount>
|
||
: “
|
||
<emphasis id="B976E049FFFB890DED5B6F5C81B3FCAF" bold="true" box="[608,710,834,858]" pageId="13" pageNumber="12">Borneo</emphasis>
|
||
,
|
||
<collectingRegion id="49C6F2B9FFFB890DEDE26F5C805FFCAF" box="[729,810,834,858]" country="Malaysia" name="Sabah" pageId="13" pageNumber="12">Sabah</collectingRegion>
|
||
,
|
||
<quantity id="4CFA91BEFFFB890DEC016F5C80E5FCAF" box="[826,912,834,858]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="13" pageNumber="12" unit="m" value="1240.0">1240m</quantity>
|
||
, Crocker Range,
|
||
<date id="FFBC1A9BFFFB890DEB4B6F5C87A0FCAF" box="[1136,1237,834,858]" pageId="13" pageNumber="12" value="2013-04">IV.2013</date>
|
||
, Kota Kinabalu → Tambunan,
|
||
<geoCoordinate id="EE365A9CFFFB890DEEBB6F7F8166FC8C" box="[384,531,865,889]" degrees="5" direction="north" minutes="51" orientation="latitude" pageId="13" pageNumber="12" precision="1" seconds="33.7" value="5.859361">N5°51’33.7”</geoCoordinate>
|
||
<geoCoordinate id="EE365A9CFFFB890DED286F7F81B5FC8C" box="[531,704,865,889]" degrees="116" direction="east" minutes="17" orientation="longitude" pageId="13" pageNumber="12" precision="1" seconds="24.1" value="116.29002">E116°17’24.1”</geoCoordinate>
|
||
, General collecting” / “
|
||
<collectionCode id="ED13A49EFFFB890DECDD6F7C8737FC8F" box="[998,1090,866,890]" pageId="13" pageNumber="12">BMNH</collectionCode>
|
||
{E} 2013-58, M.V.L.Barclay, B.H.Garner, H.Mendel and A.Giusti” / “
|
||
<collectionCode id="ED13A49EFFFB890DED846F9C8069FC6F" box="[703,796,898,922]" pageId="13" pageNumber="12">BMNH</collectionCode>
|
||
(E), 1221775” (label upside-down) / “
|
||
<emphasis id="B976E049FFFB890DEBD06F9F86EDFC6C" bold="true" box="[1259,1432,897,921]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54B982F9FFFB890DEBD06F9F86E1FC6C" box="[1259,1428,897,921]" pageId="13" pageNumber="12" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEFFB6FBC8205FC4C" authorityName="Otto. Following" authorityYear="2016" box="[192,368,930,953]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEFFB6FBC8205FC4C" box="[192,368,930,953]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEE446FBC812DFC4F" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[383,600,930,954]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cuneiformis">
|
||
<emphasis id="B976E049FFFB890DEE446FBC817AFC4F" box="[383,527,930,954]" italics="true" pageId="13" pageNumber="12">cuneiformis</emphasis>
|
||
, Otto
|
||
</taxonomicName>
|
||
, det. R.L. Otto, 2015” (
|
||
<specimenCount id="9D04F7D2FFFB890DEC496FBC80F3FC4C" box="[882,902,930,953]" pageId="13" pageNumber="12" type="male">♂</specimenCount>
|
||
handwritten behind species name on label) [red printed label].
|
||
<typeStatus id="54B982F9FFFB890DEE946FDC816BFC2F" box="[431,542,962,986]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||
deposited in
|
||
<collectionCode id="ED13A49EFFFB890DEDFA6FDC8068FC2C" box="[705,797,962,985]" pageId="13" pageNumber="12">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB681C81DEFB0F" blockId="13.[192,1440,1025,1274]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB681C8221FBEF" bold="true" box="[192,340,1026,1050]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54B982F9FFFB890DEFFB681C823BFBEF" box="[192,334,1026,1050]" pageId="13" pageNumber="12" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D04F7D2FFFB890DEE60681C820BFBEC" box="[347,382,1026,1050]" pageId="13" pageNumber="12" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D04F7D2FFFB890DEEB6681C82DAFBEC" box="[397,431,1026,1050]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
|
||
:
|
||
<emphasis id="B976E049FFFB890DEE85681F8115FBEC" bold="true" box="[446,608,1025,1049]" pageId="13" pageNumber="12">
|
||
<collectingCountry id="F3157CCBFFFB890DEE85681F812EFBEC" box="[446,603,1025,1049]" name="Malaysia" pageId="13" pageNumber="12">MALAYSIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D04F7D2FFFB890DED5C681C81FFFBEC" box="[615,650,1026,1050]" pageId="13" pageNumber="12" type="male">1♂</specimenCount>
|
||
, “Borneo, 15–
|
||
<date id="FFBC1A9BFFFB890DEC08681F80DAFBEC" box="[819,943,1025,1049]" pageId="13" pageNumber="12" value="1993-04-27">27.4.1993</date>
|
||
,
|
||
<collectingRegion id="49C6F2B9FFFB890DEC81681C877DFBEF" box="[954,1032,1026,1050]" country="Malaysia" name="Sabah" pageId="13" pageNumber="12">Sabah</collectingRegion>
|
||
Crocker Mt., Gunong Emas env., Jenis and Strba leg.” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green framed white label) / “
|
||
<emphasis id="B976E049FFFB890DEEE8685F810EFBAC" bold="true" box="[467,635,1089,1113]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54B982F9FFFB890DEEE8685F8102FBAC" box="[467,631,1089,1113]" pageId="13" pageNumber="12" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEDB0685C804EFBAC" authorityName="Otto. Following" authorityYear="2016" box="[651,827,1090,1113]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEDB0685C804EFBAC" box="[651,827,1090,1113]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEC71685C8757FBAF" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[842,1058,1090,1114]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cuneiformis">
|
||
<emphasis id="B976E049FFFB890DEC71685C80AFFBAF" box="[842,986,1090,1114]" italics="true" pageId="13" pageNumber="12">cuneiformis</emphasis>
|
||
, Otto
|
||
</taxonomicName>
|
||
, det. R.L. Otto, 2015” (
|
||
<specimenCount id="9D04F7D2FFFB890DEA06685C8624FBAC" box="[1341,1361,1090,1113]" pageId="13" pageNumber="12" type="male">♂</specimenCount>
|
||
handwritten behind species name on label) [yellow printed label];
|
||
<specimenCount id="9D04F7D2FFFB890DEC89687C80A0FB8C" box="[946,981,1122,1146]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
|
||
, “W MALAYSIA –
|
||
<collectingRegion id="49C6F2B9FFFB890DEBF9687C8656FB8F" box="[1218,1315,1122,1146]" country="Malaysia" name="Pahang" pageId="13" pageNumber="12">Pahang</collectingRegion>
|
||
, Cameron Highlands Mts., Tanah Rata,
|
||
<quantity id="4CFA91BEFFFB890DED14689F81BAFB6C" box="[559,719,1153,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="13" pageNumber="12" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">1000–1500m</quantity>
|
||
,
|
||
<date id="FFBC1A9BFFFB890DEDE0689C8019FB6F" box="[731,876,1154,1178]" pageId="13" pageNumber="12" value="1997-04-02" valueMax="1997-04-09" valueMin="1997-04-02">2–9.iv.1997</date>
|
||
, P. Čechovský leg.” / “Collection of the Global, Eucnemid Research Project, (Robert L. Otto)” (green framed white label) / “
|
||
<emphasis id="B976E049FFFB890DEB0368BF87AAFB4C" bold="true" box="[1080,1247,1185,1209]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54B982F9FFFB890DEB0368BF87AFFB4C" box="[1080,1242,1185,1209]" pageId="13" pageNumber="12" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEBD768BC86ECFB4C" authorityName="Otto. Following" authorityYear="2016" box="[1260,1433,1186,1209]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEBD768BC86ECFB4C" box="[1260,1433,1186,1209]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0247D8FFFB890DEFFB68DC82ECFB2F" authority=", Otto" authorityName=", Otto" authorityYear="2017" box="[192,409,1218,1242]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="cuneiformis">
|
||
<emphasis id="B976E049FFFB890DEFFB68DC8225FB2F" box="[192,336,1218,1242]" italics="true" pageId="13" pageNumber="12">cuneiformis</emphasis>
|
||
, Otto
|
||
</taxonomicName>
|
||
, det. R.L. Otto, 2015” (
|
||
<specimenCount id="9D04F7D2FFFB890DED8968DC81B3FB2C" box="[690,710,1218,1241]" pageId="13" pageNumber="12" type="female">♀</specimenCount>
|
||
handwritten behind species name on label) [yellow printed label].
|
||
<typeStatus id="54B982F9FFFB890DEE2868FC82E5FB0F" box="[275,400,1250,1274]" pageId="13" pageNumber="12" type="paratype">Paratypes</typeStatus>
|
||
are retained in
|
||
<collectionCode id="ED13A49EFFFB890DED6C68FC81DDFB0F" box="[599,680,1250,1274]" pageId="13" pageNumber="12">GERP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186FD0FFFB890DEFFB693F87F5FAAF" pageId="13" pageNumber="12" type="distribution">
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB693F87F5FAAF" blockId="13.[192,1441,1313,1370]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB693F8201FACC" bold="true" box="[192,372,1313,1337]" pageId="13" pageNumber="12">Distribution.</emphasis>
|
||
This species is known from two localities; one of which is the mountainous region on Peninsular
|
||
<collectingCountry id="F3157CCBFFFB890DEE74695C82B5FAAF" box="[335,448,1346,1370]" name="Malaysia" pageId="13" pageNumber="12">Malaysia</collectingCountry>
|
||
and the second, a mountain range on Bornean
|
||
<collectingCountry id="F3157CCBFFFB890DEB3C695C8709FAAF" box="[1031,1148,1346,1370]" name="Malaysia" pageId="13" pageNumber="12">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186FD0FFFB890DEFFB699F8286FA2F" pageId="13" pageNumber="12" type="biology_ecology">
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB699F8286FA2F" blockId="13.[192,1440,1409,1498]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB699F8244FA6C" bold="true" box="[192,305,1409,1433]" pageId="13" pageNumber="12">Biology.</emphasis>
|
||
One specimen was taken in the low to mid-elevation tropical highland forest on the peninsula. Two specimens were taken from a montane forest near
|
||
<collectingRegion id="49C6F2B9FFFB890DEC5E69BC80C6FA4F" box="[869,947,1442,1466]" country="Malaysia" name="Sabah" pageId="13" pageNumber="12">Sabah</collectingRegion>
|
||
in the northern areas of Borneo. Larvae and pupae are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186FD0FFFB890DEFFB6A1C8765F9CF" pageId="13" pageNumber="12" type="etymology">
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB6A1C8765F9CF" blockId="13.[192,1440,1538,1594]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB6A1C8229F9EF" bold="true" box="[192,348,1538,1562]" pageId="13" pageNumber="12">Etymology.</emphasis>
|
||
Specific epithet is derived from the overall general form of the new species, that being strongly cuneiform, compared with two other congeners of the group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186FD0FFFB890DEFFB6A7C8299F88C" pageId="13" pageNumber="12" type="discussion">
|
||
<paragraph id="8BBD3C5BFFFB890DEFFB6A7C8299F88C" blockId="13.[192,1441,1634,1914]" pageId="13" pageNumber="12">
|
||
<emphasis id="B976E049FFFB890DEFFB6A7C8231F98F" bold="true" box="[192,324,1634,1658]" pageId="13" pageNumber="12">Remarks.</emphasis>
|
||
Although many groups within the tribe exhibit some form of sexual dimorphism among each species; none is more extreme as
|
||
<taxonomicName id="4C0247D8FFFB890DED686A9C8076F96C" authorityName="Otto. Following" authorityYear="2016" box="[595,771,1666,1689]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DED686A9C8076F96C" box="[595,771,1666,1689]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
, with stark differences in antennal structures and last visible ventrite between males and females in the recently described group. Males are highly diagnostic. Females are a bit more difficult to diagnose. Female
|
||
<taxonomicName id="4C0247D8FFFB890DEC786ADC8086F92C" authorityName="Otto. Following" authorityYear="2016" box="[835,1011,1730,1753]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEC786ADC8086F92C" box="[835,1011,1730,1753]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
are very similar to
|
||
<taxonomicName id="4C0247D8FFFB890DEBD86ADC861FF92F" authorityName="Berthold" authorityYear="1827" box="[1251,1386,1730,1754]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEBD86ADC861FF92F" box="[1251,1386,1730,1754]" italics="true" pageId="13" pageNumber="12">Nematodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0247D8FFFB890DEFFB6AFC8208F90F" authorityName="Bonvouloir" authorityYear="1871" box="[192,381,1762,1786]" class="Insecta" family="Eucnemidae" genus="Trigonopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEFFB6AFC8208F90F" box="[192,381,1762,1786]" italics="true" pageId="13" pageNumber="12">Trigonopleurus</emphasis>
|
||
</taxonomicName>
|
||
. Female
|
||
<taxonomicName id="4C0247D8FFFB890DEED56AFC81EBF90C" authorityName="Otto. Following" authorityYear="2016" box="[494,670,1762,1785]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEED56AFC81EBF90C" box="[494,670,1762,1785]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
differs from any
|
||
<taxonomicName id="4C0247D8FFFB890DEC4B6AFC8723F90F" box="[880,1110,1762,1786]" class="Insecta" family="Eucnemidae" genus="Nematodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B976E049FFFB890DEC4B6AFC8082F90F" box="[880,1015,1762,1786]" italics="true" pageId="13" pageNumber="12">Nematodes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
based on the metathoracic episterna, apically wide in
|
||
<taxonomicName id="4C0247D8FFFB890DED296B1C81B7F8EC" authorityName="Otto. Following" authorityYear="2016" box="[530,706,1794,1817]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DED296B1C81B7F8EC" box="[530,706,1794,1817]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
and parallel-sided in
|
||
<taxonomicName id="4C0247D8FFFB890DECED6B1C8728F8EF" authorityName="Berthold" authorityYear="1827" box="[982,1117,1794,1818]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DECED6B1C8728F8EF" box="[982,1117,1794,1818]" italics="true" pageId="13" pageNumber="12">Nematodes</emphasis>
|
||
</taxonomicName>
|
||
. Female
|
||
<taxonomicName id="4C0247D8FFFB890DEBE86B1C86F6F8EC" authorityName="Otto. Following" authorityYear="2016" box="[1235,1411,1794,1817]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEBE86B1C86F6F8EC" box="[1235,1411,1794,1817]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
is further diagnosed from
|
||
<taxonomicName id="4C0247D8FFFB890DEED36B3C81D0F8CF" authorityName="Bonvouloir" authorityYear="1871" box="[488,677,1826,1850]" class="Insecta" family="Eucnemidae" genus="Trigonopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEED36B3C81D0F8CF" box="[488,677,1826,1850]" italics="true" pageId="13" pageNumber="12">Trigonopleurus</emphasis>
|
||
</taxonomicName>
|
||
based on the combined lengths of antennomeres IV and V in relation to antennomere III; IV and V together shorter than III in
|
||
<taxonomicName id="4C0247D8FFFB890DECD76B5C87DCF8AF" authorityName="Bonvouloir" authorityYear="1871" box="[1004,1193,1858,1882]" class="Insecta" family="Eucnemidae" genus="Trigonopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DECD76B5C87DCF8AF" box="[1004,1193,1858,1882]" italics="true" pageId="13" pageNumber="12">Trigonopleurus</emphasis>
|
||
</taxonomicName>
|
||
versus as long as or longer in
|
||
<taxonomicName id="4C0247D8FFFB890DEE0E6B7C8290F88C" authorityName="Otto. Following" authorityYear="2016" box="[309,485,1890,1913]" class="Insecta" family="Eucnemidae" genus="Miruantennus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976E049FFFB890DEE0E6B7C8290F88C" box="[309,485,1890,1913]" italics="true" pageId="13" pageNumber="12">Miruantennus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |