139 lines
17 KiB
XML
139 lines
17 KiB
XML
<document id="47D574C746A8665D754FB92A55F0ACEB" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFD8FFDB6194FE620012F19E" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus trituberculatus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="37" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="004E0E0119838F55ADAFD38CC70B2F1F">
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<mods:title id="A891C615E0C48FB835734A15105FA42F">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="2CF006B16D188A6E6DC4F208EB0BD40D" type="personal">
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<mods:roleTerm id="BA3A8C3BB9F8867893BA2DE8FD3E0D76">Author</mods:roleTerm>
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<mods:namePart id="A18CD6D79EC68E634EB87278AA1A90FC">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="9C6E827927F37230BF9A9B95BDC35701">text</mods:typeOfResource>
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<mods:title id="0068F9746F9E5EA0574BF4674D162FE9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0A0A1150BCBAF887FF66168B1BC865C5">
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<mods:date id="BD3DA3B6C86E728FE1F7B96076268C05">2021</mods:date>
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<mods:detail id="3BD494B643888D5E38821989494C3E9C" type="pubDate">
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<mods:number id="F959B489E74F916DE12B11C2D94CA4B0">2021-06-28</mods:number>
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<mods:detail id="27193E974302B1B14999FC8FE7AE429C" type="volume">
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<mods:number id="050CB9F8B4CD4802452A08AF90A4F00B">4994</mods:number>
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<mods:detail id="EF51986CF3F6250718E6EB54885EB71B" type="issue">
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<mods:number id="6F5ECF1FB36DD5B92AD35DA6574A4DBC">1</mods:number>
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<mods:start id="6B6F0E065746B6F914B9B875C836D437">1</mods:start>
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<mods:end id="EB4142C25FBE4F89385F0DE42AF6D959">93</mods:end>
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<mods:classification id="8E8A7E2DC77AB6BDD428000F502FD59A">journal article</mods:classification>
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<mods:identifier id="412E8687A5757EC4B6101343B23AD802" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="7BEFF0AF2680E07B0725554BBFD10503" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="85498558DA3EB721D23BAD3C35CF3DF7" type="Zenodo-Dep">5043261</mods:identifier>
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<treatment id="03A187DEFFD8FFDB6194FE620012F19E" ID-DOI="http://doi.org/10.5281/zenodo.5043984" ID-Zenodo-Dep="5043984" LSID="urn:lsid:plazi:treatment:03A187DEFFD8FFDB6194FE620012F19E" httpUri="http://treatment.plazi.org/id/03A187DEFFD8FFDB6194FE620012F19E" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection id="C3126543FFD8FFDB6194FE620620F6AB" box="[151,639,402,429]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BB736C8FFD8FFDB6194FE620620F6AB" blockId="36.[151,639,402,465]" box="[151,639,402,429]" pageId="36" pageNumber="37">
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<heading id="D0FF81A4FFD8FFDB6194FE620620F6AB" bold="true" box="[151,639,402,429]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
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<emphasis id="B97CEADAFFD8FFDB6194FE620620F6AB" bold="true" box="[151,639,402,429]" pageId="36" pageNumber="37">
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<taxonomicName id="4C084D4BFFD8FFDB6194FE6205BDF6AB" ID-CoL="8Y4RQ" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,482,402,429]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="trituberculatus" status="sp. nov.">
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<emphasis id="B97CEADAFFD8FFDB6194FE620542F6AA" bold="true" box="[151,285,402,429]" italics="true" pageId="36" pageNumber="37">Hydrochus</emphasis>
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trituberculatus
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFD8FFDB60ECFE630620F6AB" box="[495,639,402,429]" pageId="36" pageNumber="37" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD8FFDB6194FE470568F6D6" box="[151,311,438,465]" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BB736C8FFD8FFDB6194FE470568F6D6" blockId="36.[151,639,402,465]" box="[151,311,438,465]" pageId="36" pageNumber="37">
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<figureCitation id="13332A4DFFD8FFDB6194FE4704BAF6D6" box="[151,229,439,465]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="36" pageNumber="37">Figs. 2</figureCitation>
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,
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<figureCitation id="13332A4DFFD8FFDB61F2FE470550F6D6" box="[241,271,439,465]" captionStart="FIGURES 30–31" captionStartId="67.[151,264,1944,1968]" captionTargetBox="[163,1425,201,1882]" captionTargetId="figure-15@67.[151,1435,181,1920]" captionTargetPageId="67" captionText="FIGURES 30–31. Habitus and male genitalia of holotypes. 30. Hydrochus canalis n. sp. 31. Hydrochus trituberculatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043314" httpUri="https://zenodo.org/record/5043314/files/figure.png" pageId="36" pageNumber="37">31</figureCitation>
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,
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<figureCitation id="13332A4DFFD8FFDB6018FE460568F6D6" box="[283,311,438,465]" captionStart="FIGURES 55–56" captionStartId="80.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1920]" captionTargetId="figure-28@80.[165,1422,181,1920]" captionTargetPageId="80" captionText="FIGURES 55–56. Geographical distributions. 55. Endemics and near endemics. 56. H. bicarinatus and H. merkli." figureDoi="http://doi.org/10.5281/zenodo.5043352" httpUri="https://zenodo.org/record/5043352/files/figure.png" pageId="36" pageNumber="37">55</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD8FFDB6194FE0F0574F53A" pageId="36" pageNumber="37" type="materials_examined">
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<paragraph id="8BB736C8FFD8FFDB6194FE0F0574F53A" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">
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<emphasis id="B97CEADAFFD8FFDB6194FE0F0514F51F" bold="true" box="[151,331,510,537]" pageId="36" pageNumber="37">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFD8FFDB6056FE0F0574F53A" collectingDate="1956-05-23" collectionCode="NHMLAC" collectorName="E. Formoso & F. S. Truxal" country="Brazil" location="Goias" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
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<emphasis id="B97CEADAFFD8FFDB6056FE0F062AF51F" bold="true" box="[341,629,510,537]" pageId="36" pageNumber="37">
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<typeStatus id="54B3886AFFD8FFDB6056FE0F059DF51E" box="[341,450,511,537]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="F31F7658FFD8FFDB632FFE0E062AF51F" box="[556,629,510,536]" name="Brazil" pageId="36" pageNumber="37">Brazil</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="49CCF82AFFD8FFDB6384FE0F0694F51E" box="[647,715,511,537]" country="Brazil" name="Goias" pageId="36" pageNumber="37">Goias</collectingRegion>
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, “24 kil.
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<collectorName id="26FD531EFFD8FFDB6238FE0F079CF51E" box="[827,963,511,537]" pageId="36" pageNumber="37">E. Formoso</collectorName>
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, Go., Brazil
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<date id="FFB61008FFD8FFDB6558FE0F00A2F51E" box="[1115,1277,510,537]" pageId="36" pageNumber="37" value="1956-05-23">
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<collectingDate id="EFF2E9E0FFD8FFDB6558FE0F00A2F51E" box="[1115,1277,510,537]" pageId="36" pageNumber="37" value="1956-05-23">May 23, 1956</collectingDate>
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</date>
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<collectorName id="26FD531EFFD8FFDB6404FE0F01CCF51E" box="[1287,1427,511,537]" pageId="36" pageNumber="37">F. S. Truxal</collectorName>
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” (
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<collectionCode id="ED19AE0DFFD8FFDB619CFDD30540F53A" box="[159,287,547,573]" pageId="36" pageNumber="37">NHMLAC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD8FFDB61C4FDB60789F5CA" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="8BB736C8FFD8FFDB61C4FDB60789F5CA" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">
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<emphasis id="B97CEADAFFD8FFDB61C4FDB6058BF567" bold="true" box="[199,468,582,608]" pageId="36" pageNumber="37">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of moderate size (ca.
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<quantity id="4CF09B2DFFD8FFDB6468FDB70498F582" metricMagnitude="-3" metricUnit="m" metricValue="3.48" pageId="36" pageNumber="37" unit="mm" value="3.48">3.48 mm</quantity>
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), the dark grey to black color, the moderately deep pronotal depressions, the moderately low and relatively impunctate pronotal reliefs, the presence of three calluses on each elytron, and the male genitalia (
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<figureCitation id="13332A4DFFD8FFDB65D3FD7F017BF5AE" box="[1232,1316,655,681]" captionStart="FIGURES 30–31" captionStartId="67.[151,264,1944,1968]" captionTargetBox="[163,1425,201,1882]" captionTargetId="figure-15@67.[151,1435,181,1920]" captionTargetPageId="67" captionText="FIGURES 30–31. Habitus and male genitalia of holotypes. 30. Hydrochus canalis n. sp. 31. Hydrochus trituberculatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043314" httpUri="https://zenodo.org/record/5043314/files/figure.png" pageId="36" pageNumber="37">Fig. 31</figureCitation>
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; described below). However, reliable determinations will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD8FFDB61C4FD2606C3F172" pageId="36" pageNumber="37" type="description">
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<paragraph id="8BB736C8FFD8FFDB61C4FD260508F4EA" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">
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<emphasis id="B97CEADAFFD8FFDB61C4FD26050EF5F7" bold="true" box="[199,337,726,752]" pageId="36" pageNumber="37">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFD8FFDB60A7FD270659F5F6" box="[420,518,727,753]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.48/1.02; head width 0.85; pronotum l/w 0.82/0.89; PA 0.84; PB 0.62; elytra 2.18/1.02. Habitus as illustrated (
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<figureCitation id="13332A4DFFD8FFDB654CFD0B00FBF412" box="[1103,1188,763,789]" captionStart="FIGURES 30–31" captionStartId="67.[151,264,1944,1968]" captionTargetBox="[163,1425,201,1882]" captionTargetId="figure-15@67.[151,1435,181,1920]" captionTargetPageId="67" captionText="FIGURES 30–31. Habitus and male genitalia of holotypes. 30. Hydrochus canalis n. sp. 31. Hydrochus trituberculatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043314" httpUri="https://zenodo.org/record/5043314/files/figure.png" pageId="36" pageNumber="37">Fig. 31</figureCitation>
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). Body size moderate (ca.
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<quantity id="4CF09B2DFFD8FFDB61CAFCEF056CF43E" box="[201,307,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.48" pageId="36" pageNumber="37" unit="mm" value="3.48">3.48 mm</quantity>
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). Dorsum grey to black, with slight to moderately strong iridescence, elytra with black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
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<superScript id="7C7D9B80FFD8FFDB6384FC9606CBF473" attach="left" box="[647,660,870,884]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFD8FFDB657AFC9600D9F473" attach="left" box="[1145,1158,870,884]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFD8FFDB6012FC7A0541F49F" attach="left" box="[273,286,906,920]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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markedly raised, short, only ca. 3 punctures on each side; each elytron also with another callus on 5
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<superScript id="7C7D9B80FFD8FFDB648AFC7A01C9F49F" attach="right" box="[1417,1430,906,920]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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, anterior to usual callus, and another callus on 7
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<superScript id="7C7D9B80FFD8FFDB63ACFC5E06E3F4BB" attach="left" box="[687,700,942,956]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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interseries, ca. opposite middle of distance between two calluses on 5
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<superScript id="7C7D9B80FFD8FFDB61CBFC22048AF4E7" attach="left" box="[200,213,978,992]" fontSize="6" pageId="36" pageNumber="37">th</superScript>
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interseries.
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</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4FC070045F35E" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">Pronotum very slightly wider than long (ratio ca. 25/23), widest slightly behind anterior margin, narrowed at base, sides sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep, much more densely punctate than low reliefs between depressions, latter almost entirely impunctate.</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4FB930746F3A6" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">Elytra with spaces between strial punctures ca. 1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with angulation.</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4FB5B068FF3EE" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">Ventral characters: Mentum densely punctate, with basomedian fovea; submentum very finely sparsely punctate, without foveae; metaventrite markedly tumid.</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4FB0300E5F10E" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">
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Male genitalia (
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<figureCitation id="13332A4DFFD8FFDB607AFB030594F20A" box="[377,459,1267,1293]" captionStart="FIGURES 30–31" captionStartId="67.[151,264,1944,1968]" captionTargetBox="[163,1425,201,1882]" captionTargetId="figure-15@67.[151,1435,181,1920]" captionTargetPageId="67" captionText="FIGURES 30–31. Habitus and male genitalia of holotypes. 30. Hydrochus canalis n. sp. 31. Hydrochus trituberculatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043314" httpUri="https://zenodo.org/record/5043314/files/figure.png" pageId="36" pageNumber="37">Fig. 31</figureCitation>
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) general characters: genitalia very narrow, very elongate, length ca. 8x greatest width, wider in dorsal/ventral views than lateral (ratio ca. 2/1), basal piece as long as parameres; paramere tips not extending beyond aedeagus tip, tips arcuate toward midline, moderately sharply pointed, plsa absent; in lateral view parameres arcuate toward ventrad in distal 1/2, width ca. equal dorsal/ventral width, moderately sharply pointed apically; pseudoparameres very short, narrow, spike-like; aedeagus very narrow, almost parallel-sided for entire length, slightly widened apically and at base, the latter strongly sclerotized but not separate lobe; basal piece in dorsal/ventral views very narrow, sides very weakly arcuate if at all, length nearly four times width, in lateral view even narrower, very slightly arcuate, sides nearly parallel, orifice small, slightly asymmetrical.
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</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4F9E301CAF12A" blockId="36.[151,1437,510,1689]" box="[199,1429,1555,1581]" pageId="36" pageNumber="37">Dorsal surface: adtl absent; pdmm widely separated, only contiguous with aedeagus at very base; adbl absent.</paragraph>
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<paragraph id="8BB736C8FFD8FFDB61C4F9C706C3F172" blockId="36.[151,1437,510,1689]" pageId="36" pageNumber="37">Ventral surface: avtl subapical, delimiting gonopore; pvmm separation similar to pdmm in distal ½, parameres wider in proximal ½, overlapping alm at base.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD8FFDB61C4F98F0012F19E" box="[199,1101,1663,1689]" pageId="36" pageNumber="37" type="etymology">
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<paragraph id="8BB736C8FFD8FFDB61C4F98F0012F19E" blockId="36.[151,1437,510,1689]" box="[199,1101,1663,1689]" pageId="36" pageNumber="37">
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<emphasis id="B97CEADAFFD8FFDB61C4F98F0517F19E" bold="true" box="[199,328,1663,1689]" pageId="36" pageNumber="37">Etymology</emphasis>
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. Named in reference to the three prominent calluses on each elytron.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |