treatments-xml/data/03/A1/87/03A187DEFFCFFFCC6194FED607F1F454.xml
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<document id="0C42A8A9687CDD09A70DCAB728942BE9" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFCFFFCC6194FED607F1F454" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus soekhnandanae Makhan" docType="treatment" docVersion="3" lastPageNumber="52" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="52" updateTime="1698978858223" updateUser="plazi">
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<mods:title id="824E2B460E047A88D32EE1C5850AD8B9">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="ACA38BE4B00CBFBA6D819C97342DB2D4">Perkins, Philip D.</mods:namePart>
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<mods:date id="FC2BEE2F5DF2B9061B713C38CF76C8FC">2021</mods:date>
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<mods:number id="AB2FFC590D1B931439551387245A3C1B">4994</mods:number>
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<mods:identifier id="6ACCD137A7F41D796D3F33D347B72AC6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A187DEFFCFFFCC6194FED607F1F454" LSID="urn:lsid:plazi:treatment:03A187DEFFCFFFCC6194FED607F1F454" httpUri="http://treatment.plazi.org/id/03A187DEFFCFFFCC6194FED607F1F454" lastPageNumber="52" pageId="51" pageNumber="52">
<subSubSection id="C3126543FFCFFFCC6194FED6060BF646" box="[151,596,294,321]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BB736C8FFCFFFCC6194FED6060BF646" blockId="51.[151,596,294,357]" box="[151,596,294,321]" pageId="51" pageNumber="52">
<heading id="D0FF81A4FFCFFFCC6194FED6060BF646" bold="true" box="[151,596,294,321]" fontSize="11" level="2" pageId="51" pageNumber="52" reason="3">
<taxonomicName id="4C084D4BFFCFFFCC6194FED6060BF646" ID-CoL="3N6C8" authority="Makhan" authorityName="Makhan" box="[151,596,294,321]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B97CEADAFFCFFFCC6194FED6060BF646" bold="true" box="[151,596,294,321]" pageId="51" pageNumber="52">
<emphasis id="B97CEADAFFCFFFCC6194FED605BCF646" bold="true" box="[151,483,294,321]" italics="true" pageId="51" pageNumber="52">Hydrochus soekhnandanae</emphasis>
Makhan
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFCFFFCC6194FEBB0552F662" box="[151,269,331,357]" pageId="51" pageNumber="52" type="description">
<paragraph id="8BB736C8FFCFFFCC6194FEBB0552F662" blockId="51.[151,596,294,357]" box="[151,269,331,357]" pageId="51" pageNumber="52">
<figureCitation id="13332A4DFFCFFFCC6194FEBB04BAF662" box="[151,229,331,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1944,1968]" captionTargetBox="[174,1420,192,1883]" captionTargetId="figure-15@9.[156,1431,181,1920]" captionTargetPageId="9" captionText="FIGURE 4. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043272" httpUri="https://zenodo.org/record/5043272/files/figure.png" pageId="51" pageNumber="52">Figs. 4</figureCitation>
,
<figureCitation id="13332A4DFFCFFFCC61F2FEBB0552F662" box="[241,269,331,357]" captionStart="FIGURE 71" captionStartId="90.[151,250,1796,1820]" captionTargetBox="[166,1425,227,1738]" captionTargetId="figure-28@90.[151,1435,181,1773]" captionTargetPageId="90" captionText="FIGURE 71. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043384" httpUri="https://zenodo.org/record/5043384/files/figure.png" pageId="51" pageNumber="52">71</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFCFFFCC6194FE7F071DF551" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="8BB736C8FFCFFFCC6194FE7F063DF6A0" blockId="51.[151,1207,399,454]" box="[151,610,399,423]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB1811E6FFCFFFCC6194FE7F063DF6A0" box="[151,610,399,423]" pageId="51" pageNumber="52">
<treatmentCitation id="0AA910D9FFCFFFCC6194FE7F063DF6A0" author="Makhan, D." box="[151,610,399,423]" page="98" pageId="51" pageNumber="52" year="1992">
<taxonomicName id="4C084D4BFFCFFFCC6194FE7F063DF6A0" ID-CoL="3N6C8" authority="Makhan 1992: 98" authorityName="Makhan" authorityPageNumber="98" authorityYear="1992" box="[151,610,399,423]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B97CEADAFFCFFFCC6194FE7F05F5F6A0" box="[151,426,399,423]" italics="true" pageId="51" pageNumber="52">Hydrochus soekhnandanae</emphasis>
<bibRefCitation id="EF994B39FFCFFFCC60B3FE7F063DF6A0" author="Makhan, D." box="[432,610,399,423]" pageId="51" pageNumber="52" pagination="95 - 103" refId="ref50247" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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</paragraph>
<paragraph id="8BB736C8FFCFFFCC6194FE5E00E8F6C1" blockId="51.[151,1207,399,454]" box="[151,1207,430,454]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB1811E6FFCFFFCC6194FE5E00E8F6C1" box="[151,1207,430,454]" pageId="51" pageNumber="52">
<taxonomicName id="4C084D4BFFCFFFCC6194FE5E0584F6C1" ID-CoL="6MK94" authority="Orchymont" authorityName="Orchymont" box="[151,475,430,454]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B97CEADAFFCFFFCC6194FE5E0501F6C1" box="[151,350,430,454]" italics="true" pageId="51" pageNumber="52">Hydrochus pupillus</emphasis>
Orchymont
</taxonomicName>
:
<treatmentCitation id="0AA910D9FFCFFFCC60E7FE5E06C2F6C1" author="Oliva, A." box="[484,669,430,454]" page="331" pageId="51" pageNumber="52" year="1996">
<bibRefCitation id="EF994B39FFCFFFCC60E7FE5E06C2F6C1" author="Oliva, A." box="[484,669,430,454]" pageId="51" pageNumber="52" pagination="301 - 341" refId="ref50713" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996: 331)</bibRefCitation>
</treatmentCitation>
; incorrect synonymy by
<bibRefCitation id="EF994B39FFCFFFCC6294FE5E0042F6C1" author="Oliva, A." box="[919,1053,430,454]" pageId="51" pageNumber="52" pagination="301 - 341" refId="ref50713" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
;
<bibRefCitation id="EF994B39FFCFFFCC6524FE5E00E8F6C1" author="Perkins, P. D." box="[1063,1207,430,454]" pageId="51" pageNumber="52" pagination="1 - 59" refId="ref50913" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins 2019c</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736C8FFCFFFCC6194FE04071DF551" blockId="51.[151,1437,500,851]" pageId="51" pageNumber="52">
Originally described from
<collectingCountry id="F31F7658FFCFFFCC60C3FE040670F509" box="[448,559,500,526]" name="Suriname" pageId="51" pageNumber="52">Suriname</collectingCountry>
. Additional locality and deposition data also published for
<collectingCountry id="F31F7658FFCFFFCC65BDFE04016EF509" box="[1214,1329,500,526]" name="Venezuela" pageId="51" pageNumber="52">Venezuela</collectingCountry>
(
<bibRefCitation id="EF994B39FFCFFFCC6443FE040481F535" author="Perkins, P. D." pageId="51" pageNumber="52" pagination="1 - 59" refId="ref50913" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins, 2019c</bibRefCitation>
) and
<collectingCountry id="F31F7658FFCFFFCC601FFDE9052AF534" box="[284,373,537,563]" name="Panama" pageId="51" pageNumber="52">Panama</collectingCountry>
(
<bibRefCitation id="EF994B39FFCFFFCC6080FDE8067CF535" author="Perkins, P. D." box="[387,547,536,562]" pageId="51" pageNumber="52" pagination="459 - 503" refId="ref51126" refString="Perkins, P. D. (2021) Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae). Zootaxa, 4974 (3), 459 - 503. https: // doi. org / 10.11646 / zootaxa. 4974.3.2" type="journal article" year="2021">Perkins, 2021</bibRefCitation>
). Habitus and male genitalia images are given in
<bibRefCitation id="EF994B39FFCFFFCC654FFDE8015EF535" author="Perkins, P. D." box="[1100,1281,536,562]" pageId="51" pageNumber="52" pagination="1 - 59" refId="ref50913" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins (2019c)</bibRefCitation>
. New locality and deposition data for
<collectingCountry id="F31F7658FFCFFFCC60A3FDCC05BBF551" box="[416,484,572,598]" name="Brazil" pageId="51" pageNumber="52">Brazil</collectingCountry>
and
<collectingCountry id="F31F7658FFCFFFCC6319FDCD06DDF550" box="[538,642,573,599]" name="Paraguay" pageId="51" pageNumber="52">Paraguay</collectingCountry>
are given below.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFCFFFCC61C4FD900552F40C" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BB736C8FFCFFFCC61C4FD900552F40C" blockId="51.[151,1437,500,851]" pageId="51" pageNumber="52">
<emphasis id="B97CEADAFFCFFFCC61C4FD9005B8F57D" bold="true" box="[199,487,608,634]" pageId="51" pageNumber="52">Material examined (14):</emphasis>
<materialsCitation id="3B603C95FFCFFFCC60EDFD900115F57C" box="[494,1354,608,635]" collectingDate="1969-07-03" collectionCode="NMNH" collectorName="E. Itabuna &amp; P. &amp; P. Spangler" country="Brazil" location="Bahia" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Bahia">
<emphasis id="B97CEADAFFCFFFCC60EDFD900668F57D" bold="true" box="[494,567,608,634]" pageId="51" pageNumber="52">
<collectingCountry id="F31F7658FFCFFFCC60EDFD900668F57D" box="[494,567,608,634]" name="Brazil" pageId="51" pageNumber="52">Brazil</collectingCountry>
</emphasis>
:
<collectingRegion id="49CCF82AFFCFFFCC6346FD9006D6F57D" box="[581,649,608,634]" country="Brazil" name="Bahia" pageId="51" pageNumber="52">Bahia</collectingRegion>
,
<quantity id="4CF09B2DFFCFFFCC6397FD900688F57C" box="[660,727,608,635]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="km" value="15.0">15km</quantity>
.
<collectorName id="26FD531EFFCFFFCC63E2FD910708F57D" box="[737,855,608,635]" pageId="51" pageNumber="52">E. Itabuna</collectorName>
,
<date id="FFB61008FFCFFFCC6262FD90078AF57D" box="[865,981,608,634]" pageId="51" pageNumber="52" value="1969-07-03">
<collectingDate id="EFF2E9E0FFCFFFCC6262FD90078AF57D" box="[865,981,608,634]" pageId="51" pageNumber="52" value="1969-07-03">3.vii.1969</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFCFFFCC6513FD91009BF57D" box="[1040,1220,608,635]" pageId="51" pageNumber="52">P. &amp; P. Spangler</collectorName>
(1
<collectionCode id="ED19AE0DFFCFFFCC65E4FD91011DF57C" box="[1255,1346,609,635]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="51" pageNumber="52" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFCFFFCC6456FD9107AEF5C5" collectingDate="2018-02-24" collectionCode="INPA, SEMC" collectorName="Benetti &amp; Team, BR" country="Brazil" elevation="706" latitude="-11.65164" location="Barragem de Angelim" longLatPrecision="1" longitude="-40.95309" municipality="Morro do Chapeu" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Bahia">
<collectingMunicipality id="6BD3ACB2FFCFFFCC6456FD91054BF599" pageId="51" pageNumber="52">Morro do Chapéu</collectingMunicipality>
,
<location id="8ED76013FFCFFFCC6023FD750645F599" LSID="urn:lsid:plazi:treatment:03A187DEFFCFFFCC6194FED607F1F454:8ED76013FFCFFFCC6023FD750645F599" box="[288,538,644,671]" country="Brazil" latitude="-11.65164" longLatPrecision="1" longitude="-40.95309" municipality="Morro do Chapeu" name="Barragem de Angelim" pageId="51" pageNumber="52" stateProvince="Bahia">Barragem de Angelim</location>
on BA-422,
<locationDeviation id="9A052FEAFFCFFFCC63B2FD75071FF598" box="[689,832,644,671]" location="Ventura" pageId="51" pageNumber="52">E of Ventura</locationDeviation>
<geoCoordinate id="EE3C500FFFCFFFCC624AFD7407E5F598" box="[841,954,644,671]" degrees="11.65164" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="1" value="-11.65164">-11.65164</geoCoordinate>
<geoCoordinate id="EE3C500FFFCFFFCC62C0FD740067F598" box="[963,1080,644,671]" degrees="40.95309" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="1" value="-40.95309">-40.95309</geoCoordinate>
,
<quantity id="4CF09B2DFFCFFFCC6547FD7400D8F599" box="[1092,1159,644,670]" metricMagnitude="2" metricUnit="m" metricValue="7.06" pageId="51" pageNumber="52" unit="m" value="706.0">
<elevation id="0025D1FBFFCFFFCC6547FD7400D8F599" box="[1092,1159,644,670]" metricMagnitude="2" metricUnit="m" metricValue="7.06" pageId="51" pageNumber="52" unit="m" value="706.0">706m</elevation>
</quantity>
,
<date id="FFB61008FFCFFFCC6590FD740157F599" box="[1171,1288,644,670]" pageId="51" pageNumber="52" value="2018-02-24">
<collectingDate id="EFF2E9E0FFCFFFCC6590FD740157F599" box="[1171,1288,644,670]" pageId="51" pageNumber="52" value="2018-02-24">24.ii.2018</collectingDate>
</date>
, reservoir by road, lots of vegetation,
<collectorName id="26FD531EFFCFFFCC60AAFD5805A5F5C5" box="[425,506,680,706]" pageId="51" pageNumber="52">Benetti</collectorName>
&amp;
<collectorName id="26FD531EFFCFFFCC631DFD5906CEF5C5" box="[542,657,680,707]" pageId="51" pageNumber="52">Team, BR</collectorName>
18-0224-04B (9
<collectionCode id="ED19AE0DFFCFFFCC6248FD5807D2F5C5" box="[843,909,680,706]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="51" pageNumber="52">INPA</collectionCode>
,
<collectionCode id="ED19AE0DFFCFFFCC629BFD5807B6F5C5" box="[920,1001,680,706]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="51" pageNumber="52" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B603C95FFCFFFCC62FFFD5807C8F5E0" collectingDate="2004-11-24" collectingDateMax="2004-11-25" collectingDateMin="2004-11-24" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.55" location="Estancia Dimas" longLatPrecision="1246" longitude="-55.216667" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Alto Parana">
<emphasis id="B97CEADAFFCFFFCC62FFFD580030F5C5" bold="true" box="[1020,1135,680,706]" pageId="51" pageNumber="52">
<collectingCountry id="F31F7658FFCFFFCC62FFFD580030F5C5" box="[1020,1135,680,706]" name="Paraguay" pageId="51" pageNumber="52">Paraguay</collectingCountry>
</emphasis>
:
<collectingRegion id="49CCF82AFFCFFFCC657EFD58015AF5C4" box="[1149,1285,680,707]" country="Paraguay" name="Alto Parana" pageId="51" pageNumber="52">Alto Parana</collectingRegion>
,
<location id="8ED76013FFCFFFCC6412FD580498F5E0" LSID="urn:lsid:plazi:treatment:03A187DEFFCFFFCC6194FED607F1F454:8ED76013FFCFFFCC6412FD580498F5E0" country="Paraguay" latitude="-25.55" longLatPrecision="1246" longitude="-55.216667" name="Estancia Dimas" pageId="51" pageNumber="52" stateProvince="Alto Parana">Estancia Dimas</location>
, (
<geoCoordinate id="EE3C500FFFCFFFCC61D9FD3C0566F5E0" box="[218,313,716,743]" degrees="25" direction="south" minutes="33" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="-25.55">25°33S</geoCoordinate>
,
<geoCoordinate id="EE3C500FFFCFFFCC6040FD3C05F5F5E0" box="[323,426,716,743]" degrees="55" direction="west" minutes="13" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="-55.216667">55°13W</geoCoordinate>
),
<date id="FFB61008FFCFFFCC60C3FD3C063BF5E0" box="[448,612,716,743]" pageId="51" pageNumber="52" value="2004-11-24" valueMax="2004-11-25" valueMin="2004-11-24">
<collectingDate id="EFF2E9E0FFCFFFCC60C3FD3C063BF5E0" box="[448,612,716,743]" pageId="51" pageNumber="52" value="2004-11-24" valueMax="2004-11-25" valueMin="2004-11-24">2425.xi.2004</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFCFFFCC639EFD3D077DF5E1" box="[669,802,716,743]" pageId="51" pageNumber="52">U. Drechsel</collectorName>
(1
<collectionCode id="ED19AE0DFFCFFFCC6247FD3D07D3F5E0" box="[836,908,717,743]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="52">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C95FFCFFFCC62A0FD3D0557F40C" collectingDate="2005-02-17" collectingDateMax="2005-02-22" collectingDateMin="2005-02-17" collectionCode="NMW" collectorName="U. Drechsel" country="Paraguay" latitude="-25.55" location="Estancia Dimas" longLatPrecision="1246" longitude="-55.216667" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Alto Parana">
same locality,
<date id="FFB61008FFCFFFCC6542FD3C0080F5E0" box="[1089,1247,716,743]" pageId="51" pageNumber="52" value="2005-02-17" valueMax="2005-02-22" valueMin="2005-02-17">
<collectingDate id="EFF2E9E0FFCFFFCC6542FD3C0080F5E0" box="[1089,1247,716,743]" pageId="51" pageNumber="52" value="2005-02-17" valueMax="2005-02-22" valueMin="2005-02-17">1722.ii.2005</collectingDate>
</date>
, leg.
<collectorName id="26FD531EFFCFFFCC641BFD3D01C2F5E1" box="[1304,1437,716,743]" pageId="51" pageNumber="52">U. Drechsel</collectorName>
(3
<collectionCode id="ED19AE0DFFCFFFCC61B5FD0104A1F40C" box="[182,254,753,779]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="52">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126543FFCFFFCC61C4FCE407F1F454" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8BB736C8FFCFFFCC61C4FCE407F1F454" blockId="51.[151,1437,500,851]" pageId="51" pageNumber="52">
<emphasis id="B97CEADAFFCFFFCC61C4FCE40564F429" bold="true" box="[199,315,788,814]" pageId="51" pageNumber="52">Remarks.</emphasis>
The male genitalia of
<taxonomicName id="4C084D4BFFCFFFCC633AFCE50755F429" box="[569,778,788,814]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B97CEADAFFCFFFCC633AFCE50755F429" box="[569,778,788,814]" italics="true" pageId="51" pageNumber="52">H. soekhnandanae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C084D4BFFCFFFCC6242FCE507E1F429" box="[833,958,788,814]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B97CEADAFFCFFFCC6242FCE507E1F429" box="[833,958,788,814]" italics="true" pageId="51" pageNumber="52">H. pupillus</emphasis>
</taxonomicName>
distinctively differ. Refer to the images of
<taxonomicName id="4C084D4BFFCFFFCC6194FCC90538F455" box="[151,359,824,850]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="soekhnandanae">
<emphasis id="B97CEADAFFCFFFCC6194FCC90538F455" box="[151,359,824,850]" italics="true" pageId="51" pageNumber="52">H. soekhnandanae</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF994B39FFCFFFCC6088FCC80661F455" author="Perkins, P. D." box="[395,574,824,850]" pageId="51" pageNumber="52" pagination="1 - 59" refId="ref50913" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins (2019c)</bibRefCitation>
and
<taxonomicName id="4C084D4BFFCFFFCC6377FCC906AFF455" box="[628,752,824,850]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="pupillus">
<emphasis id="B97CEADAFFCFFFCC6377FCC906AFF455" box="[628,752,824,850]" italics="true" pageId="51" pageNumber="52">H. pupillus</emphasis>
</taxonomicName>
herein (
<figureCitation id="13332A4DFFCFFFCC624EFCC807FEF454" box="[845,929,824,851]" captionStart="FIGURES 5051" captionStartId="77.[151,264,1944,1968]" captionTargetBox="[175,1422,203,1876]" captionTargetId="figure-15@77.[151,1435,181,1920]" captionTargetPageId="77" captionText="FIGURES 5051. Habitus and male genitalia. 50. Hydrochus pupillus Orchymont, paratype. 51. Hydrochus drechseli Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043340" httpUri="https://zenodo.org/record/5043340/files/figure.png" pageId="51" pageNumber="52">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>