162 lines
17 KiB
XML
162 lines
17 KiB
XML
<document id="D671AAC2C19D44D4623467647C721F28" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFCCFFCF6194FE2A05C8F4E0" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus pietersenae Makhan (HNHM 1993" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="49" updateTime="1698978858223" updateUser="plazi">
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<mods:title id="1A5BB868C659F5EAAA66D7BC1376674E">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="7ACBD859A41223E92C24462292D18EC6" type="personal">
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<mods:namePart id="536D2DFFD34FD19EB365E263A25B4BE7">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="AD4FB0DCD65EF85414674FFD38F45DD9">text</mods:typeOfResource>
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<mods:title id="ADA6E13D8FB8944D2F398E15A26EA89B">Zootaxa</mods:title>
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<mods:part id="9C6771F339BE3DF92089A515BEEDEDFE">
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<mods:date id="ED9C1F1A58BEFC0AA739BB2AE4DA86E2">2021</mods:date>
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<mods:number id="3A683A611F1855849BE38B70F1AD1F87">2021-06-28</mods:number>
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<mods:detail id="6DED12A9E775F13AF3558565F0F79CAE" type="volume">
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<mods:number id="1FC8EEB3FDF2DF32051513B5BBA612BA">4994</mods:number>
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<mods:end id="2BB0C7467FA95C966CEDE47D4013D092">93</mods:end>
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<mods:classification id="516B68CB0829ABBDDA38ECDCBF70C2B5">journal article</mods:classification>
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<mods:identifier id="057E06262418E9783AD8139331D56E74" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="B337044B947838CB8921119A47BB08C4" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03A187DEFFCCFFCF6194FE2A05C8F4E0" LSID="urn:lsid:plazi:treatment:03A187DEFFCCFFCF6194FE2A05C8F4E0" httpUri="http://treatment.plazi.org/id/03A187DEFFCCFFCF6194FE2A05C8F4E0" lastPageNumber="49" pageId="48" pageNumber="49">
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<subSubSection id="C3126543FFCCFFCF6194FE2A0641F6F2" box="[151,542,474,501]" pageId="48" pageNumber="49" type="nomenclature">
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<paragraph id="8BB736C8FFCCFFCF6194FE2A0641F6F2" blockId="48.[151,542,474,537]" box="[151,542,474,501]" pageId="48" pageNumber="49">
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<heading id="D0FF81A4FFCCFFCF6194FE2A0641F6F2" bold="true" box="[151,542,474,501]" fontSize="11" level="2" pageId="48" pageNumber="49" reason="3">
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<taxonomicName id="4C084D4BFFCCFFCF6194FE2A0641F6F2" ID-CoL="6MK92" authority="Makhan" authorityName="Makhan (HNHM" authorityYear="1993" box="[151,542,474,501]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pietersenae">
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<emphasis id="B97CEADAFFCCFFCF6194FE2A0641F6F2" bold="true" box="[151,542,474,501]" pageId="48" pageNumber="49">
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<emphasis id="B97CEADAFFCCFFCF6194FE2A05F2F6F2" bold="true" box="[151,429,474,501]" italics="true" pageId="48" pageNumber="49">Hydrochus pietersenae</emphasis>
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Makhan
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFCCFFCF6194FE0F0568F51E" box="[151,311,510,537]" pageId="48" pageNumber="49" type="description">
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<paragraph id="8BB736C8FFCCFFCF6194FE0F0568F51E" blockId="48.[151,542,474,537]" box="[151,311,510,537]" pageId="48" pageNumber="49">
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<figureCitation id="13332A4DFFCCFFCF6194FE0F04BAF51E" box="[151,229,510,537]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[161,1435,181,1891]" captionTargetId="figure-28@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043276" httpUri="https://zenodo.org/record/5043276/files/figure.png" pageId="48" pageNumber="49">Figs. 5</figureCitation>
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,
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<figureCitation id="13332A4DFFCCFFCF61F2FE0F0550F51E" box="[241,271,511,537]" captionStart="FIGURES 42–43" captionStartId="73.[151,264,1944,1968]" captionTargetBox="[177,1428,185,1892]" captionTargetId="figure-15@73.[151,1435,181,1920]" captionTargetPageId="73" captionText="FIGURES 42–43. Habitus and male genitalia. 42. Hydrochus astrictus Perkins. 43. Hydrochus pietersenae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043328" httpUri="https://zenodo.org/record/5043328/files/figure.png" pageId="48" pageNumber="49">43</figureCitation>
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,
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<figureCitation id="13332A4DFFCCFFCF6018FE0F0568F51E" box="[283,311,511,537]" captionStart="FIGURE 70" captionStartId="89.[151,250,1862,1886]" captionTargetBox="[156,1432,201,1824]" captionTargetId="figure-15@89.[151,1435,181,1837]" captionTargetPageId="89" captionText="FIGURE 70. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043382" httpUri="https://zenodo.org/record/5043382/files/figure.png" pageId="48" pageNumber="49">70</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3126543FFCCFFCF6194FDB305DEF5C0" pageId="48" pageNumber="49" type="reference_group">
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<paragraph id="8BB736C8FFCCFFCF6194FDB30666F55C" blockId="48.[151,569,579,603]" box="[151,569,579,603]" pageId="48" pageNumber="49">
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<treatmentCitationGroup id="AB1811E6FFCCFFCF6194FDB30666F55C" box="[151,569,579,603]" pageId="48" pageNumber="49">
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<treatmentCitation id="0AA910D9FFCCFFCF6194FDB30666F55C" author="Makhan, D." box="[151,569,579,603]" page="68" pageId="48" pageNumber="49" year="1993">
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<taxonomicName id="4C084D4BFFCCFFCF6194FDB30666F55C" ID-CoL="6MK92" authority="Makhan 1993: 68" authorityName="Makhan (HNHM" authorityYear="1993" box="[151,569,579,603]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pietersenae">
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<emphasis id="B97CEADAFFCCFFCF6194FDB305DEF55C" box="[151,385,579,603]" italics="true" pageId="48" pageNumber="49">Hydrochus pietersenae</emphasis>
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<bibRefCitation id="EF994B39FFCCFFCF6084FDB30666F55C" author="Makhan, D." box="[391,569,579,603]" pageId="48" pageNumber="49" pagination="65 - 70" refId="ref50274" refString="Makhan, D. (1993) Nine new Hydrochus species from South and Central America. Annales historico-naturales musei nationalis hungarici, 85, 65 - 70." type="journal article" year="1993">Makhan 1993: 68</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB736C8FFCCFFCF6194FD7905DEF5C0" blockId="48.[151,1436,649,999]" pageId="48" pageNumber="49">
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<materialsCitation id="3B603C95FFCCFFCF6194FD790522F5C0" country="Bolivia" location="Originally" municipality="Other" pageId="48" pageNumber="49" specimenCount="2" typeStatus="holotype">
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||
<location id="8ED76013FFCCFFCF6194FD790555F5A4" LSID="urn:lsid:plazi:treatment:03A187DEFFCCFFCF6194FE2A05C8F4E0:8ED76013FFCCFFCF6194FD790555F5A4" box="[151,266,649,675]" country="Bolivia" municipality="Other" name="Originally" pageId="48" pageNumber="49">Originally</location>
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described from
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<collectingCountry id="F31F7658FFCCFFCF60C4FD790642F5A4" box="[455,541,649,675]" name="Bolivia" pageId="48" pageNumber="49">Bolivia</collectingCountry>
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||
.
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<collectingMunicipality id="6BD3ACB2FFCCFFCF632AFD790636F5A4" box="[553,617,649,675]" pageId="48" pageNumber="49">Other</collectingMunicipality>
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than the
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<typeStatus id="54B3886AFFCCFFCF63D5FD790767F5A4" box="[726,824,649,675]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="54B3886AFFCCFFCF6270FD7A0781F5A3" box="[883,990,650,676]" pageId="48" pageNumber="49" type="paratype">paratypes</typeStatus>
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only a single male was examined, and imaged (
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<figureCitation id="13332A4DFFCCFFCF61FAFD5D0517F5CF" box="[249,328,685,712]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[161,1435,181,1891]" captionTargetId="figure-28@10.[151,1435,181,1919]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043276" httpUri="https://zenodo.org/record/5043276/files/figure.png" pageId="48" pageNumber="49">Figs. 5</figureCitation>
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,
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<figureCitation id="13332A4DFFCCFFCF6057FD5D052BF5C0" box="[340,372,685,711]" captionStart="FIGURES 42–43" captionStartId="73.[151,264,1944,1968]" captionTargetBox="[177,1428,185,1892]" captionTargetId="figure-15@73.[151,1435,181,1920]" captionTargetPageId="73" captionText="FIGURES 42–43. Habitus and male genitalia. 42. Hydrochus astrictus Perkins. 43. Hydrochus pietersenae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043328" httpUri="https://zenodo.org/record/5043328/files/figure.png" pageId="48" pageNumber="49">43</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3126543FFCCFFCF61C4FD2107B1F408" pageId="48" pageNumber="49" type="materials_examined">
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||
<paragraph id="8BB736C8FFCCFFCF61C4FD2107B1F408" blockId="48.[151,1436,649,999]" pageId="48" pageNumber="49">
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||
<emphasis id="B97CEADAFFCCFFCF61C4FD2105BDF5EC" bold="true" box="[199,482,721,747]" pageId="48" pageNumber="49">Material examined (1):</emphasis>
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||
<materialsCitation id="3B603C95FFCCFFCF60EFFD2107B5F408" collectingDate="2004-04-23" collectingDateMax="2004-04-30" collectingDateMin="2004-04-23" collectionCode="SEMC" collectorName="A. Cline & J. Wappes" country="Bolivia" elevation="400" latitude="-17.483334" location="Fauna" longLatPrecision="1278" longitude="-63.55" municipality="Hotel Flora" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Santa Cruz">
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||
<emphasis id="B97CEADAFFCCFFCF60EFFD21061FF5EC" bold="true" box="[492,576,721,747]" pageId="48" pageNumber="49">
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||
<collectingCountry id="F31F7658FFCCFFCF60EFFD21061FF5EC" box="[492,576,721,747]" name="Bolivia" pageId="48" pageNumber="49">Bolivia</collectingCountry>
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||
</emphasis>
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||
:
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<collectingRegion id="49CCF82AFFCCFFCF6351FD21068BF5EC" box="[594,724,721,747]" country="Bolivia" name="Santa Cruz" pageId="48" pageNumber="49">Santa Cruz</collectingRegion>
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,
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<locationDeviation id="9A052FEAFFCCFFCF63E0FD210057F5EC" box="[739,1032,721,748]" location="Buena Vista" pageId="48" pageNumber="49">
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||
<quantity id="4CF09B2DFFCCFFCF63E0FD21076AF5EC" box="[739,821,721,747]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="48" pageNumber="49" unit="km" value="3.7">3.7 km</quantity>
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SSE Buena Vista
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</locationDeviation>
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,
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<collectingMunicipality id="6BD3ACB2FFCCFFCF6515FD2100C5F5EC" box="[1046,1178,721,747]" pageId="48" pageNumber="49">Hotel Flora</collectingMunicipality>
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y
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<location id="8ED76013FFCCFFCF65BEFD22015BF5EB" LSID="urn:lsid:plazi:treatment:03A187DEFFCCFFCF6194FE2A05C8F4E0:8ED76013FFCCFFCF65BEFD22015BF5EB" box="[1213,1284,722,748]" country="Bolivia" latitude="-17.483334" longLatPrecision="1278" longitude="-63.55" municipality="Hotel Flora" name="Fauna" pageId="48" pageNumber="49" stateProvince="Santa Cruz">Fauna</location>
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, elev.
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<quantity id="4CF09B2DFFCCFFCF6451FD2101C3F5EB" box="[1362,1436,721,748]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="m" value="400.0">
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||
<elevation id="0025D1FBFFCCFFCF6451FD2101C3F5EB" box="[1362,1436,721,748]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="49" unit="m" value="400.0">400 m</elevation>
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</quantity>
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(
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<geoCoordinate id="EE3C500FFFCCFFCF61A3FD0504A0F408" box="[160,255,757,783]" degrees="17" direction="south" minutes="29" orientation="latitude" pageId="48" pageNumber="49" precision="925" value="-17.483334">17°29’S</geoCoordinate>
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,
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||
<geoCoordinate id="EE3C500FFFCCFFCF6008FD05052EF417" box="[267,369,757,784]" degrees="63" direction="west" minutes="33" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="-63.55">63°33’W</geoCoordinate>
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),
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<date id="FFB61008FFCCFFCF608AFD050674F408" box="[393,555,757,783]" pageId="48" pageNumber="49" value="2004-04-23" valueMax="2004-04-30" valueMin="2004-04-23">
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||
<collectingDate id="EFF2E9E0FFCCFFCF608AFD050674F408" box="[393,555,757,783]" pageId="48" pageNumber="49" value="2004-04-23" valueMax="2004-04-30" valueMin="2004-04-23">23–30.iv.2004</collectingDate>
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||
</date>
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||
, leg.
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<collectorName id="26FD531EFFCCFFCF6366FD06069AF408" box="[613,709,757,784]" pageId="48" pageNumber="49">A. Cline</collectorName>
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and
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<collectorName id="26FD531EFFCCFFCF63F8FD060734F417" box="[763,875,758,784]" pageId="48" pageNumber="49">J. Wappes</collectorName>
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||
(1
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||
<collectionCode id="ED19AE0DFFCCFFCF6293FD0507BEF408" box="[912,993,757,783]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="48" pageNumber="49" type="Museum">SEMC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFCCFFCF61C4FCE905C8F4E0" pageId="48" pageNumber="49" type="discussion">
|
||
<paragraph id="8BB736C8FFCCFFCF61C4FCE905C8F4E0" blockId="48.[151,1436,649,999]" pageId="48" pageNumber="49">
|
||
<emphasis id="B97CEADAFFCCFFCF61C4FCE90564F434" bold="true" box="[199,315,793,819]" pageId="48" pageNumber="49">Remarks.</emphasis>
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||
I have examined the male genitalia of the
|
||
<typeStatus id="54B3886AFFCCFFCF6229FCE907D3F434" box="[810,908,793,819]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D0EFD41FFCCFFCF62C5FCE90036F433" box="[966,1129,793,820]" pageId="48" pageNumber="49" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
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||
of
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||
<taxonomicName id="4C084D4BFFCCFFCF6591FCEA055CF450" authority="Makhan (HNHM)" authorityName="Makhan (HNHM" authorityYear="1993" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pietersenae">
|
||
<emphasis id="B97CEADAFFCCFFCF6591FCEA0168F434" box="[1170,1335,794,819]" italics="true" pageId="48" pageNumber="49">H. pietersenae</emphasis>
|
||
Makhan (HNHM)
|
||
</taxonomicName>
|
||
. All
|
||
<specimenCount id="9D0EFD41FFCCFFCF6038FCCD05BBF450" box="[315,484,829,855]" pageId="48" pageNumber="49" type="generic">five specimens</specimenCount>
|
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have the form of the distal piece as illustrated herein (
|
||
<figureCitation id="13332A4DFFCCFFCF655FFCCD00ECF450" box="[1116,1203,829,855]" captionStart="FIGURES 42–43" captionStartId="73.[151,264,1944,1968]" captionTargetBox="[177,1428,185,1892]" captionTargetId="figure-15@73.[151,1435,181,1920]" captionTargetPageId="73" captionText="FIGURES 42–43. Habitus and male genitalia. 42. Hydrochus astrictus Perkins. 43. Hydrochus pietersenae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043328" httpUri="https://zenodo.org/record/5043328/files/figure.png" pageId="48" pageNumber="49">Fig. 43</figureCitation>
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||
). The male genitalia drawing in
|
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<bibRefCitation id="EF994B39FFCCFFCF6014FC910598F47C" author="Makhan, D." box="[279,455,865,891]" pageId="48" pageNumber="49" pagination="389 - 393" refId="ref50546" refString="Makhan, D. (2001) A new genus, Soesilius, and a new species of Hydrochidae (Coleoptera) from America. Russian Entomological Journal, 10, 389 - 393." type="journal article" year="2001">Makhan (2001)</bibRefCitation>
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||
, which does not include two of the three processes of the distal piece, is inaccurate and misleading. Images of the male genitalia of the
|
||
<typeStatus id="54B3886AFFCCFFCF63A5FC750757F498" box="[678,776,901,927]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
are given herein. The
|
||
<typeStatus id="54B3886AFFCCFFCF6503FC75003DF498" box="[1024,1122,901,927]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
is in poor condition, broken due to the poor specimen technique of a prior worker. Therefore, a different specimen, from
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||
<collectingCountry id="F31F7658FFCCFFCF65A2FC5900A9F4C4" box="[1185,1270,937,963]" name="Bolivia" pageId="48" pageNumber="49">Bolivia</collectingCountry>
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, has been used for the habitus images.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |