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<document id="563F9CCD978F8FCB4DCEF93D7CA4FA6C" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A9297FFFB6FF03C1AB9C0FFE91" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scydmoraphes" docType="treatment" docVersion="9" lastPageNumber="71" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="71" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="87ABDB4959351AB554298AB8293E1F59">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="2D866385225B183D9677AE379F8FE755">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="2473D47E1C21FD8A31017A33F3DD0829">Perkovsky, Evgeny</mods:namePart>
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<treatment id="03A187A9297FFFB6FF03C1AB9C0FFE91" ID-DOI="http://doi.org/10.5281/zenodo.5681857" ID-GBIF-Taxon="124407830" ID-Zenodo-Dep="5681857" LSID="urn:lsid:plazi:treatment:03A187A9297FFFB6FF03C1AB9C0FFE91" httpUri="http://treatment.plazi.org/id/03A187A9297FFFB6FF03C1AB9C0FFE91" lastPageId="71" lastPageNumber="71" pageId="70" pageNumber="71">
<subSubSection id="C3126534297FFFB7FF03C1AB9C39FB13" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="8BB736BF297FFFB7FF03C1AB9CC2FB71" blockId="70.[151,406,1158,1218]" box="[151,406,1158,1184]" pageId="70" pageNumber="71">
<heading id="D0FF81D3297FFFB7FF03C1AB9CC2FB71" bold="true" box="[151,406,1158,1184]" fontSize="11" level="1" pageId="70" pageNumber="71" reason="1">
<emphasis id="B97CEAAD297FFFB7FF03C1AB9CC2FB71" bold="true" box="[151,406,1158,1184]" pageId="70" pageNumber="71">
?
<taxonomicName id="4C084D3C297FFFB7FF31C1AB9C01FB71" ID-CoL="8X7TZ" box="[165,341,1158,1184]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD297FFFB7FF31C1AB9C01FB71" bold="true" box="[165,341,1158,1184]" italics="true" pageId="70" pageNumber="71">Scydmoraphes</emphasis>
</taxonomicName>
sp. 1
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF297FFFB7FF03C1849C39FB13" blockId="70.[151,406,1158,1218]" box="[151,365,1193,1218]" pageId="70" pageNumber="71">
(
<figureCitation id="13332A3A297FFFB7FF0BC1849DA1FB13" box="[159,245,1193,1218]" captionStart="FIGURES 21 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="70" pageNumber="71">Figs 32</figureCitation>
,
<figureCitation id="13332A3A297FFFB7FE95C1849C31FB13" box="[257,357,1193,1218]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">215216</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3126534297FFFB7FF03C1DC9E9EFAFF" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="8BB736BF297FFFB7FF03C1DC9E9EFAFF" blockId="70.[151,1437,1265,2010]" pageId="70" pageNumber="71">
<materialsCitation id="3B603CE2297FFFB7FF03C1DC9E9EFAFF" ID-GBIF-Occurrence="1300093514" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="70" pageNumber="71" specimenCount="1" stateProvince="Rovno">
<emphasis id="B97CEAAD297FFFB7FF03C1DC9E04FADB" bold="true" box="[151,848,1265,1290]" pageId="70" pageNumber="71">
<collectingCounty id="62D64E33297FFFB7FF03C1DC9DABFADB" box="[151,255,1265,1290]" pageId="70" pageNumber="71">Material</collectingCounty>
studied.
<collectingMunicipality id="6BD3ACC5297FFFB7FEE7C1DC9F5DFADB" box="[371,521,1265,1290]" pageId="70" pageNumber="71">Late Eocene</collectingMunicipality>
of
<location id="8ED76064297FFFB7FDA2C1DC9FC4FADB" LSID="urn:lsid:plazi:treatment:03A187A9297FFFB6FF03C1AB9C0FFE91:8ED76064297FFFB7FDA2C1DC9FC4FADB" box="[566,656,1265,1290]" county="Material" municipality="Late Eocene" name="Europe" pageId="70" pageNumber="71" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D297FFFB7FD34C1DC9FBAFADB" box="[672,750,1265,1290]" country="Ukraine" name="Rivne" pageId="70" pageNumber="71">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, subtriangular piece of amber
<quantity id="4CF09B5A297FFFB7FED0C03B9CC7FAFC" box="[324,403,1302,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="70" pageNumber="71" unit="mm" value="12.0">12 mm</quantity>
long (
<figureCitation id="13332A3A297FFFB7FE4FC0389F7AFAFF" box="[475,558,1301,1326]" captionStart="FIGURES 21 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="70" pageNumber="71">Fig. 32</figureCitation>
), collection number K-6764 (
<collectionCode id="ED19AE7A297FFFB7FCEDC03B9EE9FAFF" box="[889,957,1302,1326]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="70" pageNumber="71">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126534297FFFB7FF53C0149E37F8BF" pageId="70" pageNumber="71" type="description">
<paragraph id="8BB736BF297FFFB7FF53C0149807FA80" blockId="70.[151,1437,1265,2010]" box="[199,1363,1337,1362]" pageId="70" pageNumber="71">
<emphasis id="B97CEAAD297FFFB7FF53C0149C0DFA83" bold="true" box="[199,345,1337,1362]" pageId="70" pageNumber="71">Description.</emphasis>
Body (
<figureCitation id="13332A3A297FFFB7FE3EC0149F13FA83" box="[426,583,1337,1362]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Figs 215216</figureCitation>
) elongate, moderately slender and convex, dark brown; BL
<quantity id="4CF09B5A297FFFB7FB73C017981AFA80" box="[1255,1358,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="70" pageNumber="71" unit="mm" value="0.98">0.98 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF297FFFB7FF53C07099DDFA33" blockId="70.[151,1437,1265,2010]" pageId="70" pageNumber="71">
Head (
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) broadest at eyes, HL
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; vertex and frons not visible; eyes large, oval, strongly convex; tempora seem shorter than eyes. Punctures and setae on frons and vertex poorly visible. Antennae (
<figureCitation id="13332A3A297FFFB7FAFEC0AC9DAFFA6F" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Figs 215216</figureCitation>
) much shorter than half BL, AnL about
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, gradually thickened distally and relatively slender, scape and pedicel strongly elongate, shapes of remaining antennomeres difficult to define.
</paragraph>
<paragraph id="8BB736BF297FFFB7FF53C0C39947F90F" blockId="70.[151,1437,1265,2010]" pageId="70" pageNumber="71">
Pronotum (
<figureCitation id="13332A3A297FFFB7FED1C0C09CB6F9D7" box="[325,482,1517,1542]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Figs 215216</figureCitation>
) elongate, broadest at base but with nearly parallel-sided lateral margins in posterior half; PL
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, PW
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; anterior margin strongly and evenly rounded and laterally confluent with lateral margins, so that anterior pronotal corners are not marked; sides rounded in anterior third and nearly straight in posterior half; posterior corners distinct, nearly right-angled, blunt; posterior margin nearly straight; base with one pair of shallow and diffused lateral pits (
<figureCitation id="13332A3A297FFFB7FD16C3509FBDF947" box="[642,745,1661,1686]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297FFFB7FD6EC3509E70F947" box="[762,804,1661,1686]" italics="true" pageId="70" pageNumber="71">abp</emphasis>
) connected by complete and uninterrupted transverse antebasal groove (
<figureCitation id="13332A3A297FFFB7FEF0C38C9C92F96B" box="[356,454,1697,1722]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297FFFB7FE46C38C9CA8F96B" box="[466,508,1697,1722]" italics="true" pageId="70" pageNumber="71">abg</emphasis>
). Punctures on pronotal disc (
<figureCitation id="13332A3A297FFFB7FCDAC38C9EFAF96B" box="[846,942,1697,1722]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
) small but distinct, separated by spaces 23 × as wide as diameters of punctures; setae (
<figureCitation id="13332A3A297FFFB7FDE9C3E89F89F90F" box="[637,733,1733,1758]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
) sparse, short and suberect.
</paragraph>
<paragraph id="8BB736BF297FFFB7FF53C3C49E86F89B" blockId="70.[151,1437,1265,2010]" pageId="70" pageNumber="71">
Elytra (
<figureCitation id="13332A3A297FFFB7FE8EC3C49CE2F8D3" box="[282,438,1769,1794]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Figs 215216</figureCitation>
) elongate oval, moderately convex, broadest near middle, EL
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, EW
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, EI 1.57; elytral base distinctly broader than base of pronotum; each elytron with one distinct asetose basal fovea (
<figureCitation id="13332A3A297FFFB7FAFBC2209D91F89B" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297FFFB7FF5BC21C9DA5F89B" box="[207,241,1841,1866]" italics="true" pageId="70" pageNumber="71">bef</emphasis>
); punctures and setae (
<figureCitation id="13332A3A297FFFB7FE60C21C9F00F89B" box="[500,596,1841,1866]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Fig. 216</figureCitation>
) similar to those on pronotal disc.
</paragraph>
<paragraph id="8BB736BF297FFFB7FF53C27B9E37F8BF" blockId="70.[151,1437,1265,2010]" box="[199,867,1877,1902]" pageId="70" pageNumber="71">
Legs (
<figureCitation id="13332A3A297FFFB7FE98C2789CFDF8BF" box="[268,425,1877,1902]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="70" pageNumber="71">Figs 215216</figureCitation>
) moderately long, slender, unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C3126534297FFFB6FF53C2549C0FFE91" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8BB736BF297FFFB6FF53C2549C0FFE91" blockId="70.[151,1437,1265,2010]" lastBlockId="71.[151,1436,151,320]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<emphasis id="B97CEAAD297FFFB7FF53C2549C6FF843" bold="true" box="[199,315,1913,1938]" pageId="70" pageNumber="71">Remarks.</emphasis>
Diagnostic characters of the extant genus
<taxonomicName id="4C084D3C297FFFB7FCC6C25499CAF843" authority="Reitter, 1891" authorityName="Reitter" authorityYear="1891" box="[850,1182,1913,1938]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD297FFFB7FCC6C2549EA1F843" box="[850,1013,1913,1938]" italics="true" pageId="70" pageNumber="71">Scydmoraphes</emphasis>
<bibRefCitation id="EF994B4E297FFFB7FB91C25499CAF843" author="Reitter" box="[1029,1182,1913,1938]" pageId="70" pageNumber="71" refString="Reitter, E. (1891) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. Friedlander &amp; Son, Berlin, 420 pp. [von Heyden, L., Reitter, E. &amp; Weise, J. (Eds.)]" type="book" year="1891">Reitter, 1891</bibRefCitation>
</taxonomicName>
were illustrated and summarized in
<bibRefCitation id="EF994B4E297FFFB7FED0C2B09F70F867" author="Jaloszynski" box="[324,548,1949,1974]" pageId="70" pageNumber="71" refString="Jaloszynski, P. (2015 f) Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4007 (3), 343 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 4007.3.3" type="journal article" year="2015" yearSuffix="f">Jałoszyński (2015f)</bibRefCitation>
. Characters of the
<collectingRegion id="49CCF85D297FFFB7FD61C2B39E14F864" box="[757,832,1950,1973]" country="Ukraine" name="Rivne" pageId="70" pageNumber="71">Rovno</collectingRegion>
amber specimen K-6764 that suggest it can belong in
<taxonomicName id="4C084D3C297FFFB7FF03C2EC9C6EF80B" box="[151,314,1985,2010]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAAD297FFFB7FF03C2EC9C6EF80B" box="[151,314,1985,2010]" italics="true" pageId="70" pageNumber="71">Scydmoraphes</emphasis>
</taxonomicName>
are as follows: the pronotum elongate, with rounded anterior margin, anterior pronotal corners not marked, posterior pronotal corners distinct; lateral pronotal margins developed as sharp edges or carinae (
<figureCitation id="13332A3A297EFFB6FAA2C5BA98C3FF61" box="[1334,1431,151,176]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="71" pageNumber="72">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297EFFB6FF03C5919DEDFF04" box="[151,185,188,213]" italics="true" pageId="71" pageNumber="72">lpc</emphasis>
), the pronotal base with one pair of lateral pits (
<figureCitation id="13332A3A297EFFB6FD4BC5919E15FF04" box="[735,833,188,213]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="71" pageNumber="72">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297EFFB6FCD9C5919E23FF04" box="[845,887,188,213]" italics="true" pageId="71" pageNumber="72">abp</emphasis>
) developed as shallow and diffused impressions connected by complete transverse antebasal groove (
<figureCitation id="13332A3A297EFFB6FD73C5F29E1EFF29" box="[743,842,223,248]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="71" pageNumber="72">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297EFFB6FCC1C5F29E2BFF29" box="[853,895,223,248]" italics="true" pageId="71" pageNumber="72">abg</emphasis>
); and each elytron with one distinct and asetose basal fovea (
<figureCitation id="13332A3A297EFFB6FEBDC4299CDAFECC" box="[297,398,260,285]" captionStart="FIGURES 215 216" captionStartId="71.[151,264,1205,1228]" captionTargetBox="[151,1427,375,1184]" captionTargetId="figure@71.[151,1436,375,1184]" captionTargetPageId="71" captionText="FIGURES 215 216.? Scydmoraphes sp. 1 (K- 6764) in dorsal view. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; lpc, lateral pronotal carina." httpUri="https://zenodo.org/record/262461/files/figure.png" pageId="71" pageNumber="72">Fig. 216</figureCitation>
;
<emphasis id="B97CEAAD297EFFB6FE0FC4299CE9FECC" box="[411,445,260,285]" italics="true" pageId="71" pageNumber="72">bef</emphasis>
). However, the most important ventral characters are not visible and the placement in genus is putative.
</paragraph>
</subSubSection>
</treatment>
</document>