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<document id="937243679EA6E796015E28F49D1A605F" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92913FFDBFF03C0169CAAF80B" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rovnoscydmus" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="43" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="7EA8BECBAB7FB9AD339B8FCCEA4B0549">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="24D127F13A7FB08E6D30B417F70C3DD9">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="4AF2A9D98D6B397BDF574BF1801D6144">Perkovsky, Evgeny</mods:namePart>
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<mods:date id="BBEA4F4A32D49E776537F05373D7D996">2016</mods:date>
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<treatment id="03A187A92913FFDBFF03C0169CAAF80B" ID-DOI="http://doi.org/10.5281/zenodo.5681770" ID-GBIF-Taxon="124407850" ID-Zenodo-Dep="5681770" LSID="urn:lsid:plazi:treatment:03A187A92913FFDBFF03C0169CAAF80B" httpUri="http://treatment.plazi.org/id/03A187A92913FFDBFF03C0169CAAF80B" lastPageNumber="43" pageId="42" pageNumber="43">
<subSubSection id="C31265342913FFDBFF03C0169C39FAA7" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BB736BF2913FFDBFF03C0169F41FA84" blockId="42.[151,533,1339,1398]" box="[151,533,1339,1365]" pageId="42" pageNumber="43">
<heading id="D0FF81D32913FFDBFF03C0169F41FA84" bold="true" box="[151,533,1339,1365]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B97CEAAD2913FFDBFF03C0169F41FA84" bold="true" box="[151,533,1339,1365]" pageId="42" pageNumber="43">
<taxonomicName id="4C084D3C2913FFDBFF31C0169C0EFA84" ID-CoL="8XPPK" box="[165,346,1339,1365]" class="Insecta" family="Staphylinidae" genus="Rovnoscydmus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2913FFDBFF31C0169C0EFA84" bold="true" box="[165,346,1339,1365]" italics="true" pageId="42" pageNumber="43">Rovnoscydmus</emphasis>
</taxonomicName>
sp. specimen 2
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF2913FFDBFF03C0709C39FAA7" blockId="42.[151,533,1339,1398]" box="[151,365,1373,1398]" pageId="42" pageNumber="43">
(
<figureCitation id="13332A3A2913FFDBFF0BC0709DA1FAA7" box="[159,245,1373,1398]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="42" pageNumber="43">Figs 15</figureCitation>
,
<figureCitation id="13332A3A2913FFDBFE95C0709C31FAA7" box="[257,357,1373,1398]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">126129</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31265342913FFDBFF03C0889E2AFA33" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BB736BF2913FFDBFF03C0889E2AFA33" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
<materialsCitation id="3B603CE22913FFDBFF03C0889E2AFA33" ID-GBIF-Occurrence="1300093487" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Rovno">
<emphasis id="B97CEAAD2913FFDBFF03C0889E63FA6F" bold="true" box="[151,823,1445,1470]" pageId="42" pageNumber="43">
<collectingCounty id="62D64E332913FFDBFF03C0889DABFA6F" box="[151,255,1445,1470]" pageId="42" pageNumber="43">Material</collectingCounty>
studied.
<collectingMunicipality id="6BD3ACC52913FFDBFEF8C0889CAAFA6F" box="[364,510,1445,1470]" pageId="42" pageNumber="43">Late Eocene</collectingMunicipality>
of
<location id="8ED760642913FFDBFDB1C0889F2BFA6F" LSID="urn:lsid:plazi:treatment:03A187A92913FFDBFF03C0169CAAF80B:8ED760642913FFDBFDB1C0889F2BFA6F" box="[549,639,1445,1470]" county="Material" municipality="Late Eocene" name="Europe" pageId="42" pageNumber="43" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D2913FFDBFD1FC0889F8DFA6F" box="[651,729,1445,1470]" country="Ukraine" name="Rivne" pageId="42" pageNumber="43">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, irregular piece of amber
<quantity id="4CF09B5A2913FFDBFF70C0E49C6EFA30" box="[228,314,1481,1506]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="42" pageNumber="43" unit="mm" value="8.5">8.5 mm</quantity>
long (
<figureCitation id="13332A3A2913FFDBFE15C0E49C80FA33" box="[385,468,1481,1506]" captionStart="FIGURES 1 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="42" pageNumber="43">Fig. 15</figureCitation>
), collection number K-26348 (
<collectionCode id="ED19AE7A2913FFDBFCBAC0E79E26FA33" box="[814,882,1482,1506]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="42" pageNumber="43">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31265342913FFDBFF53C0C09E37F8BF" pageId="42" pageNumber="43" type="description">
<paragraph id="8BB736BF2913FFDBFF53C0C09D9EF9F8" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
<emphasis id="B97CEAAD2913FFDBFF53C0C09C0DF9D7" bold="true" box="[199,345,1517,1542]" pageId="42" pageNumber="43">Description.</emphasis>
Body (
<figureCitation id="13332A3A2913FFDBFE25C0C09F04F9D7" box="[433,592,1517,1542]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Figs 126129</figureCitation>
) elongate, slender and relatively strongly convex, brown; BL about
<quantity id="4CF09B5A2913FFDBFAFFC0C39D92F9F8" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="42" pageNumber="43" unit="mm" value="0.83">0.83 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF2913FFDBFF53C3189F99F947" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
Head (
<figureCitation id="13332A3A2913FFDBFE87C3189CE6F99F" box="[275,434,1589,1614]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Figs 128129</figureCitation>
) only partly visible, broadest at eyes, HW
<quantity id="4CF09B5A2913FFDBFC31C31B995BF99C" box="[933,1039,1590,1614]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="mm" value="0.2">0.20 mm</quantity>
; vertex transverse; anterior part of head poorly visible; compound eyes large, strongly convex, oval; tempora seem shorter than eyes. Punctures and setae on head not visible. Antennae poorly visible.
</paragraph>
<paragraph id="8BB736BF2913FFDBFF53C38F9984F8D3" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
Pronotum (
<figureCitation id="13332A3A2913FFDBFED3C38C9CFDF96B" box="[327,425,1697,1722]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Fig. 128</figureCitation>
) slightly elongate, broadest near middle; PL
<quantity id="4CF09B5A2913FFDBFC26C38C994FF968" box="[946,1051,1697,1722]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="42" pageNumber="43" unit="mm" value="0.25">0.25 mm</quantity>
, PW
<quantity id="4CF09B5A2913FFDBFBC8C38F9992F968" box="[1116,1222,1698,1722]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="42" pageNumber="43" unit="mm" value="0.23">0.23 mm</quantity>
; punctures on disc (
<figureCitation id="13332A3A2913FFDBFF0BC3E89C54F90F" box="[159,256,1733,1758]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Fig. 128</figureCitation>
) small but distinct, separated by spaces 24 × as wide as diameters of punctures; setae (
<figureCitation id="13332A3A2913FFDBFB66C3E89807F90F" box="[1266,1363,1733,1758]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Fig. 128</figureCitation>
) short and sparse, nearly recumbent, on sides of pronotum replaced by distinct thick bristles (
<figureCitation id="13332A3A2913FFDBFBF5C3C49990F8D3" box="[1121,1220,1769,1794]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Fig. 129</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF2913FFDBFF53C220995AF89B" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
Elytra (
<figureCitation id="13332A3A2913FFDBFE8FC2209CEEF8F7" box="[283,442,1805,1830]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Figs 126, 128</figureCitation>
) elongate oval, strongly convex, broadest slightly in front of middle, EL
<quantity id="4CF09B5A2913FFDBFB68C2209830F8F4" box="[1276,1380,1805,1830]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="42" pageNumber="43" unit="mm" value="0.58">0.58 mm</quantity>
, EW
<quantity id="4CF09B5A2913FFDBFF03C21C9DA9F898" box="[151,253,1841,1866]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="42" pageNumber="43" unit="mm" value="0.35">0.35 mm</quantity>
, EI 1.64; punctures and setae similar to those on pronotum (
<figureCitation id="13332A3A2913FFDBFC0BC21C9955F89B" box="[927,1025,1841,1866]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Fig. 128</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF2913FFDBFF53C27B9E37F8BF" blockId="42.[151,1437,1445,2010]" box="[199,867,1877,1902]" pageId="42" pageNumber="43">
Legs (
<figureCitation id="13332A3A2913FFDBFE98C2789CFCF8BF" box="[268,424,1877,1902]" captionStart="FIGURES 126 129" captionStartId="44.[151,264,859,882]" captionTargetBox="[158,1415,203,825]" captionTargetId="figure@44.[158,1423,203,826]" captionTargetPageId="44" captionText="FIGURES 126 129. Rovnoscydmus sp. specimen 2 (K- 26348). Dorsal (126, 128) and ventral (127, 129) view. Abbreviations: abg, antebasal groove; bef, basal elytral fovea; bst, basisternal part of prosternum; hy, hypomeron; lbr, labrum; md, mandible; mtvp, metaventral intercoxal process; mxp 2 4, maxillary palpomere II IV; nr, ' neck' region; st 3 8, abdominal sternite III VIII; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262439/files/figure.png" pageId="42" pageNumber="43">Figs 126129</figureCitation>
) moderately long, slender, unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C31265342913FFDBFF53C2549CAAF80B" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BB736BF2913FFDBFF53C2549CAAF80B" blockId="42.[151,1437,1445,2010]" pageId="42" pageNumber="43">
<emphasis id="B97CEAAD2913FFDBFF53C2549C6FF843" bold="true" box="[199,315,1913,1938]" pageId="42" pageNumber="43">Remarks.</emphasis>
This inclusion is similar to specimen 1, but as it is only partly visible, a detailed comparison is not possible. Also in this specimen the ventral side of prothorax and pterothorax is well exposed, although details of the mesoventrite are not visible.
</paragraph>
</subSubSection>
</treatment>
</document>