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<mods:title id="8C2D2E1B92BD15DC094B675D97938E36">Illustrated redescription of Haliplus (Liaphlus) arrowi Guignot, 1936 (Coleoptera: Haliplidae) from the Western Ghats, India, and notes on the closely related H. angustifrons Régimbart, 1892</mods:title>
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<mods:namePart id="5F0C48B46D80651547DEBCF30E630CB5">Sheth, Sayali D.</mods:namePart>
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<treatment id="03A187A0FFE3FF855F865D8DFA738F5D" ID-DOI="http://doi.org/10.5281/zenodo.5681897" ID-GBIF-Taxon="121061631" ID-Zenodo-Dep="5681897" LSID="urn:lsid:plazi:treatment:03A187A0FFE3FF855F865D8DFA738F5D" httpUri="http://treatment.plazi.org/id/03A187A0FFE3FF855F865D8DFA738F5D" lastPageId="7" lastPageNumber="362" pageId="6" pageNumber="361" scope_family="Haliplidae" scope_order="Coleoptera">
<subSubSection id="C312653DFFE3FF845F865D8DFCC58DB1" pageId="6" pageNumber="361" type="nomenclature">
<paragraph id="8BB736B6FFE3FF845F865D8DFD118CCF" blockId="6.[151,766,1708,1734]" box="[151,766,1708,1734]" pageId="6" pageNumber="361">
<heading id="D0FF81DAFFE3FF845F865D8DFD118CCF" bold="true" box="[151,766,1708,1734]" fontSize="11" level="1" pageId="6" pageNumber="361" reason="1">
<taxonomicName id="4C084D35FFE3FF845F865D8DFD118CCF" ID-CoL="C3S7L" authority="Regimbart, 1892" authorityName="Regimbart" authorityYear="1892" box="[151,766,1708,1734]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="angustifrons" subGenus="Liaphlus">
<emphasis id="B97CEAA4FFE3FF845F865D8DFD118CCF" bold="true" box="[151,766,1708,1734]" pageId="6" pageNumber="361">
<emphasis id="B97CEAA4FFE3FF845F865D8DFDC98CCF" bold="true" box="[151,550,1708,1734]" italics="true" pageId="6" pageNumber="361">Haliplus (Liaphlus) angustifrons</emphasis>
<bibRefCitation id="EF994B47FFE3FF845D3C5D8DFD118CCF" author="Regimbart" box="[557,766,1708,1734]" pageId="6" pageNumber="361" refString="Regimbart, M. (1892) Insectes du Bengale Occidental. Annales de la Societe Entomologiques de Belgique, 36, 112 - 121." type="journal article" year="1892">Régimbart, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB736B6FFE3FF845F865DD2FCC58DB1" blockId="6.[151,1436,1778,1976]" pageId="6" pageNumber="361">
Known distribution from the Oriental region:
<collectingCountry id="F31F7626FFE3FF845DBE5DD3FD068D02" box="[687,745,1778,1803]" name="India" pageId="6" pageNumber="361">India</collectingCountry>
(Punjab; Himachal Pradesh- Simla hills; Jharkhand-Ranchi district (previously in Bihar now in Jharkhand), Hazaribagh district (previously in Bihar now in Jharkhand), Santhal paraganas district (previously in Bihar now in Jharkhand); Kolkata-West Bengal; Odisha (previously Orissa)- Cuttak district, Balasore district, Mayurbanj district; Madhya Pradesh- Jabalpur; Rajasthan- Pilani; Arunachal Pradesh; Maharashtra; Karnataka),
<collectingCountry id="F31F7626FFE3FF845DC25C5FFCAD8D9F" box="[723,834,1918,1942]" name="Myanmar" pageId="6" pageNumber="361">Myanmar</collectingCountry>
,
<collectingCountry id="F31F7626FFE3FF845C485C5CFC708D9F" box="[857,927,1917,1942]" name="Nepal" pageId="6" pageNumber="361">Nepal</collectingCountry>
,
<collectingCountry id="F31F7626FFE3FF845CA65C5CFBF68D9F" box="[951,1049,1917,1942]" name="Pakistan" pageId="6" pageNumber="361">Pakistan</collectingCountry>
,
<collectingCountry id="F31F7626FFE3FF845B205C5FFB858D9C" box="[1073,1130,1918,1941]" name="Laos" pageId="6" pageNumber="361">Laos</collectingCountry>
,
<collectingCountry id="F31F7626FFE3FF845B935C5CFB0B8D9F" box="[1154,1252,1917,1942]" name="Vietnam" pageId="6" pageNumber="361">Vietnam</collectingCountry>
,
<collectingCountry id="F31F7626FFE3FF845BED5C5CFA8F8D9F" box="[1276,1376,1917,1942]" name="Thailand" pageId="6" pageNumber="361">Thailand</collectingCountry>
and
<collectingCountry id="F31F7626FFE3FF845F865CBEFEF58DB1" box="[151,282,1951,1976]" name="Bangladesh" pageId="6" pageNumber="361">Bangladesh</collectingCountry>
(
<bibRefCitation id="EF994B47FFE3FF845E395CBEFE268DB1" author="Vazirani" box="[296,457,1951,1976]" pageId="6" pageNumber="361" refString="Vazirani, T. G. (1984) The fauna of India: Coleoptera. Family Gyrinidae and family Haliplidae, Zoological Survey of India. 140 pp" type="book" year="1984">Vazirani 1984</bibRefCitation>
;
<bibRefCitation id="EF994B47FFE3FF845EC45CBEFD898DB1" author="Vondel" box="[469,614,1951,1976]" pageId="6" pageNumber="361" refString="Vondel, B. J. Van (1993) Revision of the Liaphlus species of the Oriental region excluding China (Coleoptera: Haliplidae). Tijdschrift voor Entomologie, 136, 289 - 316." type="journal article" year="1993">Vondel 1993</bibRefCitation>
,
<bibRefCitation id="EF994B47FFE3FF845D605CBEFD448DB1" author="Vondel" box="[625,683,1951,1976]" pageId="6" pageNumber="361" refString="Vondel, B. J. van &amp; Volpe, V. (2015) Additional notes on Haliplus volpei Vondel (Coleoptera, Haliplidae). Latissimus, 36, 5 - 7." type="journal article" year="2015">2015</bibRefCitation>
) (
<figureCitation id="13332A33FFE3FF845DD05CBEFCF18DB1" box="[705,798,1951,1976]" captionStart="FIGURE 1" captionStartId="1.[151,250,1651,1673]" captionTargetBox="[193,1390,740,1630]" captionTargetId="figure@1.[193,1390,740,1630]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map for H. arrowi and H. angustifrons with previous and latest localities from the Oriental region. Red dots and blue squares—recent and previous records of H. arrowi from the Oriental region, respectively. Yellow trianglesknown records of H. angustifrons from the Oriental region." httpUri="https://zenodo.org/record/262466/files/figure.png" pageId="6" pageNumber="361">Figure1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C312653DFFE2FF855FD65BB6FC9A8BCF" pageId="7" pageNumber="362" type="materials_examined">
<paragraph id="8BB736B6FFE2FF855FD65BB6FC5F8ADA" blockId="7.[151,1436,151,454]" pageId="7" pageNumber="362">
<emphasis id="B97CEAA4FFE2FF855FD65BB6FE798AB9" bold="true" box="[199,406,151,176]" pageId="7" pageNumber="362">Material studied.</emphasis>
SDS &amp; HVG studied single specimen (male) from Bhugaon Lake, Pune;
<collectionCode id="ED19AE73FFE2FF855BF05BB9FAC88AA6" box="[1249,1319,152,175]" pageId="7" pageNumber="362">BJVV</collectionCode>
examined the
<typeStatus id="54B38814FFE2FF855FD15B9AFF1F8ADA" box="[192,240,187,211]" pageId="7" pageNumber="362">type</typeStatus>
specimen and additional specimens from
<collectingCountry id="F31F7626FFE2FF855DD95B9BFCED8ADA" box="[712,770,186,211]" name="India" pageId="7" pageNumber="362">India</collectingCountry>
(
<bibRefCitation id="EF994B47FFE2FF855C1E5B9BFC4C8ADA" author="Vondel" box="[783,931,186,211]" pageId="7" pageNumber="362" refString="Vondel, B. J. Van (1993) Revision of the Liaphlus species of the Oriental region excluding China (Coleoptera: Haliplidae). Tijdschrift voor Entomologie, 136, 289 - 316." type="journal article" year="1993">Vondel 1993</bibRefCitation>
).
</paragraph>
<paragraph id="8BB736B6FFE2FF855FD65BFCFC9A8BCF" blockId="7.[151,1436,151,454]" pageId="7" pageNumber="362">
<emphasis id="B97CEAA4FFE2FF855FD65BFCFD508AFF" bold="true" box="[199,703,221,246]" pageId="7" pageNumber="362">
Distinguishing characters from
<taxonomicName id="4C084D35FFE2FF855D555BFFFD5A8AFC" box="[580,693,221,245]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855D555BFFFD5A8AFC" bold="true" box="[580,693,221,245]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
:
</emphasis>
Head overall brown, dorsally eyes closer than that in
<taxonomicName id="4C084D35FFE2FF855A355BFFFA7D8AFC" box="[1316,1426,222,245]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855A355BFFFA7D8AFC" box="[1316,1426,222,245]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
- distance between eyes 0.7 times width of one eye; pronotum darker on disc than on sides (
<figureCitation id="13332A33FFE2FF855BD05BDEFACA8B11" box="[1217,1317,255,280]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
A); elytral maculation extensive against moderately dense to rare maculation in
<taxonomicName id="4C084D35FFE2FF855CDC5A02FBAF8B33" box="[973,1088,291,314]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855CDC5A02FBAF8B33" box="[973,1088,291,314]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
(
<figureCitation id="13332A33FFE2FF855B455A03FB538B32" box="[1108,1212,290,315]" captionStart="FIGURE 4" captionStartId="5.[151,250,1948,1970]" captionText="FIGURE 4. Variation in the elytral maculation (scale 1 mm): H. arrowi (A, C I); H. angustifrons (B)." pageId="7" pageNumber="362">Figure 4</figureCitation>
B), twenty rows of primary and secondary punctures present per elytron, 30 punctures in first primary-puncture row; ventrally brown, prosternum posteriorly with deeper pit than that in
<taxonomicName id="4C084D35FFE2FF855DCE5A48FCA18B76" box="[735,846,360,384]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855DCE5A48FCA18B76" box="[735,846,360,384]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
(
<figureCitation id="13332A33FFE2FF855C4F5A46FC2F8B89" box="[862,960,359,384]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
B), metaventrite with two deeper and less connected pits anterolaterally (
<figureCitation id="13332A33FFE2FF855D135AABFD868BAA" box="[514,617,394,419]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
D); protarsi narrower and slender (
<figureCitation id="13332A33FFE2FF855B1C5AABFB9C8BAA" box="[1037,1139,394,419]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
E); median lobe of male genitalia narrower and less spatulate in apical region (
<figureCitation id="13332A33FFE2FF855DE35A8CFCBD8BCF" box="[754,850,429,454]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
C).
</paragraph>
</subSubSection>
<caption id="DF77663EFFE2FF855F865937FE46889E" ID-Table-UUID="DF77663EFFE2FF855F865937FE46889E" httpUri="http://table.plazi.org/id/DF77663EFFE2FF855F865937FE46889E" pageId="7" pageNumber="362" targetBox="[151,1235,690,1122]" targetIsTable="true" targetPageId="7">
<paragraph id="8BB736B6FFE2FF855F865937FE46889E" blockId="7.[151,1436,533,663]" pageId="7" pageNumber="362">
<emphasis id="B97CEAA4FFE2FF855F865937FEE18824" bold="true" box="[151,270,534,557]" pageId="7" pageNumber="362">TABLE 1.</emphasis>
Measurements of
<taxonomicName id="4C084D35FFE2FF855EF65934FDAD8824" box="[487,578,533,557]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAA4FFE2FF855EF65934FDAD8824" box="[487,578,533,557]" italics="true" pageId="7" pageNumber="362">Haliplus</emphasis>
</taxonomicName>
specimens: TL—total length measured from head to the elytral tip; MW maximum width of specimen measured across elytra; PL—medial length of pronotum; PW—maximum width of pronotum measured at its base; DE—minimum distance between eyes, and EW—maximum width of one eye. All measurements are in mm.
</paragraph>
</caption>
<paragraph id="8BB736B6FFE2FF855F8E5993FE0B8E6B" pageId="7" pageNumber="362">
<table id="F908C416FFE2007D5F865993FB3C8E6B" box="[151,1235,690,1122]" gridcols="3" gridrows="8" pageId="7" pageNumber="362">
<tr id="353834F4FFE2007D5F865993FB3C88EE" box="[151,1235,690,743]" gridrow="0" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F865993FE0B88EE" box="[151,484,690,743]" gridcol="0" gridrow="0" pageId="7" pageNumber="362">Characters</th>
<th id="76E95D88FFE2007D5D5D5993FCB588EE" box="[588,858,690,743]" gridcol="1" gridrow="0" pageId="7" pageNumber="362">
<taxonomicName id="4C084D35FFE2FF855D5D5992FD4188C1" box="[588,686,691,712]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855D5D5992FD4188C1" box="[588,686,691,712]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
(n=37) Mean ± standard deviation
</th>
<th id="76E95D88FFE2007D5CE95993FB3C88EE" box="[1016,1235,690,743]" gridcol="2" gridrow="0" pageId="7" pageNumber="362">
<taxonomicName id="4C084D35FFE2FF855CE95992FB7A88C1" box="[1016,1173,690,712]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B97CEAA4FFE2FF855CE95992FB7A88C1" box="[1016,1173,690,712]" italics="true" pageId="7" pageNumber="362">H. angustifrons</emphasis>
</taxonomicName>
(n=1)
</th>
</tr>
<tr id="353834F4FFE2007D5F8659DBFB3C8919" box="[151,1235,762,784]" gridrow="1" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F8659DBFE0B8919" box="[151,484,762,784]" gridcol="0" gridrow="1" pageId="7" pageNumber="362">TL</th>
<td id="76E95D88FFE2007D5D5D59DBFCB58919" box="[588,858,762,784]" gridcol="1" gridrow="1" pageId="7" pageNumber="362">3.10 ± 0.18</td>
<td id="76E95D88FFE2007D5CE959DBFB3C8919" box="[1016,1235,762,784]" gridcol="2" gridrow="1" pageId="7" pageNumber="362">3.57</td>
</tr>
<tr id="353834F4FFE2007D5F865805FB3C8933" box="[151,1235,804,826]" gridrow="2" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F865805FE0B8933" box="[151,484,804,826]" gridcol="0" gridrow="2" pageId="7" pageNumber="362">MW</th>
<td id="76E95D88FFE2007D5D5D5805FCB58933" box="[588,858,804,826]" gridcol="1" gridrow="2" pageId="7" pageNumber="362">1.71 ± 0.10</td>
<td id="76E95D88FFE2007D5CE95805FB3C8933" box="[1016,1235,804,826]" gridcol="2" gridrow="2" pageId="7" pageNumber="362">2</td>
</tr>
<tr id="353834F4FFE2007D5F86586CFB3C896A" box="[151,1235,845,867]" gridrow="3" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F86586CFE0B896A" box="[151,484,845,867]" gridcol="0" gridrow="3" pageId="7" pageNumber="362">PL</th>
<td id="76E95D88FFE2007D5D5D586CFCB5896A" box="[588,858,845,867]" gridcol="1" gridrow="3" pageId="7" pageNumber="362">0.65 ± 0.05</td>
<td id="76E95D88FFE2007D5CE9586CFB3C896A" box="[1016,1235,845,867]" gridcol="2" gridrow="3" pageId="7" pageNumber="362">0.7</td>
</tr>
<tr id="353834F4FFE2007D5F865857FB3C8985" box="[151,1235,886,908]" gridrow="4" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F865857FE0B8985" box="[151,484,886,908]" gridcol="0" gridrow="4" pageId="7" pageNumber="362">PW</th>
<td id="76E95D88FFE2007D5D5D5857FCB58985" box="[588,858,886,908]" gridcol="1" gridrow="4" pageId="7" pageNumber="362">1.37 ± 0.08</td>
<td id="76E95D88FFE2007D5CE95857FB3C8985" box="[1016,1235,886,908]" gridcol="2" gridrow="4" pageId="7" pageNumber="362">1.58</td>
</tr>
<tr id="353834F4FFE2007D5F865881FB3C89BF" box="[151,1235,928,950]" gridrow="5" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F865881FE0B89BF" box="[151,484,928,950]" gridcol="0" gridrow="5" pageId="7" pageNumber="362">DE</th>
<td id="76E95D88FFE2007D5D5D5881FCB589BF" box="[588,858,928,950]" gridcol="1" gridrow="5" pageId="7" pageNumber="362">0.26 ± 0.02</td>
<td id="76E95D88FFE2007D5CE95881FB3C89BF" box="[1016,1235,928,950]" gridcol="2" gridrow="5" pageId="7" pageNumber="362">0.216</td>
</tr>
<tr id="353834F4FFE2007D5F8658E8FB3C89D6" box="[151,1235,969,991]" gridrow="6" pageId="7" pageNumber="362">
<th id="76E95D88FFE2007D5F8658E8FE0B89D6" box="[151,484,969,991]" gridcol="0" gridrow="6" pageId="7" pageNumber="362">EW</th>
<td id="76E95D88FFE2007D5D5D58E8FCB589D6" box="[588,858,969,991]" gridcol="1" gridrow="6" pageId="7" pageNumber="362">0.23 ± 0.03</td>
<td id="76E95D88FFE2007D5CE958E8FB3C89D6" box="[1016,1235,969,991]" gridcol="2" gridrow="6" pageId="7" pageNumber="362">0.3</td>
</tr>
<tr id="353834F4FFE2007D5F865F66FB3C8E6B" box="[151,1235,1095,1122]" gridrow="7" pageId="7" pageNumber="362" rowspan-1="1" rowspan-2="1">
<th id="76E95D88FFE2007D5F865F66FE0B8E6B" box="[151,484,1095,1122]" gridcol="0" gridrow="7" pageId="7" pageNumber="362">
Key to the
<taxonomicName id="4C084D35FFE2FF855E0E5F66FE688E68" box="[287,391,1095,1121]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAA4FFE2FF855E0E5F66FE688E68" bold="true" box="[287,391,1095,1121]" italics="true" pageId="7" pageNumber="362">Haliplus</emphasis>
</taxonomicName>
species
</th>
</tr>
</table>
</paragraph>
<subSubSection id="C312653DFFE2FF855F865FAFFA738F5D" pageId="7" pageNumber="362" type="key">
<key id="FF996187FFE2FF855F865FAFFA738F5D" pageId="7" pageNumber="362">
<keyStep id="30FC2E13FFE2FF855F865FAFFA738F5D" pageId="7" pageNumber="362">
<paragraph id="8BB736B6FFE2FF855F865FAFFA738EE0" blockId="7.[151,1436,1164,1364]" pageId="7" pageNumber="362">
<keyLead id="30F99583FFE2FF855F865FAFFA738EE0" pageId="7" pageNumber="362">
1. Pronotal base protruding (
<figureCitation id="13332A33FFE2FF855ECB5FAFFDC38EAD" box="[474,556,1166,1188]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
A), 1st row of secondary punctures on elytra dense and sometimes irregular at base, distance between eyes is 0.7 times width of one eye, prosternal process anteriorly as wide as posteriorly, anterolateral metasternal pits deeper and more separated (
<figureCitation id="13332A33FFE2FF855EFC5FF5FDD18EE3" box="[493,574,1236,1258]" captionStart="FIGURE 5. H" captionStartId="6.[151,250,1575,1597]" captionTargetBox="[158,1431,211,1538]" captionTargetId="figure@6.[151,1436,195,1551]" captionTargetPageId="6" captionText="FIGURE 5. H. angustifrons, A &amp; B—Dorsal &amp; ventral habitus; C—male genitalia (scale 100 µm); D—SEM of anterolateral metasternal pits; E—SEM of male protarsi." httpUri="https://zenodo.org/record/262469/files/figure.png" pageId="7" pageNumber="362">Figure 5</figureCitation>
D)........................................................
<taxonomicName id="4C084D35FFE2FF855A175FF5FA738EE0" box="[1286,1436,1236,1257]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="B97CEAA4FFE2FF855A175FF5FA738EE0" box="[1286,1436,1236,1257]" italics="true" pageId="7" pageNumber="362">H. angustifrons</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB736B6FFE2FF855F865FDBFA738F5D" blockId="7.[151,1436,1164,1364]" pageId="7" pageNumber="362">
<keyLead id="30F99583FFE2FF855F865FDBFA738F5D" pageId="7" pageNumber="362">
- Pronotal base not protruding (
<figureCitation id="13332A33FFE2FF855EEE5FD8FDBD8F06" box="[511,594,1273,1295]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[209,1356,187,1897]" captionTargetId="figure@4.[180,1385,158,1929]" captionTargetPageId="4" captionText="FIGURE 3. A—Dighi pond, Pune; B—Savitribai Phule Pune University pond, Pune; H. arrowi, C—dorsal &amp; ventral view (length 3 mm); D—Hind leg (scale 500 µm); E—Abdomen ventral (scale 100 µm); F—median lobe (scale 100 µm); G &amp; Hright &amp; left parameres, respectively (scale 100 µm)." httpUri="https://zenodo.org/record/262468/files/figure.png" pageId="7" pageNumber="362">Figure 3</figureCitation>
C), 1st row of secondary punctures on elytra sparse at base, between eyes distance is 1.1 times width of one eye, prosternal process wider posteriorly than anteriorly, anterolateral metasternal pits shallow and less separated (
<figureCitation id="13332A33FFE2FF855E395E1FFE958F5D" box="[296,378,1342,1364]" captionStart="FIGURE 2. H" captionStartId="3.[151,250,1952,1974]" captionTargetBox="[168,1412,176,1926]" captionTargetId="figure@3.[168,1412,161,1926]" captionTargetPageId="3" captionText="FIGURE 2. H. arrowi (all SEM images), A—Head; B—Pronotum; C—Elytra; D—Elytral epipleura &amp; anterolateral metasternal region; E—pro and metasternal region; F—metacoxal plates; G magnified view of a setiferous puncture; H— Adhesive structures on male protarsomeres." httpUri="https://zenodo.org/record/262467/files/figure.png" pageId="7" pageNumber="362">Figure 2</figureCitation>
D)..............................................................................
<taxonomicName id="4C084D35FFE2FF855A2F5E1EFA738F5D" box="[1342,1436,1343,1364]" class="Insecta" family="Haliplidae" genus="Haliplus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="arrowi">
<emphasis id="B97CEAA4FFE2FF855A2F5E1EFA738F5D" box="[1342,1436,1343,1364]" italics="true" pageId="7" pageNumber="362">H. arrowi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>