treatments-xml/data/03/88/AD/0388AD2AFFF4F23AFF6CFF2D4DD24DF5.xml

<document id="7D5BDC5A3E178955CCE6EFEA9527C8C1" ID-CLB-Dataset="30608" ID-DOI="10.11646/zootaxa.4388.2.5" ID-GBIF-Dataset="35d94d8a-8367-4e0f-8746-9d22bea27b0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187953" ID-ZooBank="857FAFBA-2AC1-4C61-AA6D-2E089342D92B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520060309145" checkinUser="plazi" docAuthor="Kpan, Tokouaho Flora, Kouamé, N’Goran Germain, Barej, Michael F., Adeba, Patrick Joël, Emmrich, Mike, Ofori-Boateng, Caleb &amp; Rödel, Mark-Oliver" docDate="2018" docId="0388AD2AFFF4F23AFF6CFF2D4DD24DF5" docLanguage="en" docName="zootaxa.4388.2.5.pdf" docOrigin="Zootaxa 4388 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynobatrachus tanoeensis Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="233" masterDocId="FFB1D552FFF7F236FFFBFFBA4C214D21" masterDocTitle="A new Puddle Frog, genus Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae), from the eastern part of the Upper Guinea biodiversity hotspot, West Africa" masterLastPageNumber="237" masterPageNumber="221" pageNumber="224" updateTime="1698526925980" updateUser="ExternalLinkService">
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<mods:title id="C7CBC32649529A5F04891D40C7E27CFA">A new Puddle Frog, genus Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae), from the eastern part of the Upper Guinea biodiversity hotspot, West Africa</mods:title>
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<mods:namePart id="57A5E1D34A85217EEE1FDBC675001274">Kpan, Tokouaho Flora</mods:namePart>
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<mods:namePart id="C327DEE48B8EBEA666733BEF777398EB">Kouamé, N’Goran Germain</mods:namePart>
</mods:name>
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<mods:namePart id="AFF591001770872CDA84FDEC039E76A1">Barej, Michael F.</mods:namePart>
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<mods:namePart id="4E1D32515A5D0094E29757FF6325ACE4">Adeba, Patrick Joël</mods:namePart>
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<mods:namePart id="E8BFB4E71B6E90C8F383EEFB9045D95C">Emmrich, Mike</mods:namePart>
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<mods:namePart id="13BAF018DB60EF477CFE37CC862D4861">Ofori-Boateng, Caleb</mods:namePart>
</mods:name>
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<mods:namePart id="F65226E4184C1758F4E1C1F2C65D9849">Rödel, Mark-Oliver</mods:namePart>
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<mods:number id="CD4F5EC3E9EEF929A2710FEF67F3DAC2">4388</mods:number>
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<emphasis id="B955C02EFFF4F235FF6CFF2D4E754D93" bold="true" box="[151,596,151,178]" pageId="3" pageNumber="224">
<taxonomicName id="4C2167BFFFF4F235FF6CFF2D4DCE4D90" ID-CoL="4GWB9" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[151,495,151,177]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF4F235FF6CFF2D4DCE4D90" bold="true" box="[151,495,151,177]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF4F235FE0DFF224E754D93" box="[502,596,152,178]" pageId="3" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E1C3CFFF4F235FF6CFF004CD94DF2" blockId="3.[151,596,151,211]" box="[151,248,186,211]" pageId="3" pageNumber="224">
<figureCitation id="131A00B9FFF4F235FF6CFF004CD94DF2" box="[151,248,186,211]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1911,1933]" captionStartId-1="5.[151,250,668,690]" captionTargetBox-0="[196,1367,193,1876]" captionTargetBox-1="[188,1398,193,647]" captionTargetId-0="figure@4.[196,1367,193,1876]" captionTargetId-1="figure@5.[188,1398,193,647]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." captionText-1="FIGURE 3. Dorsal and ventral views of adult Phrynobatrachus tanoeensis sp. nov. male (ZMB 86965) from Ebonloa, southwestern Ghana." httpUri-0="https://zenodo.org/record/1187957/files/figure.png" httpUri-1="https://zenodo.org/record/1187959/files/figure.png" pageId="3" pageNumber="224">Figs 2–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B4FB7FFF4F235FF6CFEB8484E4F86" pageId="3" pageNumber="224" type="materials_examined">
<paragraph id="8B9E1C3CFFF4F235FF6CFEB84E784C42" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
<materialsCitation id="3B491661FFF4F235FF6CFEB84E784C42" ID-GBIF-Occurrence="1828873297" collectingDate="2010-07-17" collectingDateMax="2010-07-29" collectingDateMin="2010-07-17" collectionCode="ZMB" collectorName="T. F. Kpan &amp; Adeba." country="Ivory Coast" location="GenBank" municipality="Tanoe-Ehy Swamp Forest" pageId="3" pageNumber="224" specimenCount="1" typeStatus="holotype">
<typeStatus id="549AA29EFFF4F235FF6CFEB84D254C3A" box="[151,260,258,283]" pageId="3" pageNumber="224" type="holotype">
<emphasis id="B955C02EFFF4F235FF6CFEB84D254C3A" bold="true" box="[151,260,258,283]" pageId="3" pageNumber="224">Holotype</emphasis>
</typeStatus>
. 
<collectionCode id="ED3084F9FFF4F235FEEAFEB94D6C4C3B" box="[273,333,259,282]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="224">ZMB</collectionCode>
80870 (field and tissue #: YA 14-03, 
<location id="8EFE4AE7FFF4F235FD0BFEB84F7F4C3A" LSID="urn:lsid:plazi:treatment:0388AD2AFFF4F23AFF6CFF2D4DD24DF5:8EFE4AE7FFF4F235FD0BFEB84F7F4C3A" box="[752,862,258,283]" country="Ivory Coast" municipality="Tanoe-Ehy Swamp Forest" name="GenBank" pageId="3" pageNumber="224">GenBank</location>
#: 
<accessionNumber id="947281DFFFF4F235FC8CFEB84FDD4C3A" box="[887,1020,258,283]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG209126" pageId="3" pageNumber="224">MG209126</accessionNumber>
), adult male, 
<collectingCountry id="F3365CACFFF4F235FB6AFEB949364C3A" box="[1169,1303,259,283]" name="Ivory Coast" pageId="3" pageNumber="224">Ivory Coast</collectingCountry>
, 
<collectingMunicipality id="6BFA8646FFF4F235FAD9FEB84D1F4C1F" pageId="3" pageNumber="224">Tanoé-Ehy Swamp Forest</collectingMunicipality>
, 05°11’181’’N, 02°53’011’’W (date WGS84), swamp forest dominated by 
<taxonomicName id="4C2167BFFFF4F235FB5DFE9C48D64C1F" box="[1190,1271,294,318]" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="B955C02EFFF4F235FB5DFE9C48D64C1F" box="[1190,1271,294,318]" italics="true" pageId="3" pageNumber="224">Raphia</emphasis>
</taxonomicName>
palms, 
<collectingDate id="EFDBC314FFF4F235FAADFE9D4D264C42" pageId="3" pageNumber="224" value="2010-07-17" valueMax="2010-07-29" valueMin="2010-07-17">
<date id="FF9F3AFCFFF4F235FAADFE9D4D264C42" pageId="3" pageNumber="224" value="2010-07-17" valueMax="2010-07-29" valueMin="2010-07-17">17–29 July 2010</date>
</collectingDate>
, coll. 
<collectorName id="26D479EAFFF4F235FEB0FEF14D984C43" box="[331,441,331,354]" pageId="3" pageNumber="224">T.F. Kpan</collectorName>
&amp; P.J. 
<collectorName id="26D479EAFFF4F235FDF2FEF04E784C42" box="[521,601,330,355]" pageId="3" pageNumber="224">Adeba.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9E1C3CFFF4F235FF3CFED449B14F36" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
<materialsCitation id="3B491661FFF4F235FF3CFED448584CA6" ID-GBIF-Occurrence="1828873298" box="[199,1145,366,391]" collectingDate="2010-07-17" collectingDateMax="2010-07-29" collectingDateMin="2010-07-17" collectionCode="ZMB" collectorName="T. F. Kpan &amp; Adeba." country="Ivory Coast" location="GenBank" municipality="Tanoe-Ehy Swamp Forest" pageId="3" pageNumber="224" specimenCode="ZMB 80871" specimenCount="1" typeStatus="paratype">
<typeStatus id="549AA29EFFF4F235FF3CFED44D1E4CA6" box="[199,319,366,391]" pageId="3" pageNumber="224" type="paratype">
<emphasis id="B955C02EFFF4F235FF3CFED44D1E4CA6" bold="true" box="[199,319,366,391]" pageId="3" pageNumber="224">Paratypes</emphasis>
</typeStatus>
. 
<specimenCode id="DB87B447FFF4F235FEB6FED54DF94CA6" box="[333,472,367,391]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="224">ZMB 80871</specimenCode>
(YA14-01, 
<accessionNumber id="947281DFFFF4F235FDA5FED44EC54CA6" box="[606,740,366,391]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG209125" pageId="3" pageNumber="224">MG209125</accessionNumber>
), adult male, same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B491661FFF4F235FB7FFED54FD54CEE" ID-GBIF-Occurrence="1828873358" collectingDate="2014-10-01" collectingDateMax="2014-10-03" collectingDateMin="2014-10-01" collectionCode="ZMB" collectorName="T. F. Kpan" country="Ivory Coast" elevation="17" latitude="5.197117" location="Dohouan village" longLatPrecision="1" longitude="-2.8856666" municipality="Nouamou" pageId="3" pageNumber="224" specimenCode="ZMB 86953-86955" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB87B447FFF4F235FB7FFED549424CA6" box="[1156,1379,366,391]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="224">ZMB 86953–86955</specimenCode>
(KK 1–3), three adult males, all with damages of dorsal skin, 
<collectingCountry id="F3365CACFFF4F235FCA4FE294FCF4C8A" box="[863,1006,403,427]" name="Ivory Coast" pageId="3" pageNumber="224">Ivory Coast</collectingCountry>
, 
<location id="8EFE4AE7FFF4F235FBFBFE2848E34C8A" LSID="urn:lsid:plazi:treatment:0388AD2AFFF4F23AFF6CFF2D4DD24DF5:8EFE4AE7FFF4F235FBFBFE2848E34C8A" box="[1024,1218,402,427]" country="Ivory Coast" latitude="5.197117" longLatPrecision="1" longitude="-2.8856666" municipality="Nouamou" name="Dohouan village" pageId="3" pageNumber="224">Dohouan village</location>
forest, 
<collectingMunicipality id="6BFA8646FFF4F235FADCFE2949B94C8B" box="[1319,1432,403,426]" pageId="3" pageNumber="224">Nouamou</collectingMunicipality>
, 
<geoCoordinate id="EE157AFBFFF4F235FF6CFE0C4D074CEE" box="[151,294,438,463]" direction="north" orientation="latitude" pageId="3" pageNumber="224" precision="1" value="5.197117">05°11.827'N</geoCoordinate>
, 
<geoCoordinate id="EE157AFBFFF4F235FECAFE0C4DE44CEE" box="[305,453,438,463]" direction="west" orientation="longitude" pageId="3" pageNumber="224" precision="1" value="-2.8856666">02°53.140'W</geoCoordinate>
, 
<elevation id="000CFB0FFFF4F235FE2BFE0D4E134CEE" box="[464,562,438,463]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="224" unit="m" value="17.0">
<quantity id="4CD9B1D9FFF4F235FE2BFE0D4E284CEF" box="[464,521,439,463]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="224" unit="m" value="17.0">17 m</quantity>
asl
</elevation>
, 
<collectingDate id="EFDBC314FFF4F235FDC6FE0D4F2B4CEE" box="[573,778,438,463]" pageId="3" pageNumber="224" value="2014-10-01" valueMax="2014-10-03" valueMin="2014-10-01">
<date id="FF9F3AFCFFF4F235FDC6FE0D4F2B4CEE" box="[573,778,438,463]" pageId="3" pageNumber="224" value="2014-10-01" valueMax="2014-10-03" valueMin="2014-10-01">1–3 October 2014</date>
</collectingDate>
, coll. 
<collectorName id="26D479EAFFF4F235FCB6FE0D4F9A4CEF" box="[845,955,439,462]" pageId="3" pageNumber="224">T.F. Kpan</collectorName>
&amp; N
</materialsCitation>
.G. Kouamé; ZMB 86956–86960 (KK 4–8; ZMB 86959= 
<accessionNumber id="947281DFFFF4F235FE8FFE614DD84CD2" box="[372,505,475,499]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG209122" pageId="3" pageNumber="224">MG209122</accessionNumber>
), five adult males, ZMB 86961–86963 (KK 9–11; ZMB 8961= 
<accessionNumber id="947281DFFFF4F235FB34FE6149724CD2" box="[1231,1363,475,499]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MG209123" pageId="3" pageNumber="224">MG209123</accessionNumber>
, ZMB 86963= MK209124), three females, 05°11.718'N, 02°52.854'W, 
<quantity id="4CD9B1D9FFF4F235FC91FE454F824F37" box="[874,931,511,535]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="224" unit="m" value="20.0">20 m</quantity>
asl, all other data as in ZMB 86953–86955.
</paragraph>
<paragraph id="8B9E1C3CFFF4F235FF3CFD98484E4F86" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
<emphasis id="B955C02EFFF4F235FF3CFD984D9B4F1A" bold="true" box="[199,442,546,571]" pageId="3" pageNumber="224">Additional material.</emphasis>
ZMB 86964 (KK 12), juvenile, 05°11.718'N, 02°52.854'W, 
<quantity id="4CD9B1D9FFF4F235FB9BFD9948BB4F1B" box="[1120,1178,547,571]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="224" unit="m" value="20.0">20 m</quantity>
asl, all other data as in ZMB 86953–86955; 
<materialsCitation id="3B491661FFF4F235FE7CFDFD484E4F86" ID-GBIF-Occurrence="1828873357" collectionCode="ZMB" collectorName="C. Ofori-Boateng." country="Ghana" elevation="16" latitude="5.033411" location="Essiama" longLatPrecision="1" longitude="-2.5516388" municipality="Ebonloa" pageId="3" pageNumber="224" specimenCode="GAW009" specimenCount="1" stateProvince="Western Region">
<collectionCode id="ED3084F9FFF4F235FE7CFDFD4DE54F7F" box="[391,452,583,606]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="224">ZMB</collectionCode>
86965 (
<specimenCode id="DB87B447FFF4F235FDDFFDFD4EB54F7E" box="[548,660,583,607]" collectionCode="GAW" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12967" name="Eastern Botanical Society of Glasgow" pageId="3" pageNumber="224">GAW009</specimenCode>
), adult male, skin damaged or missing on left flank and right thigh, 
<collectingCountry id="F3365CACFFF4F235FF6CFDD04CC24FA2" box="[151,227,618,643]" name="Ghana" pageId="3" pageNumber="224">Ghana</collectingCountry>
, 
<collectingRegion id="49E5D2DEFFF4F235FF0BFDD14D884FA2" box="[240,425,618,643]" country="Ghana" name="Western" pageId="3" pageNumber="224">
<collectingRegion id="49E5D2DEFFF4F235FF0BFDD14D6A4FA3" box="[240,331,619,642]" country="Sierra Leone" name="Western" pageId="3" pageNumber="224">Western</collectingRegion>
Region
</collectingRegion>
, 
<collectingMunicipality id="6BFA8646FFF4F235FE4CFDD04E364FA2" box="[439,535,618,643]" pageId="3" pageNumber="224">Ebonloa</collectingMunicipality>
, near 
<location id="8EFE4AE7FFF4F235FDA5FDD04E9F4FA2" LSID="urn:lsid:plazi:treatment:0388AD2AFFF4F23AFF6CFF2D4DD24DF5:8EFE4AE7FFF4F235FDA5FDD04E9F4FA2" box="[606,702,618,643]" country="Ghana" latitude="5.033411" longLatPrecision="1" longitude="-2.5516388" municipality="Ebonloa" name="Essiama" pageId="3" pageNumber="224" stateProvince="Western Region">Essiama</location>
, 
<geoCoordinate id="EE157AFBFFF4F235FD37FDD04F4C4FA2" box="[716,877,618,643]" direction="north" orientation="latitude" pageId="3" pageNumber="224" precision="1" value="5.033411">05°02'0.28’’N</geoCoordinate>
, 
<geoCoordinate id="EE157AFBFFF4F235FC80FDD0482F4FA2" box="[891,1038,618,643]" direction="west" orientation="longitude" pageId="3" pageNumber="224" precision="1" value="-2.5516388">02°33'05.9W</geoCoordinate>
, 
<elevation id="000CFB0FFFF4F235FBE6FDD048A54FA2" box="[1053,1156,618,643]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="224" unit="m" value="16.0">
<quantity id="4CD9B1D9FFF4F235FBE6FDD048794FA3" box="[1053,1112,618,643]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="224" unit="m" value="16.0">16 m</quantity>
asl
</elevation>
, peat/swamp forest with slow moving stream near the borders of 
<collectingCountry id="F3365CACFFF4F235FDA0FD344E854F86" box="[603,676,654,679]" name="Ghana" pageId="3" pageNumber="224">Ghana</collectingCountry>
and 
<collectingCountry id="F3365CACFFF4F235FD21FD354F404F86" box="[730,865,655,679]" name="Ivory Coast" pageId="3" pageNumber="224">Ivory Coast</collectingCountry>
, coll. 
<collectorName id="26D479EAFFF4F235FC5FFD35484E4F86" box="[932,1135,654,679]" pageId="3" pageNumber="224">C. Ofori-Boateng.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B4FB7FFF4F235FF3CFD084F1F4AFA" pageId="3" pageNumber="224" type="diagnosis">
<paragraph id="8B9E1C3CFFF4F235FF3CFD084CC24E82" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
<emphasis id="B955C02EFFF4F235FF3CFD084D1B4FEA" bold="true" box="[199,314,690,715]" pageId="3" pageNumber="224">Diagnosis</emphasis>
. The overall body shape and morphology, i.e. the presence of a tarsal tubercle, as well as the genetic data confirm that the new species is a member of the genus 
<taxonomicName id="4C2167BFFFF4F235FCCEFD6C48C64FCE" authorityName="Gunther" authorityYear="1862" box="[821,1255,726,751]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B955C02EFFF4F235FCCEFD6C4FD94FCF" box="[821,1016,726,750]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus</emphasis>
(Phrynobatrachidae)
</taxonomicName>
. From members of the externally-morphologically similar genus 
<taxonomicName id="4C2167BFFFF4F235FD2BFD41483C4E32" authorityName="Smith" authorityYear="1849" box="[720,1053,762,787]" class="Amphibia" family="Arthroleptidae" genus="Arthroleptis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B955C02EFFF4F235FD2BFD414F774E32" box="[720,854,763,787]" italics="true" pageId="3" pageNumber="224">Arthroleptis</emphasis>
(Arthroleptidae)
</taxonomicName>
, the new species differs by: the presence of scapular ridges, pedal webbing and a tarsal tubercle, and the absence of a mid-dorsal skin raphe. From members of the genus 
<taxonomicName id="4C2167BFFFF4F235FE22FCF94E7C4E7A" authorityName="Boulenger" authorityYear="1917" box="[473,605,835,859]" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B955C02EFFF4F235FE22FCF94E7C4E7A" box="[473,605,835,859]" italics="true" pageId="3" pageNumber="224">Ptychadena</emphasis>
</taxonomicName>
(
<taxonomicName id="4C2167BFFFF4F235FD86FCF84F074E7A" authorityName="Dubois" authorityYear="1987" box="[637,806,834,859]" class="Amphibia" family="Ptychadenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="family">Ptychadenidae</taxonomicName>
; juveniles sometimes morphologically similar to 
<taxonomicName id="4C2167BFFFF4F235FF6CFCDC4D7B4E5F" authorityName="Gunther" authorityYear="1862" box="[151,346,870,894]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B955C02EFFF4F235FF6CFCDC4D7B4E5F" box="[151,346,870,894]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus</emphasis>
</taxonomicName>
) they differ by a more rounded and shorter snout and the absence of parallel dorsal glandular ridges.
</paragraph>
<paragraph id="8B9E1C3CFFF4F235FF3CFC14490949BE" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
The new species can be differentiated from other members of the genus 
<taxonomicName id="4C2167BFFFF4F235FBF2FC1448EA4EE7" authorityName="Gunther" authorityYear="1862" box="[1033,1227,942,966]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B955C02EFFF4F235FBF2FC1448EA4EE7" box="[1033,1227,942,966]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus</emphasis>
</taxonomicName>
by molecular and acoustic characters (see below), as well as by the combination of the following morphological characters: adult snout–vent length larger than 
<quantity id="4CD9B1D9FFF4F235FE1EFC4D4E15492F" box="[485,564,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="224" unit="mm" value="20.0">20 mm</quantity>
but smaller than 
<quantity id="4CD9B1D9FFF4F235FD0CFC4D4F68492F" box="[759,841,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="224" unit="mm" value="30.0">30 mm</quantity>
; a dark face mask; absence of a spiny tubercle on the eyelid; narrow and partly indistinct scapular ridges or comma-shaped warts, in particular no long X-shaped pair of dorsal ridges; distinct spinulae on males’ throats and dorsal surfaces; pronounced pedal webbing; finger- and toe tips not enlarged; a black throat in adult males; a white belly with small blackish spotting in both sexes; more than one wide dark cross bar on hind legs; ventral parts of hind legs in both sexes with rosé to reddish colour.
</paragraph>
<paragraph id="8B9E1C3CFFF4F235FF3CFB114F1F4AFA" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="224">
In particular the new species is smaller (&lt;
<quantity id="4CD9B1D9FFF4F235FD59FB114ED349E2" box="[674,754,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="224" unit="mm" value="30.0">30 mm</quantity>
) than 
<taxonomicName id="4C2167BFFFF4F235FCC3FB114FE849E2" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[824,969,1195,1219]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF4F235FCC3FB114FE849E2" box="[824,969,1195,1219]" italics="true" pageId="3" pageNumber="224">P. liberiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF4F235FC04FB11491149E2" authority="(Gunther, 1858)" authorityName="Gunther" authorityYear="1858" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[1023,1328,1194,1219]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="B955C02EFFF4F235FC04FB11485549E2" box="[1023,1140,1195,1219]" italics="true" pageId="3" pageNumber="224">P. plicatus</emphasis>
(Günther, 1858)
</taxonomicName>
; it differs from 
<taxonomicName id="4C2167BFFFF4F235FF2FFB754DD049C6" authority="Perret, 1966" authorityName="Perret" authorityYear="1966" box="[212,497,1230,1255]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B955C02EFFF4F235FF2FFB754D7F49C7" box="[212,350,1230,1254]" italics="true" pageId="3" pageNumber="224">P. annulatus</emphasis>
Perret, 1966
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FE07FB754F1349C6" authority="(Peters, 1863)" authorityName="Peters" authorityYear="1863" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[508,818,1230,1255]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="B955C02EFFF4F235FE07FB754EAC49C7" box="[508,653,1230,1254]" italics="true" pageId="3" pageNumber="224">P. calcaratus</emphasis>
(Peters, 1863)
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FCBBFB75486549C6" authority="Perret, 1988" authorityName="Perret" authorityYear="1988" box="[832,1092,1231,1255]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="taiensis">
<emphasis id="B955C02EFFF4F235FCBBFB754F9149C7" box="[832,944,1231,1254]" italics="true" pageId="3" pageNumber="224">P. taiensis</emphasis>
<bibRefCitation id="EFB061CDFFF4F235FC43FB75486549C6" box="[952,1092,1231,1255]" pageId="3" pageNumber="224" refId="ref7576" refString="Perret, J. - L. (1988) Les especes de Phrynobatrachus (Anura, Ranidae) a eperon palpebral. Archives des Sciences Geneve, 41, 275 - 294." type="journal article">Perret, 1988</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FBB5FB75497049C6" authority="Guibe, 1959" authorityName="Guibe" authorityYear="1959" box="[1102,1361,1230,1255]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="villiersi">
<emphasis id="B955C02EFFF4F235FBB5FB75489E49C7" box="[1102,1215,1230,1254]" italics="true" pageId="3" pageNumber="224">P. villiersi</emphasis>
Guibé, 1959
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF4F235FA7CFB754E61482A" authority="Hillers, Zimkus &amp; Rodel, 2008" authorityName="Hillers, Zimkus &amp; Rodel" authorityYear="2008" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="pintoi">
<emphasis id="B955C02EFFF4F235FA7CFB754CF9482B" italics="true" pageId="3" pageNumber="224">P. pintoi</emphasis>
<bibRefCitation id="EFB061CDFFF4F235FF1BFB484E61482A" box="[224,576,1266,1291]" pageId="3" pageNumber="224" refId="ref7018" refString="Hillers, A., Zimkus, B. &amp; Rodel, M. - O. (2008) A new species of Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae) from north-western Guinea, West Africa. Zootaxa, 1815, 43 - 50." type="journal article">Hillers, Zimkus &amp; Rödel, 2008</bibRefCitation>
</taxonomicName>
by lacking an eyelid cornicle (spine-like wart on upper eyelid); from all the latter species and 
<taxonomicName id="4C2167BFFFF4F235FE9FFAAD4EFB480E" authority="(Chabanaud, 1921)" authorityName="Chabanaud" authorityYear="1921" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[356,730,1302,1327]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="gutturosus">
<emphasis id="B955C02EFFF4F235FE9FFAAD4DD7480F" box="[356,502,1303,1326]" italics="true" pageId="3" pageNumber="224">P. gutturosus</emphasis>
(Chabanaud, 1921)
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FD1CFAAD4922480E" authority="Rodel, Ohler &amp; Hillers, 2010" authorityName="Rodel, Ohler &amp; Hillers" authorityYear="2010" box="[743,1283,1302,1327]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="hieroglyphicus">
<emphasis id="B955C02EFFF4F235FD1CFAAD4F88480F" box="[743,937,1302,1326]" italics="true" pageId="3" pageNumber="224">P. hieroglyphicus</emphasis>
<bibRefCitation id="EFB061CDFFF4F235FC49FAAC4922480E" box="[946,1283,1302,1327]" pageId="3" pageNumber="224" refId="ref8116" refString="Rodel, M. - O., Ohler, A. &amp; Hillers, A. (2010) A new extraordinary Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae) from West Africa. Zoosystematics and Evolution, 86, 257 - 261. https: // doi. org / 10.1002 / zoos. 201000008" type="journal article">Rödel, Ohler &amp; Hillers, 2010</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C2167BFFFF4F235FABBFAAD4D514872" authority="(Chabanaud, 1921)" authorityName="Chabanaud" authorityYear="1921" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="tokba">
<emphasis id="B955C02EFFF4F235FABBFAAD49BD480F" box="[1344,1436,1302,1326]" italics="true" pageId="3" pageNumber="224">P. tokba</emphasis>
(Chabanaud, 1921)
</taxonomicName>
, the new species differs by more extensive webbing; 
<taxonomicName id="4C2167BFFFF4F235FC39FA8148D84872" authorityName="Perret" authorityYear="1966" box="[962,1273,1339,1363]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B955C02EFFF4F235FC39FA8148D84872" box="[962,1273,1339,1363]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus annulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FAFCFA814D504856" authority="(Chabanaud, 1921)" authorityName="Chabanaud" authorityYear="1921" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="fraterculus">
<emphasis id="B955C02EFFF4F235FAFCFA8149BD4872" box="[1287,1436,1339,1363]" italics="true" pageId="3" pageNumber="224">P. fraterculus</emphasis>
(Chabanaud, 1921)
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FE86FAE54E284857" authorityName="Schiotz" authorityYear="1964" box="[381,521,1374,1398]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="ghanensis">
<emphasis id="B955C02EFFF4F235FE86FAE54E284857" box="[381,521,1374,1398]" italics="true" pageId="3" pageNumber="224">P. ghanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FDE3FAE54FC34856" authority="Guibe &amp; Lamotte, 1958" authorityName="Guibe &amp; Lamotte" authorityYear="1958" box="[536,994,1374,1399]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="maculiventris">
<emphasis id="B955C02EFFF4F235FDE3FAE54EEC4857" box="[536,717,1374,1398]" italics="true" pageId="3" pageNumber="224">P. maculiventris</emphasis>
Guibé &amp; Lamotte, 1958
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FC0BFAE5486F4857" authorityName="Hillers, Zimkus &amp; Rodel" authorityYear="2008" box="[1008,1102,1375,1398]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="pintoi">
<emphasis id="B955C02EFFF4F235FC0BFAE5486F4857" box="[1008,1102,1375,1398]" italics="true" pageId="3" pageNumber="224">P. pintoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FBA6FAE548EF4857" authorityName="Perret" authorityYear="1988" box="[1117,1230,1375,1398]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="taiensis">
<emphasis id="B955C02EFFF4F235FBA6FAE548EF4857" box="[1117,1230,1375,1398]" italics="true" pageId="3" pageNumber="224">P. taiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF4F235FAFDFAE549594857" authorityName="Guibe" authorityYear="1959" box="[1286,1400,1374,1398]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="villiersi">
<emphasis id="B955C02EFFF4F235FAFDFAE549594857" box="[1286,1400,1374,1398]" italics="true" pageId="3" pageNumber="224">P. villiersi</emphasis>
</taxonomicName>
all exhibit very distinct, species-specific patterning on the belly, different to the new species which exhibits many small black spots. In contrast 
<taxonomicName id="4C2167BFFFF4F235FE1AFA1D4EFB489E" authority=", Ahl, 1924" authorityName=", Ahl" authorityYear="1924" box="[481,730,1446,1471]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B955C02EFFF4F235FE1AFA1D4E7B489F" box="[481,602,1446,1470]" italics="true" pageId="3" pageNumber="224">P. latifrons</emphasis>
, Ahl, 1924
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FD1EFA1D4FF5489E" authority="Parker, 1936" authorityName="Parker" authorityYear="1936" box="[741,980,1446,1471]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="alleni">
<emphasis id="B955C02EFFF4F235FD1EFA1D4F61489F" box="[741,832,1446,1470]" italics="true" pageId="3" pageNumber="224">P. alleni</emphasis>
Parker, 1936
</taxonomicName>
(only females, males with yellow belly), 
<taxonomicName id="4C2167BFFFF4F235FF6CFA714DCA48C2" authority="Boulenger, 1912" authorityName="Boulenger" authorityYear="1912" box="[151,491,1482,1507]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="francisci">
<emphasis id="B955C02EFFF4F235FF6CFA714D3648C2" box="[151,279,1483,1507]" italics="true" pageId="3" pageNumber="224">P. francisci</emphasis>
Boulenger, 1912
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FE06FA714F6248C2" authority="(Smith, 1849)" authorityName="Smith" authorityYear="1849" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[509,835,1482,1507]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="natalensis">
<emphasis id="B955C02EFFF4F235FE06FA714EB048C2" box="[509,657,1483,1507]" italics="true" pageId="3" pageNumber="224">P. natalensis</emphasis>
(Smith, 1849)
</taxonomicName>
, and 
<taxonomicName id="4C2167BFFFF4F235FC70FA714D7E4B26" authority="Rodel, Onadeko, Barej &amp; Sandberger, 2012" authorityName="Rodel, Onadeko, Barej &amp; Sandberger" authorityYear="2012" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="rainerguentheri">
<emphasis id="B955C02EFFF4F235FC70FA71487C48C2" box="[907,1117,1483,1507]" italics="true" pageId="3" pageNumber="224">P. rainerguentheri</emphasis>
Rödel, Onadeko, 
<bibRefCitation id="EFB061CDFFF4F235FAC7FA704D7E4B26" pageId="3" pageNumber="224" refId="ref7722" refString="Rodel, M. - O., Barej, M. F., Hillers, A., Leache, A. D., Kouame, N. G., Ofori-Boateng, C., Assemian, N. E., Tohe, B., Penner, J., Hirschfeld, M., Doumbia, J., Gonwouo, L. N., Nopper, J., Brede, C., Diaz, R., Fujita, M., Gil, M., Segniagbeto, G. H., Ernst, R. &amp; Sandberger, L. (2012) The genus Astylosternus in the Upper Guinea rainforests, West Africa, with the description of a new species (Amphibia: Anura: Arthroleptidae). Zootaxa, 3245, 1 - 29." type="journal article">Barej &amp; Sandberger, 2012</bibRefCitation>
</taxonomicName>
, have predominantly white bellies with at best a few minute black points. Whereas males of the new species have dark grey to black throats, 
<taxonomicName id="4C2167BFFFF4F235FD72F9A94EC04B0A" authorityName="Parker" authorityYear="1936" box="[649,737,1555,1579]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="alleni">
<emphasis id="B955C02EFFF4F235FD72F9A94EC04B0A" box="[649,737,1555,1579]" italics="true" pageId="3" pageNumber="224">P. alleni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF4F235FCE1F9A94FB24B0A" authorityName=", Ahl" authorityYear="1924" box="[794,915,1555,1579]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B955C02EFFF4F235FCE1F9A94FB24B0A" box="[794,915,1555,1579]" italics="true" pageId="3" pageNumber="224">P. latifrons</emphasis>
</taxonomicName>
males have yellow throats. These two species, along with all species with an eyelid cornicle, as well as 
<emphasis id="B955C02EFFF4F235FCECF98D481D4B6F" box="[791,1084,1591,1615]" italics="true" pageId="3" pageNumber="224">
<taxonomicName id="4C2167BFFFF4F235FCECF98D4FB84B6E" authorityName="Boulenger" authorityYear="1912" box="[791,921,1591,1615]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="francisci">P. francisci</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FC5FF98D48194B6F" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[932,1080,1591,1614]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="gutturosus">P. gutturosus</taxonomicName>
,
</emphasis>
<taxonomicName id="4C2167BFFFF4F235FBBFF98D4CEE4B52" authority="Rodel &amp; Ernst, 2002" authorityName="Rodel &amp; Ernst" authorityYear="2002" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="phyllophilus">
<emphasis id="B955C02EFFF4F235FBBFF98D48CA4B6F" box="[1092,1259,1590,1614]" italics="true" pageId="3" pageNumber="224">P. phyllophilus</emphasis>
<bibRefCitation id="EFB061CDFFF4F235FB09F98C4CEE4B52" pageId="3" pageNumber="224" refId="ref7646" refString="Rodel, M. - O. &amp; Ernst, R. (2002) A new Phrynobatrachus from the Upper Guinean rain forest, West Africa, including a description of a new reproductive mode for the genus. Journal of Herpetology, 36, 561 - 571. https: // doi. org / 10.1670 / 0022 - 1511 (2002) 036 [0561: ANPFTU] 2.0. CO; 2" type="journal article">Rödel &amp; Ernst, 2002</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FF26F9E14EA44B52" authority="Guibe &amp; Lamotte, 1962" authorityName="Guibe &amp; Lamotte" authorityYear="1962" box="[221,645,1626,1651]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="guineensis">
<emphasis id="B955C02EFFF4F235FF26F9E14D514B53" box="[221,368,1627,1650]" italics="true" pageId="3" pageNumber="224">P. guineensis</emphasis>
Guibé &amp; Lamotte, 1962
</taxonomicName>
, 
<emphasis id="B955C02EFFF4F235FD6FF9E148224B52" box="[660,1027,1627,1651]" italics="true" pageId="3" pageNumber="224">
<taxonomicName id="4C2167BFFFF4F235FD6FF9E14F054B52" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[660,804,1627,1651]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="natalensis">P. natalensis</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FCCBF9E14FDE4B52" authorityName="Rodel, Onadeko, Barej &amp; Sandberger" authorityYear="2012" box="[816,1023,1627,1651]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="rainerguentheri">P. rainerguentheri</taxonomicName>
,
</emphasis>
<taxonomicName id="4C2167BFFFF4F235FBF0F9E1496A4B52" authority="Parker, 1936" authorityName="Parker" authorityYear="1936" box="[1035,1355,1626,1651]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="brongersmai">
<emphasis id="B955C02EFFF4F235FBF0F9E148944B52" box="[1035,1205,1627,1651]" italics="true" pageId="3" pageNumber="224">P. brongersmai</emphasis>
Parker, 1936
</taxonomicName>
have a different body shape and a snout which is rounded in dorsal and lateral view (pointed in lateral view in the new species). In 
<taxonomicName id="4C2167BFFFF4F235FEE4F9194D5A4B9A" authorityName="Parker" authorityYear="1936" box="[287,379,1699,1723]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="alleni">
<emphasis id="B955C02EFFF4F235FEE4F9194D5A4B9A" box="[287,379,1699,1723]" italics="true" pageId="3" pageNumber="224">P. alleni</emphasis>
</taxonomicName>
the head is also relatively flatter in lateral view. 
<taxonomicName id="4C2167BFFFF4F235FC58F91948C24B9B" authorityName="Guibe &amp; Lamotte" authorityYear="1962" box="[931,1251,1699,1723]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="guineensis">
<emphasis id="B955C02EFFF4F235FC58F91948C24B9B" box="[931,1251,1699,1723]" italics="true" pageId="3" pageNumber="224">Phrynobatrachus guineensis</emphasis>
</taxonomicName>
is smaller (&lt;
<quantity id="4CD9B1D9FFF4F235FA7BF9194CE54BFE" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="224" unit="mm" value="20.0">20 mm</quantity>
) than the new species, has yellow toe and finger tips and a yellow inguinal patch. 
<taxonomicName id="4C2167BFFFF4F235FB22F97C4D034A22" authorityName="Rodel &amp; Ernst" authorityYear="2002" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="phyllophilus">
<emphasis id="B955C02EFFF4F235FB22F97C4D034A22" italics="true" pageId="3" pageNumber="224">Phrynobatrachus phyllophilus</emphasis>
</taxonomicName>
shows some black spots on belly and pectoral region but has a more compact body shape and roundish snout, toe and finger tips which are enlarged to small round discs (straight in the new species) and usually only one wide black cross bar on thigh and lower leg (several in the new species). In 
<collectingRegion id="49E5D2DEFFF4F235FBECF889486F4A6B" box="[1047,1102,1843,1866]" country="Cameroon" name="West" pageId="3" pageNumber="224">West</collectingRegion>
Africa, only 
<taxonomicName id="4C2167BFFFF4F235FB17F889496A4A6A" authorityName="Parker" authorityYear="1936" box="[1260,1355,1843,1867]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="alleni">
<emphasis id="B955C02EFFF4F235FB17F889496A4A6A" box="[1260,1355,1843,1867]" italics="true" pageId="3" pageNumber="224">P. alleni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF4F235FA7CF8894CD14A4F" baseAuthorityName="Gunther" baseAuthorityYear="1858" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="B955C02EFFF4F235FA7CF8894CD14A4F" italics="true" pageId="3" pageNumber="224">P. plicatus</emphasis>
</taxonomicName>
have scapular ridges formed like an open X, in 
<taxonomicName id="4C2167BFFFF4F235FCF8F8ED4F574A4F" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[771,886,1878,1902]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="B955C02EFFF4F235FCF8F8ED4F574A4F" box="[771,886,1878,1902]" italics="true" pageId="3" pageNumber="224">P. plicatus</emphasis>
</taxonomicName>
this X extends to posterior to half-back. The new species shares a black face mask in 
<collectingRegion id="49E5D2DEFFF4F235FDD3F8C14E7E4AB3" box="[552,607,1915,1938]" country="Cameroon" name="West" pageId="3" pageNumber="224">West</collectingRegion>
Africa only with 
<taxonomicName id="4C2167BFFFF4F235FCDDF8C14FE74AB2" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[806,966,1915,1939]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFF4F235FCDDF8C14FE74AB2" box="[806,966,1915,1939]" italics="true" pageId="3" pageNumber="224">P. intermedius</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2167BFFFF4F235FC2FF8C148694AB2" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[980,1096,1915,1939]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="B955C02EFFF4F235FC2FF8C148694AB2" box="[980,1096,1915,1939]" italics="true" pageId="3" pageNumber="224">P. plicatus</emphasis>
</taxonomicName>
and some 
<taxonomicName id="4C2167BFFFF4F235FB3AF8C149704AB2" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[1217,1361,1915,1939]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF4F235FB3AF8C149704AB2" box="[1217,1361,1915,1939]" italics="true" pageId="3" pageNumber="224">P. liberiensis</emphasis>
</taxonomicName>
and 
<emphasis id="B955C02EFFF4F235FA7CF8C14D6E4A97" italics="true" pageId="3" pageNumber="224">
<taxonomicName id="4C2167BFFFF4F235FA7CF8C14D6A4A97" authorityName="Rodel, Onadeko, Barej &amp; Sandberger" authorityYear="2012" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="rainerguentheri">P. rainerguentheri</taxonomicName>
.
</emphasis>
However, the face mask is always less distinct than in 
<taxonomicName id="4C2167BFFFF4F235FC34F82548674A97" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[975,1094,1950,1974]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="224" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="B955C02EFFF4F235FC34F82548674A97" box="[975,1094,1950,1974]" italics="true" pageId="3" pageNumber="224">P. plicatus</emphasis>
</taxonomicName>
(see discussion for a more in depth comparison with the genetically most similar species).
</paragraph>
</subSubSection>
<caption id="DF5E4CB4FFF3F232FF6CF8CD4D3D4AC8" httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="4" pageNumber="225" startId="4.[151,250,1911,1933]" targetBox="[196,1367,193,1876]" targetPageId="4">
<paragraph id="8B9E1C3CFFF3F232FF6CF8CD4D3D4AC8" blockId="4.[151,1436,1911,2025]" pageId="4" pageNumber="225">
<emphasis id="B955C02EFFF3F232FF6CF8CD4D374AAD" bold="true" box="[151,278,1911,1933]" pageId="4" pageNumber="225">FIGURE 2.</emphasis>
Dorsal and ventral views of 
<taxonomicName id="4C2167BFFFF3F232FDB5F8CD4F524AAF" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[590,883,1911,1934]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="225" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF3F232FDB5F8CD4F524AAF" box="[590,883,1911,1934]" italics="true" pageId="4" pageNumber="225">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF3F232FC87F8C24FF14AAC" box="[892,976,1912,1933]" pageId="4" pageNumber="225" rank="species">
<emphasis id="B955C02EFFF3F232FC87F8C24FF14AAC" bold="true" box="[892,976,1912,1933]" pageId="4" pageNumber="225">sp. nov.</emphasis>
</taxonomicNameLabel>
from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female.
</paragraph>
</caption>
<caption id="DF5E4CB4FFF2F233FF6CFD264D144FF0" httpUri="https://zenodo.org/record/1187959/files/figure.png" pageId="5" pageNumber="226" startId="5.[151,250,668,690]" targetBox="[188,1398,193,647]" targetPageId="5">
<paragraph id="8B9E1C3CFFF2F233FF6CFD264D144FF0" blockId="5.[151,1435,668,721]" pageId="5" pageNumber="226">
<emphasis id="B955C02EFFF2F233FF6CFD264D324F90" bold="true" box="[151,275,668,690]" pageId="5" pageNumber="226">FIGURE 3.</emphasis>
Dorsal and ventral views of adult 
<taxonomicName id="4C2167BFFFF2F233FD8EFD264FB64F93" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[629,919,668,690]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="226" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF2F233FD8EFD264FB64F93" box="[629,919,668,690]" italics="true" pageId="5" pageNumber="226">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF2F233FC66FD274FCF4F93" box="[925,1006,669,690]" pageId="5" pageNumber="226" rank="species">
<emphasis id="B955C02EFFF2F233FC66FD274FCF4F93" bold="true" box="[925,1006,669,690]" pageId="5" pageNumber="226">sp. nov.</emphasis>
</taxonomicNameLabel>
male (ZMB 86965) from Ebonloa, southwestern Ghana.
</paragraph>
</caption>
<subSubSection id="C33B4FB7FFF2F23EFF3CFD4748434E7C" lastPageId="8" lastPageNumber="229" pageId="5" pageNumber="226" type="description">
<paragraph id="8B9E1C3CFFF2F233FF3CFD474D1948BF" blockId="5.[151,1437,765,2015]" pageId="5" pageNumber="226">
<emphasis id="B955C02EFFF2F233FF3CFD474E2C4E37" bold="true" box="[199,525,765,790]" pageId="5" pageNumber="226">
Description of the 
<typeStatus id="549AA29EFFF2F233FE5DFD474E2C4E37" box="[422,525,765,790]" pageId="5" pageNumber="226" type="holotype">holotype</typeStatus>
</emphasis>
(measurements in millimetres mm). Adult male with longish, but compact body shape, short snout and large eyes; snout–vent length 22.1; snout rounded in dorsal, moderately pointed in lateral view; head width 6.9; thigh length 8.5, shorter than crus length, 12.0; foot length including longest toe 15.4; eye diameter 2.8, larger than horizontal diameter of tympanum, 1.6; tympanum distinct, round with round median annuli, diameter 0.5; interorbital distance 2.5; distance eye–nostrils 1.9; distance eye–snout 3.0; nostrils much closer to snout than to eye; canthus rostralis rounded; loreal region straight; distinct, slightly bent supratympanal ridge extending from posterior edge of eye to forearm insertion; dorsal skin fine granular, spinulae on posterior part of back indistinct; spinulae on dorsal surfaces of thighs and lower legs; two pairs of narrow, indistinct, commashaped warts; the anterior pair originating slightly posterior to eyelids, slightly converging and extending to about forearm insertion; the posterior pair being shorter and slightly diverging; a few minute round tubercles on eyelids; throat smooth with parallel skin folds along mandibles; whitish spinulae on throat and anterior part of breast indistinct; skin of other ventral surfaces smooth; palmar surface of hand with large, oval palmar and more slender, longish thenar tubercle; fingers with large, undivided oval subarticular tubercles, additional tubercles on hands absent; finger tips roundish, not enlarged to discs; outer part of thumb with indistinct thin nuptial pad; palmar webbing absent; relative finger length: II = IV ý I &lt;III; small round tarsal tubercle present; large, slender oval inner metatarsal tubercle, 1.1, reaching about half length of shortest toe, 2.2; outer metatarsal tubercle round and minute; subarticular tubercles slender oval, not divided; relative toe length: I &lt;II &lt;V &lt;III &lt;IV; pedal webbing pronounced; webbing formula: I(0.5), II(1-0.5), III(1.5-1), IV(2-2), V(1); toe tips very slightly expanded without forming discs.
</paragraph>
<paragraph id="8B9E1C3CFFF2F233FF3CFA134F424A0B" blockId="5.[151,1437,765,2015]" pageId="5" pageNumber="226">
<emphasis id="B955C02EFFF2F233FF3CFA134E1048E3" bold="true" box="[199,561,1449,1474]" pageId="5" pageNumber="226">
Colour of the 
<typeStatus id="549AA29EFFF2F233FE8FFA134DFA48E3" box="[372,475,1449,1474]" pageId="5" pageNumber="226" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsal surfaces light beige to yellowish brown; top of snout with Y-shaped dark figure – the single arm pointing towards snout, and further dark spots; margin of snout and upper eyelids with ill-defined light band; upper eyelids dark brown, connected by dark brown inter-orbital band, this band bordered anteriorly and posteriorly by narrow clear bands; the two pairs of narrow comma shaped scapular ridges marked darker, the posterior pair connected by V-shaped black marking; a pair of white spots at the tips of this V; posterior back with a few further blackish markings; dorsal surfaces of thighs (4) and lower legs (3) with black cross bars; arms, hands and feet with less distinct black cross bars; lateral side of snout, upper lip and area below supratympanal ridge black with some minute white spots on lip; supratympanal ridge dorsally bordered by white band; posterior to this band a wide black lateral band, followed by further black spots on whitish base colour; lower mandible, throat and anterior pectoral region almost uniform dark grey; belly white, densely beset with black spots; posterior belly and lower surfaces of thighs and lower legs rosé.
</paragraph>
<paragraph id="8B9E1C3CFFF2F230FF3CF88F4E3F4C45" blockId="5.[151,1437,765,2015]" lastBlockId="6.[151,1436,151,501]" lastPageId="6" lastPageNumber="227" pageId="5" pageNumber="226">
<emphasis id="B955C02EFFF2F233FF3CF88F4EB84A6F" bold="true" box="[199,665,1845,1870]" pageId="5" pageNumber="226">
Colour of the 
<typeStatus id="549AA29EFFF2F233FE88F88F4DF84A6F" box="[371,473,1845,1870]" pageId="5" pageNumber="226" type="holotype">holotype</typeStatus>
in preservation
</emphasis>
(after seven years in 75% ethanol). Back light brown, a few white spots where skin is rubbed off; snout tip and upper eyelids with some black mottling; loreal triangle from nostrils to anterior eye and tympanal area darker; scapular ridges and supratympanal ridge darker; black and light lateral pattern faded; upper arm with two dark cross bars, lower side reddish brown; lower arm with one dark cross bar close to hand; palmar surfaces of hands uniform light reddish brown; lower side of mandibles light blackish grey; throat skin dark blackish grey with minute white points; belly and pectoral region whitish with dark brown spotting, denser towards throat and flanks; vent surrounded by flat black triangle, bordered by narrow light line; posterior part of thighs uniform light brown; dorsal part of thighs banded with 3-4 light and dark transversal bands; lower legs with three dark cross bars; foot and toes brown with indistinct darker pattern; ventral parts of thigh whitish beige with brownish mottling; ventral sides of lower legs with very dense reddish brown spotting; part of foot darker brown than dorsal part.
</paragraph>
<paragraph id="8B9E1C3CFFF1F230FF3CFED54DBC4CD5" blockId="6.[151,1436,151,501]" pageId="6" pageNumber="227">
<emphasis id="B955C02EFFF1F230FF3CFED54D144CA9" bold="true" box="[199,309,367,392]" pageId="6" pageNumber="227">Variation</emphasis>
. 
<typeStatus id="549AA29EFFF1F230FEB3FECA4D944CA9" box="[328,437,368,392]" pageId="6" pageNumber="227" type="paratype">Paratypes</typeStatus>
are similar to 
<typeStatus id="549AA29EFFF1F230FD9AFED54EE74CA9" box="[609,710,367,392]" pageId="6" pageNumber="227" type="holotype">holotype</typeStatus>
, but mostly with less contrasting dorsal pattern. Size (SVL) of adult males ranges from 
<quantity id="4CD9B1D9FFF1F230FE54FE2E4E724C8D" box="[431,595,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="2.225" metricValueMax="2.34" metricValueMin="2.11" pageId="6" pageNumber="227" unit="mm" value="22.25" valueMax="23.4" valueMin="21.1">21.1–23.4 mm</quantity>
(mean ± sd: 21.9 ± 
<quantity id="4CD9B1D9FFF1F230FCC3FE2E4FB34C8D" box="[824,914,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="227" unit="mm" value="0.8">0.8 mm</quantity>
; N= 10). They are smaller than females, these ranging from 
<quantity id="4CD9B1D9FFF1F230FECCFE0D4DFC4CEE" box="[311,477,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" metricValueMax="2.66" metricValueMin="2.38" pageId="6" pageNumber="227" unit="mm" value="25.2" valueMax="26.6" valueMin="23.8">23.8–26.6 mm</quantity>
(mean ± sd: 25.6 ± 
<quantity id="4CD9B1D9FFF1F230FD34FE0D4F0D4CEE" box="[719,812,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="227" unit="mm" value="1.5">1.5 mm</quantity>
; N= 3; 
<tableCitation id="C6A32987FFF1F230FC73FE0D48264CF1" box="[904,1031,439,464]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="6.[151,239,548,571]" captionStartId-1="7.[151,239,328,351]" captionText-0="TABLE 1. Measures of the Phrynobatrachus tanoeensis sp. nov. type series; measures in mm; ZMB= ZMB accession number; ref. mat.= referred material; M= male, F= female, j= juvenile; SVL= snout–vent length; HW= head width; FL= thigh length; TL= crus length; FTL= foot length incl. longest toe; ED= horizontal eye diameter; TD= horizontal tympanum diameter; IOD= interorbital distance; EN= distance anterior corner of eye to nostril; ES= distance anterior corner of eye to snout-tip." captionText-1="TABLE 2. Summary measures (in mm) and indices of adult Phrynobatrachus tanoeensis sp. nov., P. intermedius and P. liberiensis; given are mean ± standard deviation and range (min-max); M= male, F= female, N= sample size; SVL= snout–vent length; HW= head width; FL= thigh length; TL= crus length; FTL= foot length incl. longest toe; ED= horizontal eye diameter; TD= horizontal tympanum diameter; IOD= interorbital distance; EN= distance anterior corner of eye to nostril; ES= distance anterior corner of eye to snout-tip." pageId="6" pageNumber="227">Tables 1–2</tableCitation>
). Further measures and indices are summarized in 
<tableCitation id="C6A32987FFF1F230FEBFFE664DB64CD5" box="[324,407,476,501]" captionStart="TABLE 1" captionStartId="6.[151,239,548,571]" captionText="TABLE 1. Measures of the Phrynobatrachus tanoeensis sp. nov. type series; measures in mm; ZMB= ZMB accession number; ref. mat.= referred material; M= male, F= female, j= juvenile; SVL= snout–vent length; HW= head width; FL= thigh length; TL= crus length; FTL= foot length incl. longest toe; ED= horizontal eye diameter; TD= horizontal tympanum diameter; IOD= interorbital distance; EN= distance anterior corner of eye to nostril; ES= distance anterior corner of eye to snout-tip." pageId="6" pageNumber="227">Table 1</tableCitation>
.
</paragraph>
<caption id="DF5E4CB4FFF1F230FF6CFD9E4D8D4F9A" pageId="6" pageNumber="227">
<paragraph id="8B9E1C3CFFF1F230FF6CFD9E4D8D4F9A" blockId="6.[151,1436,547,699]" pageId="6" pageNumber="227">
<emphasis id="B955C02EFFF1F230FF6CFD9E4D2A4F1A" bold="true" box="[151,267,548,571]" pageId="6" pageNumber="227">TABLE 1.</emphasis>
Measures of the 
<taxonomicName id="4C2167BFFFF1F230FE33FD994ED84F1A" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[456,761,547,571]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="227" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF1F230FE33FD994ED84F1A" box="[456,761,547,571]" italics="true" pageId="6" pageNumber="227">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF1F230FCF9FD9E4F794F1A" box="[770,856,548,571]" pageId="6" pageNumber="227" rank="species">
<emphasis id="B955C02EFFF1F230FCF9FD9E4F794F1A" bold="true" box="[770,856,548,571]" pageId="6" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
type series; measures in mm; ZMB= ZMB accession number; ref. mat.= referred material; M= male, F= female, j= juvenile; SVL= snout–vent length; HW= head width; FL= thigh length; TL= crus length; FTL= foot length incl. longest toe; ED= horizontal eye diameter; TD= horizontal tympanum diameter; IOD= interorbital distance; EN= distance anterior corner of eye to nostril; ES= distance anterior corner of eye to snout-tip.
</paragraph>
</caption>
<paragraph id="8B9E1C3CFFF1F230FF64FD6D495E4A27" pageId="6" pageNumber="227">
<table id="F921EE9CFFF10DC9FF64FD6C49B34A27" box="[159,1426,726,1798]" gridcols="14" gridrows="16" pageId="6" pageNumber="227">
<tr id="35111E7EFFF10DC9FF64FD6C49B34FCD" box="[159,1426,726,748]" gridrow="0" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FD6C4CFF4FCD" box="[159,222,726,748]" gridcol="0" gridrow="0" pageId="6" pageNumber="227">ZMB</th>
<th id="76C07702FFF10DC9FEFAFD6C4D784FCD" box="[257,345,726,748]" gridcol="1" gridrow="0" pageId="6" pageNumber="227">status</th>
<th id="76C07702FFF10DC9FE7FFD6C4D874FCD" box="[388,422,726,748]" gridcol="2" gridrow="0" pageId="6" pageNumber="227">sex</th>
<th id="76C07702FFF10DC9FE3AFD6C4DD14FCD" box="[449,496,726,748]" gridcol="3" gridrow="0" pageId="6" pageNumber="227">SVL</th>
<th id="76C07702FFF10DC9FDECFD6C4E604FCD" box="[535,577,726,748]" gridcol="4" gridrow="0" pageId="6" pageNumber="227">HW</th>
<th id="76C07702FFF10DC9FD9CFD6C4EB24FCD" box="[615,659,726,748]" gridcol="5" gridrow="0" pageId="6" pageNumber="227">FL</th>
<th id="76C07702FFF10DC9FD4DFD6C4EC34FCD" box="[694,738,726,748]" gridcol="6" gridrow="0" pageId="6" pageNumber="227">TL</th>
<th id="76C07702FFF10DC9FCFEFD6C4F104FCD" box="[773,817,726,748]" gridcol="7" gridrow="0" pageId="6" pageNumber="227">FTL</th>
<th id="76C07702FFF10DC9FCA0FD6C4F5C4FCD" box="[859,893,726,748]" gridcol="8" gridrow="0" pageId="6" pageNumber="227">ED</th>
<th id="76C07702FFF10DC9FC65FD6C4FE14FCD" box="[926,960,726,748]" gridcol="9" gridrow="0" pageId="6" pageNumber="227">TD</th>
<th id="76C07702FFF10DC9FC1AFD6C482F4FCD" box="[993,1038,726,748]" gridcol="10" gridrow="0" pageId="6" pageNumber="227">IOD</th>
<th id="76C07702FFF10DC9FBCEFD6C48764FCD" box="[1077,1111,726,748]" gridcol="11" gridrow="0" pageId="6" pageNumber="227">EN</th>
<th id="76C07702FFF10DC9FB83FD6C48B94FCD" box="[1144,1176,726,748]" gridcol="12" gridrow="0" pageId="6" pageNumber="227">ES</th>
<th id="76C07702FFF10DC9FB4DFD6C49B34FCD" box="[1206,1426,726,748]" gridcol="13" gridrow="0" pageId="6" pageNumber="227">Webbing formula</th>
</tr>
<tr id="35111E7EFFF10DC9FF64FCBA49B34E15" box="[159,1426,768,820]" gridrow="1" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FCBA4CFF4E15" box="[159,222,768,820]" gridcol="0" gridrow="1" pageId="6" pageNumber="227">80870</th>
<td id="76C07702FFF10DC9FEFAFCBA4D784E15" box="[257,345,768,820]" gridcol="1" gridrow="1" pageId="6" pageNumber="227">holotype</td>
<td id="76C07702FFF10DC9FE7FFCBA4D874E15" box="[388,422,768,820]" gridcol="2" gridrow="1" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFCBA4DD14E15" box="[449,496,768,820]" gridcol="3" gridrow="1" pageId="6" pageNumber="227">21.4</td>
<td id="76C07702FFF10DC9FDECFCBA4E604E15" box="[535,577,768,820]" gridcol="4" gridrow="1" pageId="6" pageNumber="227">6.9</td>
<td id="76C07702FFF10DC9FD9CFCBA4EB24E15" box="[615,659,768,820]" gridcol="5" gridrow="1" pageId="6" pageNumber="227">8.5</td>
<td id="76C07702FFF10DC9FD4DFCBA4EC34E15" box="[694,738,768,820]" gridcol="6" gridrow="1" pageId="6" pageNumber="227">12.0</td>
<td id="76C07702FFF10DC9FCFEFCBA4F104E15" box="[773,817,768,820]" gridcol="7" gridrow="1" pageId="6" pageNumber="227">15.4</td>
<td id="76C07702FFF10DC9FCA0FCBA4F5C4E15" box="[859,893,768,820]" gridcol="8" gridrow="1" pageId="6" pageNumber="227">2.8</td>
<td id="76C07702FFF10DC9FC65FCBA4FE14E15" box="[926,960,768,820]" gridcol="9" gridrow="1" pageId="6" pageNumber="227">1.6</td>
<td id="76C07702FFF10DC9FC1AFCBA482F4E15" box="[993,1038,768,820]" gridcol="10" gridrow="1" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FBCEFCBA48764E15" box="[1077,1111,768,820]" gridcol="11" gridrow="1" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FB83FCBA48B94E15" box="[1144,1176,768,820]" gridcol="12" gridrow="1" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FB4DFCBA49B34E15" box="[1206,1426,768,820]" gridcol="13" gridrow="1" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FCF249B34E5D" box="[159,1426,840,892]" gridrow="2" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FCF24CFF4E5D" box="[159,222,840,892]" gridcol="0" gridrow="2" pageId="6" pageNumber="227">80871</th>
<td id="76C07702FFF10DC9FEFAFCF24D784E5D" box="[257,345,840,892]" gridcol="1" gridrow="2" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFCF24D874E5D" box="[388,422,840,892]" gridcol="2" gridrow="2" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFCF24DD14E5D" box="[449,496,840,892]" gridcol="3" gridrow="2" pageId="6" pageNumber="227">21.6</td>
<td id="76C07702FFF10DC9FDECFCF24E604E5D" box="[535,577,840,892]" gridcol="4" gridrow="2" pageId="6" pageNumber="227">6.9</td>
<td id="76C07702FFF10DC9FD9CFCF24EB24E5D" box="[615,659,840,892]" gridcol="5" gridrow="2" pageId="6" pageNumber="227">8.1</td>
<td id="76C07702FFF10DC9FD4DFCF24EC34E5D" box="[694,738,840,892]" gridcol="6" gridrow="2" pageId="6" pageNumber="227">11.7</td>
<td id="76C07702FFF10DC9FCFEFCF24F104E5D" box="[773,817,840,892]" gridcol="7" gridrow="2" pageId="6" pageNumber="227">16.6</td>
<td id="76C07702FFF10DC9FCA0FCF24F5C4E5D" box="[859,893,840,892]" gridcol="8" gridrow="2" pageId="6" pageNumber="227">3.4</td>
<td id="76C07702FFF10DC9FC65FCF24FE14E5D" box="[926,960,840,892]" gridcol="9" gridrow="2" pageId="6" pageNumber="227">1.7</td>
<td id="76C07702FFF10DC9FC1AFCF2482F4E5D" box="[993,1038,840,892]" gridcol="10" gridrow="2" pageId="6" pageNumber="227">2.6</td>
<td id="76C07702FFF10DC9FBCEFCF248764E5D" box="[1077,1111,840,892]" gridcol="11" gridrow="2" pageId="6" pageNumber="227">2.3</td>
<td id="76C07702FFF10DC9FB83FCF248B94E5D" box="[1144,1176,840,892]" gridcol="12" gridrow="2" pageId="6" pageNumber="227">3.7</td>
<td id="76C07702FFF10DC9FB4DFCF249B34E5D" box="[1206,1426,840,892]" gridcol="13" gridrow="2" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2.5);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FC2A49B34EE5" box="[159,1426,912,964]" gridrow="3" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FC2A4CFF4EE5" box="[159,222,912,964]" gridcol="0" gridrow="3" pageId="6" pageNumber="227">86962</th>
<td id="76C07702FFF10DC9FEFAFC2A4D784EE5" box="[257,345,912,964]" gridcol="1" gridrow="3" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFC2A4D874EE5" box="[388,422,912,964]" gridcol="2" gridrow="3" pageId="6" pageNumber="227">F</td>
<td id="76C07702FFF10DC9FE3AFC2A4DD14EE5" box="[449,496,912,964]" gridcol="3" gridrow="3" pageId="6" pageNumber="227">23.8</td>
<td id="76C07702FFF10DC9FDECFC2A4E604EE5" box="[535,577,912,964]" gridcol="4" gridrow="3" pageId="6" pageNumber="227">8.5</td>
<td id="76C07702FFF10DC9FD9CFC2A4EB24EE5" box="[615,659,912,964]" gridcol="5" gridrow="3" pageId="6" pageNumber="227">8.6</td>
<td id="76C07702FFF10DC9FD4DFC2A4EC34EE5" box="[694,738,912,964]" gridcol="6" gridrow="3" pageId="6" pageNumber="227">12.4</td>
<td id="76C07702FFF10DC9FCFEFC2A4F104EE5" box="[773,817,912,964]" gridcol="7" gridrow="3" pageId="6" pageNumber="227">18.1</td>
<td id="76C07702FFF10DC9FCA0FC2A4F5C4EE5" box="[859,893,912,964]" gridcol="8" gridrow="3" pageId="6" pageNumber="227">3.1</td>
<td id="76C07702FFF10DC9FC65FC2A4FE14EE5" box="[926,960,912,964]" gridcol="9" gridrow="3" pageId="6" pageNumber="227">2.4</td>
<td id="76C07702FFF10DC9FC1AFC2A482F4EE5" box="[993,1038,912,964]" gridcol="10" gridrow="3" pageId="6" pageNumber="227">2.2</td>
<td id="76C07702FFF10DC9FBCEFC2A48764EE5" box="[1077,1111,912,964]" gridcol="11" gridrow="3" pageId="6" pageNumber="227">2.0</td>
<td id="76C07702FFF10DC9FB83FC2A48B94EE5" box="[1144,1176,912,964]" gridcol="12" gridrow="3" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FB4DFC2A49B34EE5" box="[1206,1426,912,964]" gridcol="13" gridrow="3" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(2-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FC6249B3492D" box="[159,1426,984,1036]" gridrow="4" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FC624CFF492D" box="[159,222,984,1036]" gridcol="0" gridrow="4" pageId="6" pageNumber="227">86963</th>
<td id="76C07702FFF10DC9FEFAFC624D78492D" box="[257,345,984,1036]" gridcol="1" gridrow="4" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFC624D87492D" box="[388,422,984,1036]" gridcol="2" gridrow="4" pageId="6" pageNumber="227">F</td>
<td id="76C07702FFF10DC9FE3AFC624DD1492D" box="[449,496,984,1036]" gridcol="3" gridrow="4" pageId="6" pageNumber="227">26.4</td>
<td id="76C07702FFF10DC9FDECFC624E60492D" box="[535,577,984,1036]" gridcol="4" gridrow="4" pageId="6" pageNumber="227">9.2</td>
<td id="76C07702FFF10DC9FD9CFC624EB2492D" box="[615,659,984,1036]" gridcol="5" gridrow="4" pageId="6" pageNumber="227">10.2</td>
<td id="76C07702FFF10DC9FD4DFC624EC3492D" box="[694,738,984,1036]" gridcol="6" gridrow="4" pageId="6" pageNumber="227">13.5</td>
<td id="76C07702FFF10DC9FCFEFC624F10492D" box="[773,817,984,1036]" gridcol="7" gridrow="4" pageId="6" pageNumber="227">20.1</td>
<td id="76C07702FFF10DC9FCA0FC624F5C492D" box="[859,893,984,1036]" gridcol="8" gridrow="4" pageId="6" pageNumber="227">3.2</td>
<td id="76C07702FFF10DC9FC65FC624FE1492D" box="[926,960,984,1036]" gridcol="9" gridrow="4" pageId="6" pageNumber="227">1.8</td>
<td id="76C07702FFF10DC9FC1AFC62482F492D" box="[993,1038,984,1036]" gridcol="10" gridrow="4" pageId="6" pageNumber="227">2.9</td>
<td id="76C07702FFF10DC9FBCEFC624876492D" box="[1077,1111,984,1036]" gridcol="11" gridrow="4" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FB83FC6248B9492D" box="[1144,1176,984,1036]" gridcol="12" gridrow="4" pageId="6" pageNumber="227">3.3</td>
<td id="76C07702FFF10DC9FB4DFC6249B3492D" box="[1206,1426,984,1036]" gridcol="13" gridrow="4" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FB9A49B34975" box="[159,1426,1056,1108]" gridrow="5" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FB9A4CFF4975" box="[159,222,1056,1108]" gridcol="0" gridrow="5" pageId="6" pageNumber="227">86961</th>
<td id="76C07702FFF10DC9FEFAFB9A4D784975" box="[257,345,1056,1108]" gridcol="1" gridrow="5" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFB9A4D874975" box="[388,422,1056,1108]" gridcol="2" gridrow="5" pageId="6" pageNumber="227">F</td>
<td id="76C07702FFF10DC9FE3AFB9A4DD14975" box="[449,496,1056,1108]" gridcol="3" gridrow="5" pageId="6" pageNumber="227">26.6</td>
<td id="76C07702FFF10DC9FDECFB9A4E604975" box="[535,577,1056,1108]" gridcol="4" gridrow="5" pageId="6" pageNumber="227">9.0</td>
<td id="76C07702FFF10DC9FD9CFB9A4EB24975" box="[615,659,1056,1108]" gridcol="5" gridrow="5" pageId="6" pageNumber="227">10.0</td>
<td id="76C07702FFF10DC9FD4DFB9A4EC34975" box="[694,738,1056,1108]" gridcol="6" gridrow="5" pageId="6" pageNumber="227">14.1</td>
<td id="76C07702FFF10DC9FCFEFB9A4F104975" box="[773,817,1056,1108]" gridcol="7" gridrow="5" pageId="6" pageNumber="227">21.1</td>
<td id="76C07702FFF10DC9FCA0FB9A4F5C4975" box="[859,893,1056,1108]" gridcol="8" gridrow="5" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FC65FB9A4FE14975" box="[926,960,1056,1108]" gridcol="9" gridrow="5" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FC1AFB9A482F4975" box="[993,1038,1056,1108]" gridcol="10" gridrow="5" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FBCEFB9A48764975" box="[1077,1111,1056,1108]" gridcol="11" gridrow="5" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FB83FB9A48B94975" box="[1144,1176,1056,1108]" gridcol="12" gridrow="5" pageId="6" pageNumber="227">2.7</td>
<td id="76C07702FFF10DC9FB4DFB9A49B34975" box="[1206,1426,1056,1108]" gridcol="13" gridrow="5" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FBD249B3495F" box="[159,1426,1128,1150]" gridrow="6" pageId="6" pageNumber="227" rowspan-13="1">
<th id="76C07702FFF10DC9FF64FBD24CFF495F" box="[159,222,1128,1150]" gridcol="0" gridrow="6" pageId="6" pageNumber="227">86964</th>
<td id="76C07702FFF10DC9FEFAFBD24D78495F" box="[257,345,1128,1150]" gridcol="1" gridrow="6" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFBD24D87495F" box="[388,422,1128,1150]" gridcol="2" gridrow="6" pageId="6" pageNumber="227">j</td>
<td id="76C07702FFF10DC9FE3AFBD24DD1495F" box="[449,496,1128,1150]" gridcol="3" gridrow="6" pageId="6" pageNumber="227">13.9</td>
<td id="76C07702FFF10DC9FDECFBD24E60495F" box="[535,577,1128,1150]" gridcol="4" gridrow="6" pageId="6" pageNumber="227">3.7</td>
<td id="76C07702FFF10DC9FD9CFBD24EB2495F" box="[615,659,1128,1150]" gridcol="5" gridrow="6" pageId="6" pageNumber="227">6.8</td>
<td id="76C07702FFF10DC9FD4DFBD24EC3495F" box="[694,738,1128,1150]" gridcol="6" gridrow="6" pageId="6" pageNumber="227">7.5</td>
<td id="76C07702FFF10DC9FCFEFBD24F10495F" box="[773,817,1128,1150]" gridcol="7" gridrow="6" pageId="6" pageNumber="227">10.8</td>
<td id="76C07702FFF10DC9FCA0FBD24F5C495F" box="[859,893,1128,1150]" gridcol="8" gridrow="6" pageId="6" pageNumber="227">1.5</td>
<td id="76C07702FFF10DC9FC65FBD24FE1495F" box="[926,960,1128,1150]" gridcol="9" gridrow="6" pageId="6" pageNumber="227">0.9</td>
<td id="76C07702FFF10DC9FC1AFBD2482F495F" box="[993,1038,1128,1150]" gridcol="10" gridrow="6" pageId="6" pageNumber="227">1.7</td>
<td id="76C07702FFF10DC9FBCEFBD24876495F" box="[1077,1111,1128,1150]" gridcol="11" gridrow="6" pageId="6" pageNumber="227">1.2</td>
<td id="76C07702FFF10DC9FB83FBD248B9495F" box="[1144,1176,1128,1150]" gridcol="12" gridrow="6" pageId="6" pageNumber="227">1.6</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FB2B49B349E7" box="[159,1426,1169,1222]" gridrow="7" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FB2B4CFF49E7" box="[159,222,1169,1222]" gridcol="0" gridrow="7" pageId="6" pageNumber="227">86953</th>
<td id="76C07702FFF10DC9FEFAFB2B4D7849E7" box="[257,345,1169,1222]" gridcol="1" gridrow="7" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFB2B4D8749E7" box="[388,422,1169,1222]" gridcol="2" gridrow="7" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFB2B4DD149E7" box="[449,496,1169,1222]" gridcol="3" gridrow="7" pageId="6" pageNumber="227">22.9</td>
<td id="76C07702FFF10DC9FDECFB2B4E6049E7" box="[535,577,1169,1222]" gridcol="4" gridrow="7" pageId="6" pageNumber="227">8.0</td>
<td id="76C07702FFF10DC9FD9CFB2B4EB249E7" box="[615,659,1169,1222]" gridcol="5" gridrow="7" pageId="6" pageNumber="227">10.3</td>
<td id="76C07702FFF10DC9FD4DFB2B4EC349E7" box="[694,738,1169,1222]" gridcol="6" gridrow="7" pageId="6" pageNumber="227">12.1</td>
<td id="76C07702FFF10DC9FCFEFB2B4F1049E7" box="[773,817,1169,1222]" gridcol="7" gridrow="7" pageId="6" pageNumber="227">17.5</td>
<td id="76C07702FFF10DC9FCA0FB2B4F5C49E7" box="[859,893,1169,1222]" gridcol="8" gridrow="7" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FC65FB2B4FE149E7" box="[926,960,1169,1222]" gridcol="9" gridrow="7" pageId="6" pageNumber="227">1.6</td>
<td id="76C07702FFF10DC9FC1AFB2B482F49E7" box="[993,1038,1169,1222]" gridcol="10" gridrow="7" pageId="6" pageNumber="227">2.0</td>
<td id="76C07702FFF10DC9FBCEFB2B487649E7" box="[1077,1111,1169,1222]" gridcol="11" gridrow="7" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FB83FB2B48B949E7" box="[1144,1176,1169,1222]" gridcol="12" gridrow="7" pageId="6" pageNumber="227">2.8</td>
<td id="76C07702FFF10DC9FB4DFB2B49B349E7" box="[1206,1426,1169,1222]" gridcol="13" gridrow="7" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FB6349B3482F" box="[159,1426,1241,1294]" gridrow="8" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FB634CFF482F" box="[159,222,1241,1294]" gridcol="0" gridrow="8" pageId="6" pageNumber="227">86954</th>
<td id="76C07702FFF10DC9FEFAFB634D78482F" box="[257,345,1241,1294]" gridcol="1" gridrow="8" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFB634D87482F" box="[388,422,1241,1294]" gridcol="2" gridrow="8" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFB634DD1482F" box="[449,496,1241,1294]" gridcol="3" gridrow="8" pageId="6" pageNumber="227">21.4</td>
<td id="76C07702FFF10DC9FDECFB634E60482F" box="[535,577,1241,1294]" gridcol="4" gridrow="8" pageId="6" pageNumber="227">7.4</td>
<td id="76C07702FFF10DC9FD9CFB634EB2482F" box="[615,659,1241,1294]" gridcol="5" gridrow="8" pageId="6" pageNumber="227">10.1</td>
<td id="76C07702FFF10DC9FD4DFB634EC3482F" box="[694,738,1241,1294]" gridcol="6" gridrow="8" pageId="6" pageNumber="227">12.1</td>
<td id="76C07702FFF10DC9FCFEFB634F10482F" box="[773,817,1241,1294]" gridcol="7" gridrow="8" pageId="6" pageNumber="227">17.3</td>
<td id="76C07702FFF10DC9FCA0FB634F5C482F" box="[859,893,1241,1294]" gridcol="8" gridrow="8" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FC65FB634FE1482F" box="[926,960,1241,1294]" gridcol="9" gridrow="8" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FC1AFB63482F482F" box="[993,1038,1241,1294]" gridcol="10" gridrow="8" pageId="6" pageNumber="227">2.0</td>
<td id="76C07702FFF10DC9FBCEFB634876482F" box="[1077,1111,1241,1294]" gridcol="11" gridrow="8" pageId="6" pageNumber="227">1.7</td>
<td id="76C07702FFF10DC9FB83FB6348B9482F" box="[1144,1176,1241,1294]" gridcol="12" gridrow="8" pageId="6" pageNumber="227">2.7</td>
<td id="76C07702FFF10DC9FB4DFB6349B3482F" box="[1206,1426,1241,1294]" gridcol="13" gridrow="8" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FA9B49B34877" box="[159,1426,1313,1366]" gridrow="9" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FA9B4CFF4877" box="[159,222,1313,1366]" gridcol="0" gridrow="9" pageId="6" pageNumber="227">86955</th>
<td id="76C07702FFF10DC9FEFAFA9B4D784877" box="[257,345,1313,1366]" gridcol="1" gridrow="9" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFA9B4D874877" box="[388,422,1313,1366]" gridcol="2" gridrow="9" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFA9B4DD14877" box="[449,496,1313,1366]" gridcol="3" gridrow="9" pageId="6" pageNumber="227">21.8</td>
<td id="76C07702FFF10DC9FDECFA9B4E604877" box="[535,577,1313,1366]" gridcol="4" gridrow="9" pageId="6" pageNumber="227">7.7</td>
<td id="76C07702FFF10DC9FD9CFA9B4EB24877" box="[615,659,1313,1366]" gridcol="5" gridrow="9" pageId="6" pageNumber="227">9.5</td>
<td id="76C07702FFF10DC9FD4DFA9B4EC34877" box="[694,738,1313,1366]" gridcol="6" gridrow="9" pageId="6" pageNumber="227">12.2</td>
<td id="76C07702FFF10DC9FCFEFA9B4F104877" box="[773,817,1313,1366]" gridcol="7" gridrow="9" pageId="6" pageNumber="227">16.9</td>
<td id="76C07702FFF10DC9FCA0FA9B4F5C4877" box="[859,893,1313,1366]" gridcol="8" gridrow="9" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FC65FA9B4FE14877" box="[926,960,1313,1366]" gridcol="9" gridrow="9" pageId="6" pageNumber="227">2.0</td>
<td id="76C07702FFF10DC9FC1AFA9B482F4877" box="[993,1038,1313,1366]" gridcol="10" gridrow="9" pageId="6" pageNumber="227">2.6</td>
<td id="76C07702FFF10DC9FBCEFA9B48764877" box="[1077,1111,1313,1366]" gridcol="11" gridrow="9" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FB83FA9B48B94877" box="[1144,1176,1313,1366]" gridcol="12" gridrow="9" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FB4DFA9B49B34877" box="[1206,1426,1313,1366]" gridcol="13" gridrow="9" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FAD349B348BF" box="[159,1426,1385,1438]" gridrow="10" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FAD34CFF48BF" box="[159,222,1385,1438]" gridcol="0" gridrow="10" pageId="6" pageNumber="227">86956</th>
<td id="76C07702FFF10DC9FEFAFAD34D7848BF" box="[257,345,1385,1438]" gridcol="1" gridrow="10" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFAD34D8748BF" box="[388,422,1385,1438]" gridcol="2" gridrow="10" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFAD34DD148BF" box="[449,496,1385,1438]" gridcol="3" gridrow="10" pageId="6" pageNumber="227">21.8</td>
<td id="76C07702FFF10DC9FDECFAD34E6048BF" box="[535,577,1385,1438]" gridcol="4" gridrow="10" pageId="6" pageNumber="227">8.3</td>
<td id="76C07702FFF10DC9FD9CFAD34EB248BF" box="[615,659,1385,1438]" gridcol="5" gridrow="10" pageId="6" pageNumber="227">7.9</td>
<td id="76C07702FFF10DC9FD4DFAD34EC348BF" box="[694,738,1385,1438]" gridcol="6" gridrow="10" pageId="6" pageNumber="227">10.3</td>
<td id="76C07702FFF10DC9FCFEFAD34F1048BF" box="[773,817,1385,1438]" gridcol="7" gridrow="10" pageId="6" pageNumber="227">16.5</td>
<td id="76C07702FFF10DC9FCA0FAD34F5C48BF" box="[859,893,1385,1438]" gridcol="8" gridrow="10" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FC65FAD34FE148BF" box="[926,960,1385,1438]" gridcol="9" gridrow="10" pageId="6" pageNumber="227">1.8</td>
<td id="76C07702FFF10DC9FC1AFAD3482F48BF" box="[993,1038,1385,1438]" gridcol="10" gridrow="10" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FBCEFAD3487648BF" box="[1077,1111,1385,1438]" gridcol="11" gridrow="10" pageId="6" pageNumber="227">2.3</td>
<td id="76C07702FFF10DC9FB83FAD348B948BF" box="[1144,1176,1385,1438]" gridcol="12" gridrow="10" pageId="6" pageNumber="227">2.9</td>
<td id="76C07702FFF10DC9FB4DFAD349B348BF" box="[1206,1426,1385,1438]" gridcol="13" gridrow="10" pageId="6" pageNumber="227">1(0.5);2(1-0.5); (1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FA0B49B348C7" box="[159,1426,1457,1510]" gridrow="11" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FA0B4CFF48C7" box="[159,222,1457,1510]" gridcol="0" gridrow="11" pageId="6" pageNumber="227">86957</th>
<td id="76C07702FFF10DC9FEFAFA0B4D7848C7" box="[257,345,1457,1510]" gridcol="1" gridrow="11" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFA0B4D8748C7" box="[388,422,1457,1510]" gridcol="2" gridrow="11" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFA0B4DD148C7" box="[449,496,1457,1510]" gridcol="3" gridrow="11" pageId="6" pageNumber="227">21.3</td>
<td id="76C07702FFF10DC9FDECFA0B4E6048C7" box="[535,577,1457,1510]" gridcol="4" gridrow="11" pageId="6" pageNumber="227">7.3</td>
<td id="76C07702FFF10DC9FD9CFA0B4EB248C7" box="[615,659,1457,1510]" gridcol="5" gridrow="11" pageId="6" pageNumber="227">9.8</td>
<td id="76C07702FFF10DC9FD4DFA0B4EC348C7" box="[694,738,1457,1510]" gridcol="6" gridrow="11" pageId="6" pageNumber="227">11.8</td>
<td id="76C07702FFF10DC9FCFEFA0B4F1048C7" box="[773,817,1457,1510]" gridcol="7" gridrow="11" pageId="6" pageNumber="227">17.2</td>
<td id="76C07702FFF10DC9FCA0FA0B4F5C48C7" box="[859,893,1457,1510]" gridcol="8" gridrow="11" pageId="6" pageNumber="227">2.4</td>
<td id="76C07702FFF10DC9FC65FA0B4FE148C7" box="[926,960,1457,1510]" gridcol="9" gridrow="11" pageId="6" pageNumber="227">1.4</td>
<td id="76C07702FFF10DC9FC1AFA0B482F48C7" box="[993,1038,1457,1510]" gridcol="10" gridrow="11" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FBCEFA0B487648C7" box="[1077,1111,1457,1510]" gridcol="11" gridrow="11" pageId="6" pageNumber="227">1.8</td>
<td id="76C07702FFF10DC9FB83FA0B48B948C7" box="[1144,1176,1457,1510]" gridcol="12" gridrow="11" pageId="6" pageNumber="227">2.8</td>
<td id="76C07702FFF10DC9FB4DFA0B49B348C7" box="[1206,1426,1457,1510]" gridcol="13" gridrow="11" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64FA4349B34B0F" box="[159,1426,1529,1582]" gridrow="12" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64FA434CFF4B0F" box="[159,222,1529,1582]" gridcol="0" gridrow="12" pageId="6" pageNumber="227">86958</th>
<td id="76C07702FFF10DC9FEFAFA434D784B0F" box="[257,345,1529,1582]" gridcol="1" gridrow="12" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FFA434D874B0F" box="[388,422,1529,1582]" gridcol="2" gridrow="12" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AFA434DD14B0F" box="[449,496,1529,1582]" gridcol="3" gridrow="12" pageId="6" pageNumber="227">22.7</td>
<td id="76C07702FFF10DC9FDECFA434E604B0F" box="[535,577,1529,1582]" gridcol="4" gridrow="12" pageId="6" pageNumber="227">8.0</td>
<td id="76C07702FFF10DC9FD9CFA434EB24B0F" box="[615,659,1529,1582]" gridcol="5" gridrow="12" pageId="6" pageNumber="227">9.1</td>
<td id="76C07702FFF10DC9FD4DFA434EC34B0F" box="[694,738,1529,1582]" gridcol="6" gridrow="12" pageId="6" pageNumber="227">10.9</td>
<td id="76C07702FFF10DC9FCFEFA434F104B0F" box="[773,817,1529,1582]" gridcol="7" gridrow="12" pageId="6" pageNumber="227">17.4</td>
<td id="76C07702FFF10DC9FCA0FA434F5C4B0F" box="[859,893,1529,1582]" gridcol="8" gridrow="12" pageId="6" pageNumber="227">2.9</td>
<td id="76C07702FFF10DC9FC65FA434FE14B0F" box="[926,960,1529,1582]" gridcol="9" gridrow="12" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FC1AFA43482F4B0F" box="[993,1038,1529,1582]" gridcol="10" gridrow="12" pageId="6" pageNumber="227">2.3</td>
<td id="76C07702FFF10DC9FBCEFA4348764B0F" box="[1077,1111,1529,1582]" gridcol="11" gridrow="12" pageId="6" pageNumber="227">2.4</td>
<td id="76C07702FFF10DC9FB83FA4348B94B0F" box="[1144,1176,1529,1582]" gridcol="12" gridrow="12" pageId="6" pageNumber="227">3.5</td>
<td id="76C07702FFF10DC9FB4DFA4349B34B0F" box="[1206,1426,1529,1582]" gridcol="13" gridrow="12" pageId="6" pageNumber="227">1(.05);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64F9FB49B34B57" box="[159,1426,1601,1654]" gridrow="13" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64F9FB4CFF4B57" box="[159,222,1601,1654]" gridcol="0" gridrow="13" pageId="6" pageNumber="227">86959</th>
<td id="76C07702FFF10DC9FEFAF9FB4D784B57" box="[257,345,1601,1654]" gridcol="1" gridrow="13" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FF9FB4D874B57" box="[388,422,1601,1654]" gridcol="2" gridrow="13" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AF9FB4DD14B57" box="[449,496,1601,1654]" gridcol="3" gridrow="13" pageId="6" pageNumber="227">23.4</td>
<td id="76C07702FFF10DC9FDECF9FB4E604B57" box="[535,577,1601,1654]" gridcol="4" gridrow="13" pageId="6" pageNumber="227">7.9</td>
<td id="76C07702FFF10DC9FD9CF9FB4EB24B57" box="[615,659,1601,1654]" gridcol="5" gridrow="13" pageId="6" pageNumber="227">8.2</td>
<td id="76C07702FFF10DC9FD4DF9FB4EC34B57" box="[694,738,1601,1654]" gridcol="6" gridrow="13" pageId="6" pageNumber="227">11.8</td>
<td id="76C07702FFF10DC9FCFEF9FB4F104B57" box="[773,817,1601,1654]" gridcol="7" gridrow="13" pageId="6" pageNumber="227">17.9</td>
<td id="76C07702FFF10DC9FCA0F9FB4F5C4B57" box="[859,893,1601,1654]" gridcol="8" gridrow="13" pageId="6" pageNumber="227">3.5</td>
<td id="76C07702FFF10DC9FC65F9FB4FE14B57" box="[926,960,1601,1654]" gridcol="9" gridrow="13" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FC1AF9FB482F4B57" box="[993,1038,1601,1654]" gridcol="10" gridrow="13" pageId="6" pageNumber="227">2.4</td>
<td id="76C07702FFF10DC9FBCEF9FB48764B57" box="[1077,1111,1601,1654]" gridcol="11" gridrow="13" pageId="6" pageNumber="227">2.1</td>
<td id="76C07702FFF10DC9FB83F9FB48B94B57" box="[1144,1176,1601,1654]" gridcol="12" gridrow="13" pageId="6" pageNumber="227">4.0</td>
<td id="76C07702FFF10DC9FB4DF9FB49B34B57" box="[1206,1426,1601,1654]" gridcol="13" gridrow="13" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(2-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64F93349B34B9F" box="[159,1426,1673,1726]" gridrow="14" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64F9334CFF4B9F" box="[159,222,1673,1726]" gridcol="0" gridrow="14" pageId="6" pageNumber="227">86960</th>
<td id="76C07702FFF10DC9FEFAF9334D784B9F" box="[257,345,1673,1726]" gridcol="1" gridrow="14" pageId="6" pageNumber="227">paratype</td>
<td id="76C07702FFF10DC9FE7FF9334D874B9F" box="[388,422,1673,1726]" gridcol="2" gridrow="14" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AF9334DD14B9F" box="[449,496,1673,1726]" gridcol="3" gridrow="14" pageId="6" pageNumber="227">21.1</td>
<td id="76C07702FFF10DC9FDECF9334E604B9F" box="[535,577,1673,1726]" gridcol="4" gridrow="14" pageId="6" pageNumber="227">7.1</td>
<td id="76C07702FFF10DC9FD9CF9334EB24B9F" box="[615,659,1673,1726]" gridcol="5" gridrow="14" pageId="6" pageNumber="227">9.8</td>
<td id="76C07702FFF10DC9FD4DF9334EC34B9F" box="[694,738,1673,1726]" gridcol="6" gridrow="14" pageId="6" pageNumber="227">10.7</td>
<td id="76C07702FFF10DC9FCFEF9334F104B9F" box="[773,817,1673,1726]" gridcol="7" gridrow="14" pageId="6" pageNumber="227">15.4</td>
<td id="76C07702FFF10DC9FCA0F9334F5C4B9F" box="[859,893,1673,1726]" gridcol="8" gridrow="14" pageId="6" pageNumber="227">2.7</td>
<td id="76C07702FFF10DC9FC65F9334FE14B9F" box="[926,960,1673,1726]" gridcol="9" gridrow="14" pageId="6" pageNumber="227">1.4</td>
<td id="76C07702FFF10DC9FC1AF933482F4B9F" box="[993,1038,1673,1726]" gridcol="10" gridrow="14" pageId="6" pageNumber="227">1.9</td>
<td id="76C07702FFF10DC9FBCEF93348764B9F" box="[1077,1111,1673,1726]" gridcol="11" gridrow="14" pageId="6" pageNumber="227">1.8</td>
<td id="76C07702FFF10DC9FB83F93348B94B9F" box="[1144,1176,1673,1726]" gridcol="12" gridrow="14" pageId="6" pageNumber="227">2.9</td>
<td id="76C07702FFF10DC9FB4DF93349B34B9F" box="[1206,1426,1673,1726]" gridcol="13" gridrow="14" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
<tr id="35111E7EFFF10DC9FF64F96B49B34A27" box="[159,1426,1745,1798]" gridrow="15" pageId="6" pageNumber="227">
<th id="76C07702FFF10DC9FF64F96B4CFF4A27" box="[159,222,1745,1798]" gridcol="0" gridrow="15" pageId="6" pageNumber="227">86965</th>
<td id="76C07702FFF10DC9FEFAF96B4D784A27" box="[257,345,1745,1798]" gridcol="1" gridrow="15" pageId="6" pageNumber="227">ref. mat.</td>
<td id="76C07702FFF10DC9FE7FF96B4D874A27" box="[388,422,1745,1798]" gridcol="2" gridrow="15" pageId="6" pageNumber="227">M</td>
<td id="76C07702FFF10DC9FE3AF96B4DD14A27" box="[449,496,1745,1798]" gridcol="3" gridrow="15" pageId="6" pageNumber="227">22.1</td>
<td id="76C07702FFF10DC9FDECF96B4E604A27" box="[535,577,1745,1798]" gridcol="4" gridrow="15" pageId="6" pageNumber="227">6.8</td>
<td id="76C07702FFF10DC9FD9CF96B4EB24A27" box="[615,659,1745,1798]" gridcol="5" gridrow="15" pageId="6" pageNumber="227">10.2</td>
<td id="76C07702FFF10DC9FD4DF96B4EC34A27" box="[694,738,1745,1798]" gridcol="6" gridrow="15" pageId="6" pageNumber="227">10.5</td>
<td id="76C07702FFF10DC9FCFEF96B4F104A27" box="[773,817,1745,1798]" gridcol="7" gridrow="15" pageId="6" pageNumber="227">14.8</td>
<td id="76C07702FFF10DC9FCA0F96B4F5C4A27" box="[859,893,1745,1798]" gridcol="8" gridrow="15" pageId="6" pageNumber="227">2.8</td>
<td id="76C07702FFF10DC9FC65F96B4FE14A27" box="[926,960,1745,1798]" gridcol="9" gridrow="15" pageId="6" pageNumber="227">2.2</td>
<td id="76C07702FFF10DC9FC1AF96B482F4A27" box="[993,1038,1745,1798]" gridcol="10" gridrow="15" pageId="6" pageNumber="227">2.5</td>
<td id="76C07702FFF10DC9FBCEF96B48764A27" box="[1077,1111,1745,1798]" gridcol="11" gridrow="15" pageId="6" pageNumber="227">2.1</td>
<td id="76C07702FFF10DC9FB83F96B48B94A27" box="[1144,1176,1745,1798]" gridcol="12" gridrow="15" pageId="6" pageNumber="227">3.0</td>
<td id="76C07702FFF10DC9FB4DF96B49B34A27" box="[1206,1426,1745,1798]" gridcol="13" gridrow="15" pageId="6" pageNumber="227">1(0.5);2(1-0.5); 3(1.5-1);4(2-2);5(1)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9E1C3CFFF1F231FF3CF8F04F724C3C" blockId="6.[151,1437,1866,2035]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="228" pageId="6" pageNumber="227">
The dorsal skin may vary from almost smooth to granular. Some small warts may be present on lower flanks and posterior back. In most male 
<typeStatus id="549AA29EFFF1F230FDE4F8D54EAC4AA6" box="[543,653,1903,1927]" pageId="6" pageNumber="227" type="paratype">paratypes</typeStatus>
, the posterior half of the back is beset with minute round spinulae. Similar spinulae are present on dorsal surfaces of thigh and lower leg and, less visible, on anterior part of body; the throat and the anterior part of the pectoral region likewise densely beset with whitish, round spinulae. Depending on the preservation the spinulae can be difficult to see. They are best visible when the specimen is slightly dehydrated due to too high ethanol concentration, or after the skin has been carefully towelled. Presumably the distinctiveness of the spinulae may vary as well with the breeding status of the respective male. The spinulae are most pronounced in the male from 
<collectingCountry id="F3365CACFFF0F231FDDDFF654E4E4DD9" box="[550,623,223,248]" name="Ghana" pageId="7" pageNumber="228">Ghana</collectingCountry>
(ZMB 86965; 
<figureCitation id="131A00B9FFF0F231FCE1FF654F7F4DD9" box="[794,862,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,668,690]" captionTargetBox="[188,1398,193,647]" captionTargetId="figure@5.[188,1398,193,647]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal and ventral views of adult Phrynobatrachus tanoeensis sp. nov. male (ZMB 86965) from Ebonloa, southwestern Ghana." httpUri="https://zenodo.org/record/1187959/files/figure.png" pageId="7" pageNumber="228">Fig. 3</figureCitation>
). Here they cover the entire back from vent to just behind the eyes, as well as the dorsal surfaces of the hind legs.
</paragraph>
<caption id="DF5E4CB4FFF0F231FF6CFEF24F6F4CFE" pageId="7" pageNumber="228">
<paragraph id="8B9E1C3CFFF0F231FF6CFEF24F6F4CFE" blockId="7.[151,1437,327,479]" pageId="7" pageNumber="228">
<emphasis id="B955C02EFFF0F231FF6CFEF24D2B4C7E" bold="true" box="[151,266,328,351]" pageId="7" pageNumber="228">TABLE 2.</emphasis>
Summary measures (in mm) and indices of adult 
<taxonomicName id="4C2167BFFFF0F231FCDAFEFD48734C7E" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[801,1106,327,351]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF0F231FCDAFEFD48734C7E" box="[801,1106,327,351]" italics="true" pageId="7" pageNumber="228">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF0F231FBA2FEF2488F4C7E" box="[1113,1198,328,351]" pageId="7" pageNumber="228" rank="species">
<emphasis id="B955C02EFFF0F231FBA2FEF2488F4C7E" bold="true" box="[1113,1198,328,351]" pageId="7" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C2167BFFFF0F231FB47FEF249754C7E" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[1212,1364,327,351]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFF0F231FB47FEF249754C7E" box="[1212,1364,327,351]" italics="true" pageId="7" pageNumber="228">P. intermedius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF0F231FA73FEF24D274C5E" authorityName="Barbour &amp; Loveridge" authorityYear="1927" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FA73FEF24D274C5E" italics="true" pageId="7" pageNumber="228">P. liberiensis</emphasis>
</taxonomicName>
; given are mean ± standard deviation and range (min-max); M= male, F= female, N= sample size; SVL= snout–vent length; HW= head width; FL= thigh length; TL= crus length; FTL= foot length incl. longest toe; ED= horizontal eye diameter; TD= horizontal tympanum diameter; IOD= interorbital distance; EN= distance anterior corner of eye to nostril; ES= distance anterior corner of eye to snout-tip.
</paragraph>
</caption>
<paragraph id="8B9E1C3CFFF0F231FF64FE404CCB4E71" blockId="7.[159,1402,506,931]" pageId="7" pageNumber="228">
species SVL HW FL TL FTL ED TD 
<taxonomicName id="4C2167BFFFF0F231FF64FD9F4D2B4F1B" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[159,266,549,570]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF0F231FF64FD9F4D2B4F1B" box="[159,266,549,570]" italics="true" pageId="7" pageNumber="228">tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF0F231FEEBFD9E4D414F18" box="[272,352,548,569]" pageId="7" pageNumber="228" rank="species">
<emphasis id="B955C02EFFF0F231FEEBFD9E4D414F18" bold="true" box="[272,352,548,569]" pageId="7" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
21.9 ± 0.8 7.5 ± 0.5 9.2 ± 0.9 11.6 ± 0.7 16.8 ± 0.9 2.8 ± 0.4 1.7 ± 0.2 M, N= 10 21.1‒23.4 6.9‒8.3 7.9‒10.3 10.3‒12.2 15.4‒17.9 2.4‒3.5 1.4‒2.0 
<taxonomicName id="4C2167BFFFF0F231FF64FDCC4D284FAD" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[159,265,630,652]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FF64FDCC4D284FAD" box="[159,265,630,652]" italics="true" pageId="7" pageNumber="228">liberiensis</emphasis>
</taxonomicName>
25.3 ± 1.5 8.1 ± 0.7 12.1 ± 1.1 14.3 ± 1.1 20.2 ± 1.6 3.1 ± 0.3 2.2 ± 0.2 M, N= 15 23.5‒28.9 6.2‒9.1 10.8‒14.4 12.8‒17.0 18.1‒24.2 2.6‒3.5 1.8‒2.7 
<taxonomicName id="4C2167BFFFF0F231FF64FD704D2B4FFE" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[159,266,714,735]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF0F231FF64FD704D2B4FFE" box="[159,266,714,735]" italics="true" pageId="7" pageNumber="228">tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF0F231FEEBFD734D414FFF" box="[272,352,713,734]" pageId="7" pageNumber="228" rank="species">
<emphasis id="B955C02EFFF0F231FEEBFD734D414FFF" bold="true" box="[272,352,713,734]" pageId="7" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
25.6 ± 1.5 8.9 ± 0.4 9.6 ± 0.9 13.4 ± 0.9 19.8 ± 1.5 3.1 ± 0.1 2.0 ± 0.4 F, N= 3 23.8‒26.6 8.5‒9.2 8.6‒10.2 12.4‒14.1 18.1‒21.1 3.0‒3.2 1.8‒2.4 
<taxonomicName id="4C2167BFFFF0F231FF64FCA64D364E13" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[159,279,796,818]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFF0F231FF64FCA64D364E13" box="[159,279,796,818]" italics="true" pageId="7" pageNumber="228">intermedius</emphasis>
</taxonomicName>
27.2 7.8 14.4 16.5 20.3 3.4 2.3 F, N= 1
</paragraph>
<paragraph id="8B9E1C3CFFF0F231FF64FCDE494D4E82" blockId="7.[159,1402,506,931]" pageId="7" pageNumber="228">
<taxonomicName id="4C2167BFFFF0F231FF64FCDE4D284E5B" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[159,265,868,890]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FF64FCDE4D284E5B" box="[159,265,868,890]" italics="true" pageId="7" pageNumber="228">liberiensis</emphasis>
</taxonomicName>
31.7 ± 2.4 10.1 ± 0.8 14.7 ± 1.2 17.7 ± 1.1 24.0 ± 1.7 3.3 ± 0.4 2.4 ± 0.3 F, N= 31 25.5‒35.2 8.0‒11.2 11.9‒17.2 13.9‒19.3 19.1‒26.5 2.7‒4.0 1.9‒3.1
</paragraph>
<paragraph id="8B9E1C3CFFF0F231FF6CFC7D4D294EFF" blockId="7.[151,264,967,990]" box="[151,264,967,990]" pageId="7" pageNumber="228">continued.</paragraph>
<paragraph id="8B9E1C3CFFF0F231FF64FC4048B64A45" blockId="7.[159,1386,1018,1205]" lastBlockId="7.[151,1437,1507,2036]" pageId="7" pageNumber="228">
species IOD EN ES HW/ 
<collectingRegion id="49E5D2DEFFF0F231FC7AFC404F914931" box="[897,944,1018,1040]" country="Cameroon" name="South" pageId="7" pageNumber="228">SUL</collectingRegion>
FL/ 
<collectingRegion id="49E5D2DEFFF0F231FBECFC4048674931" box="[1047,1094,1018,1040]" country="Cameroon" name="South" pageId="7" pageNumber="228">SUL</collectingRegion>
TL/ 
<collectingRegion id="49E5D2DEFFF0F231FB53FC4048F64931" box="[1192,1239,1018,1040]" country="Cameroon" name="South" pageId="7" pageNumber="228">SUL</collectingRegion>
EN/ES 
<taxonomicName id="4C2167BFFFF0F231FF64FB9E4D2B4918" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[159,266,1060,1081]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF0F231FF64FB9E4D2B4918" box="[159,266,1060,1081]" italics="true" pageId="7" pageNumber="228">tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF0F231FEEBFB9E4D414918" box="[272,352,1060,1081]" pageId="7" pageNumber="228" rank="species">
<emphasis id="B955C02EFFF0F231FEEBFB9E4D414918" bold="true" box="[272,352,1060,1081]" pageId="7" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
2.3 ± 0.3 2.0 ± 0.2 3.1 ± 0.4 0.3 ± 0 0.4 ± 0 0.5 ± 0 0.6 ± 0.1 M, N= 10 1.9‒2.6 1.7‒2.4 2.7‒4.0 0.3‒04 0.4‒05 0.5‒0.6 0.5‒0.8 
<taxonomicName id="4C2167BFFFF0F231FF64FBCC4D2849AD" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[159,265,1142,1164]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FF64FBCC4D2849AD" box="[159,265,1142,1164]" italics="true" pageId="7" pageNumber="228">liberiensis</emphasis>
</taxonomicName>
2.1 ± 0.2 1.9 ± 0.2 2.9 ± 0.3 0.3 ± 0 0.5 ± 0 0.6 ± 0 0.6 ± 0.1 M, N= 15 1.8‒2.7 1.6‒2.2 2.4‒3.7 0.2‒0.4 0.4‒0.5 0.5‒0.6 0.6‒0.8 The dorsal warts/ridges may vary from very distinct to almost absent (in live and preserved specimens); often only the anterior pair is present and then longer than in the 
<typeStatus id="549AA29EFFF0F231FCBCF9BD4F884B01" box="[839,937,1543,1568]" pageId="7" pageNumber="228" type="holotype">holotype</typeStatus>
(e.g. 
<figureCitation id="131A00B9FFF0F231FC15F9BD484B4B01" box="[1006,1130,1543,1568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">Figs. 2e, g</figureCitation>
). Dorsal and lateral colour varies from dark grey or light to dark brown to almost black, with or without lighter patterning. Sometimes a greenish tinge may be present on the back (
<figureCitation id="131A00B9FFF0F231FD77F9F54EC34B49" box="[652,738,1615,1640]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">Fig. 2d</figureCitation>
). Females exhibit more uniform dorsal coloration than males (
<figureCitation id="131A00B9FFF0F231FF64F9C94CD54BAD" box="[159,244,1651,1676]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">
Fig. 
<quantity id="4CD9B1D9FFF0F231FF2FF9CE4CD54BAD" box="[212,244,1652,1676]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="7" pageNumber="228" unit="g" value="2.0">2g</quantity>
</figureCitation>
). Black lateral bands and white lines may be present (
<figureCitation id="131A00B9FFF0F231FCAFF9C94F904BAD" box="[852,945,1651,1676]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">Figs. 2a</figureCitation>
, c-e) or absent. The dark face mask may be more or less pronounced. The juvenile specimen and some adults exhibit a pair of light dorsal spots posterior to the dorsal ridges (
<figureCitation id="131A00B9FFF0F231FECDF9014D8E4BF5" box="[310,431,1723,1748]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">Figs. 2a, d</figureCitation>
). The Throats of female vary from light grey (
<figureCitation id="131A00B9FFF0F231FC4CF90148284BF5" box="[951,1033,1723,1748]" captionStart="FIGURE 2" captionStartId="4.[151,250,1911,1933]" captionTargetBox="[196,1367,193,1876]" captionTargetId="figure@4.[196,1367,193,1876]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a &amp; b) ZMB 80870, holotype, adult male; c &amp; f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." httpUri="https://zenodo.org/record/1187957/files/figure.png" pageId="7" pageNumber="228">Fig. 2h</figureCitation>
) to brownish dark with white spots. Further greyish or blackish belly spots are particularly dense close to the forearm insertion and towards flanks. The central parts of female bellies are almost uniform white. The ventral surfaces of female hind legs are greyish to orange, in males the colour is reddish beige or rosé. The pedal webbing may vary slightly. In particular toes III and IV may show slightly less or more webbing than the 
<typeStatus id="549AA29EFFF0F231FD1CF8F14F684A45" box="[743,841,1867,1892]" pageId="7" pageNumber="228" type="holotype">holotype</typeStatus>
(in the range of 0.5 phalanx).
</paragraph>
<paragraph id="8B9E1C3CFFF0F231FF64FB704CCB4871" pageId="7" pageNumber="228">
<table id="F921EE9CFFF00DC9FF64FB73494B4871" box="[159,1386,1225,1360]" gridcols="8" gridrows="4" pageId="7" pageNumber="228">
<tr id="35111E7EFFF00DC9FF64FB73494B49FE" box="[159,1386,1225,1247]" gridrow="0" pageId="7" pageNumber="228">
<th id="76C07702FFF00DC9FF64FB734D4149FE" box="[159,352,1225,1247]" gridcol="0" gridrow="0" pageId="7" pageNumber="228">
<taxonomicName id="4C2167BFFFF0F231FF64FB704D2B49FE" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[159,266,1226,1247]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFF0F231FF64FB704D2B49FE" box="[159,266,1226,1247]" italics="true" pageId="7" pageNumber="228">tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFF0F231FEEBFB734D4149FF" box="[272,352,1225,1246]" pageId="7" pageNumber="228" rank="species">
<emphasis id="B955C02EFFF0F231FEEBFB734D4149FF" bold="true" box="[272,352,1225,1246]" pageId="7" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="76C07702FFF00DC9FE60FB734DD449FE" box="[411,501,1225,1247]" gridcol="1" gridrow="0" pageId="7" pageNumber="228">2.5 ± 0.3</th>
<th id="76C07702FFF00DC9FDD1FB734EA549FE" box="[554,644,1225,1247]" gridcol="2" gridrow="0" pageId="7" pageNumber="228">2.3 ± 0.3</th>
<th id="76C07702FFF00DC9FD42FB734F3249FE" box="[697,787,1225,1247]" gridcol="3" gridrow="0" pageId="7" pageNumber="228">3.0 ± 0.3</th>
<th id="76C07702FFF00DC9FCABFB734FBD49FE" box="[848,924,1225,1247]" gridcol="4" gridrow="0" pageId="7" pageNumber="228">0.3 ± 0</th>
<th id="76C07702FFF00DC9FC09FB73481849FE" box="[1010,1081,1225,1247]" gridcol="5" gridrow="0" pageId="7" pageNumber="228">0.4 ± 0</th>
<th id="76C07702FFF00DC9FB7AFB7348E949FE" box="[1153,1224,1225,1247]" gridcol="6" gridrow="0" pageId="7" pageNumber="228">0.5 ± 0</th>
<th id="76C07702FFF00DC9FAEBFB73494B49FE" box="[1296,1386,1225,1247]" gridcol="7" gridrow="0" pageId="7" pageNumber="228">0.8 ± 0.1</th>
</tr>
<tr id="35111E7EFFF00DC9FF64FB48494B4829" box="[159,1386,1266,1288]" gridrow="1" pageId="7" pageNumber="228">
<th id="76C07702FFF00DC9FF64FB484D414829" box="[159,352,1266,1288]" gridcol="0" gridrow="1" pageId="7" pageNumber="228">F, N= 3</th>
<td id="76C07702FFF00DC9FE60FB484DD44829" box="[411,501,1266,1288]" gridcol="1" gridrow="1" pageId="7" pageNumber="228">2.2‒2.9</td>
<td id="76C07702FFF00DC9FDD1FB484EA54829" box="[554,644,1266,1288]" gridcol="2" gridrow="1" pageId="7" pageNumber="228">2.0‒2.5</td>
<td id="76C07702FFF00DC9FD42FB484F324829" box="[697,787,1266,1288]" gridcol="3" gridrow="1" pageId="7" pageNumber="228">2.7‒3.3</td>
<td id="76C07702FFF00DC9FCABFB484FBD4829" box="[848,924,1266,1288]" gridcol="4" gridrow="1" pageId="7" pageNumber="228">0.3‒0.4</td>
<td id="76C07702FFF00DC9FC09FB4848184829" box="[1010,1081,1266,1288]" gridcol="5" gridrow="1" pageId="7" pageNumber="228">0.4</td>
<td id="76C07702FFF00DC9FB7AFB4848E94829" box="[1153,1224,1266,1288]" gridcol="6" gridrow="1" pageId="7" pageNumber="228">0.5</td>
<td id="76C07702FFF00DC9FAEBFB48494B4829" box="[1296,1386,1266,1288]" gridcol="7" gridrow="1" pageId="7" pageNumber="228">0.7‒0.9</td>
</tr>
<table id="F921EE9CFFF00DC9FF64FAA1494B4883" box="[159,1386,1307,1442]" gridrows="4" pageId="7" pageNumber="228">
<tr id="35111E7EFFF00DC9FF64FAA1494B4810" box="[159,1386,1307,1329]" gridrow="2" pageId="7" pageNumber="228">
<th id="76C07702FFF00DC9FF64FAA14D414810" box="[159,352,1307,1329]" gridcol="0" gridrow="2" pageId="7" pageNumber="228">
<taxonomicName id="4C2167BFFFF0F231FF64FAA14D364810" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[159,279,1307,1329]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFF0F231FF64FAA14D364810" box="[159,279,1307,1329]" italics="true" pageId="7" pageNumber="228">intermedius</emphasis>
</taxonomicName>
</th>
<td id="76C07702FFF00DC9FE60FAA14DD44810" box="[411,501,1307,1329]" gridcol="1" gridrow="2" pageId="7" pageNumber="228">2.6</td>
<td id="76C07702FFF00DC9FDD1FAA14EA54810" box="[554,644,1307,1329]" gridcol="2" gridrow="2" pageId="7" pageNumber="228">2.3</td>
<td id="76C07702FFF00DC9FD42FAA14F324810" box="[697,787,1307,1329]" gridcol="3" gridrow="2" pageId="7" pageNumber="228">3.9</td>
<td id="76C07702FFF00DC9FCABFAA14FBD4810" box="[848,924,1307,1329]" gridcol="4" gridrow="2" pageId="7" pageNumber="228">0.3</td>
<td id="76C07702FFF00DC9FC09FAA148184810" box="[1010,1081,1307,1329]" gridcol="5" gridrow="2" pageId="7" pageNumber="228">0.5</td>
<td id="76C07702FFF00DC9FB7AFAA148E94810" box="[1153,1224,1307,1329]" gridcol="6" gridrow="2" pageId="7" pageNumber="228">0.6</td>
<td id="76C07702FFF00DC9FAEBFAA1494B4810" box="[1296,1386,1307,1329]" gridcol="7" gridrow="2" pageId="7" pageNumber="228">0.6</td>
</tr>
<tr id="35111E7EFFF00DC9FF64FA80494B4871" box="[159,1386,1338,1360]" gridrow="3" pageId="7" pageNumber="228" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76C07702FFF00DC9FF64FA804D414871" box="[159,352,1338,1360]" gridcol="0" gridrow="3" pageId="7" pageNumber="228">F, N= 1</th>
</tr>
<paragraph id="8B9E1C3CFFF0F231FF64FAD9497D4883" pageId="7" pageNumber="228">
<taxonomicName id="4C2167BFFFF0F231FF64FAD94D284858" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[159,265,1379,1401]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FF64FAD94D284858" box="[159,265,1379,1401]" italics="true" pageId="7" pageNumber="228">liberiensis</emphasis>
</taxonomicName>
2.8 ± 0.4 2.3 ± 0.3 3.5 ± 0.3 0.3 ± 0 0.5 ± 0 0.6 ± 0 0.7 ± 0.1 F, N= 31 2.1‒3.6 1.7‒3.0 2.8‒4.0 0.3 0.4‒0.5 0.5‒0.6 0.5‒0.8
</paragraph>
</table>
</table>
</paragraph>
<paragraph id="8B9E1C3CFFF0F23EFF3CF8D54E924CA9" blockId="7.[151,1437,1507,2036]" lastBlockId="8.[151,1437,151,861]" lastPageId="8" lastPageNumber="229" pageId="7" pageNumber="228">
<emphasis id="B955C02EFFF0F231FF3CF8D54D144AA9" bold="true" box="[199,309,1903,1928]" pageId="7" pageNumber="228">Genetics.</emphasis>
We compared parts (up to 538 bp) of the 16S RNA gene from five individuals of the new species, with two sequences of 
<taxonomicName id="4C2167BFFFF0F231FE4BF82E4E774A8D" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[432,598,1940,1964]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFF0F231FE4BF82E4E774A8D" box="[432,598,1940,1964]" italics="true" pageId="7" pageNumber="228">P. intermedius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2167BFFFF0F231FD62F82E4ED74A8D" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[665,758,1940,1964]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="tokba">
<emphasis id="B955C02EFFF0F231FD62F82E4ED74A8D" box="[665,758,1940,1964]" italics="true" pageId="7" pageNumber="228">P. tokba</emphasis>
</taxonomicName>
, respectively and, considering the erroneous initial field identification, 27 individuals of 
<taxonomicName id="4C2167BFFFF0F231FDFBF8024EAD4AEE" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[512,652,1975,1999]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FDFBF8024EAD4AEE" box="[512,652,1975,1999]" italics="true" pageId="7" pageNumber="228">P. liberiensis</emphasis>
</taxonomicName>
. The 
<taxonomicName id="4C2167BFFFF0F231FD2AF8024F404AEE" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[721,865,1975,1999]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="228" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFF0F231FD2AF8024F404AEE" box="[721,865,1975,1999]" italics="true" pageId="7" pageNumber="228">P. liberiensis</emphasis>
</taxonomicName>
samples covered almost the entire known range of that species from eastern 
<collectingCountry id="F3365CACFFF0F231FE49F8614E644AD2" box="[434,581,2011,2036]" name="Sierra Leone" pageId="7" pageNumber="228">Sierra Leone</collectingCountry>
, 
<collectingCountry id="F3365CACFFF0F231FDABF8614E854AD5" box="[592,676,2011,2036]" name="Liberia" pageId="7" pageNumber="228">Liberia</collectingCountry>
, south-eastern 
<collectingCountry id="F3365CACFFF0F231FCB1F8614FBF4AD5" box="[842,926,2011,2036]" name="Guinea-Bissau" pageId="7" pageNumber="228">Guinea</collectingCountry>
, western and eastern 
<collectingCountry id="F3365CACFFF0F231FB77F866492E4AD5" box="[1164,1295,2012,2036]" name="Ivory Coast" pageId="7" pageNumber="228">Ivory Coast</collectingCountry>
and western 
<collectingCountry id="F3365CACFFFFF23EFF6CFF2D4CC14D91" box="[151,224,151,176]" name="Ghana" pageId="8" pageNumber="229">Ghana</collectingCountry>
(
<figureCitation id="131A00B9FFFFF23EFF14FF2D4D104D91" box="[239,305,151,176]" captionStart="FIGURE1" captionStartId="2.[151,250,1819,1841]" captionTargetBox="[166,1427,288,1771]" captionTargetId="figure@2.[151,1435,267,1794]" captionTargetPageId="2" captionText="FIGURE1. Neighbour-joiningtree basedonup to 538bp of the 16SRNA gene.Phrynobatrachus tanoeensissp.nov. (yellow) is sister to a clade of Phrynobatrachus species comprising P.intermedius (green) and P.liberiensis (blue); all three being sister to P.tokba (red).Phrynobatrachus liberiensis comprises two clades, one (showing little support) ranging from southern-central Ivory Coast (Banco National Park) west, through western Ivory Coast, south-eastern Guiana and Liberia to eastern Sierra Leone (Gola Rainforest National Park) (pale blue), and one comprising frogs from the Ankasa and Kakum National Parks, south-western Ghana (dark blue). Nodes supported by bootstrap NJ (above) and ML/PP (below); bootstrap value ¿ 65 and PP values&gt; 0.90 shown. Outgroups (P.cricogaster and P. natalensis) not shown." httpUri="https://zenodo.org/record/1187955/files/figure.png" pageId="8" pageNumber="229">Fig. 1</figureCitation>
). 
<taxonomicName id="4C2167BFFFFFF23EFEB2FF224EA64D91" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[329,647,152,176]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFEB2FF224EA64D91" box="[329,647,152,176]" italics="true" pageId="8" pageNumber="229">Phrynobatrachus liberiensis</emphasis>
</taxonomicName>
showed some clear intraspecific genetic structure, i.e. a western clade comprising frogs from the Ivorian Banco National Park to 
<collectingCountry id="F3365CACFFFFF23EFCD3FF064F994DF5" box="[808,952,188,213]" name="Sierra Leone" pageId="8" pageNumber="229">Sierra Leone</collectingCountry>
and a Ghanaian clade, comprising samples from Kakum and Ankasa National Parks. The intraspecific variation in 
<taxonomicName id="4C2167BFFFFFF23EFC4BFF5B48614DD9" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[944,1088,224,248]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFC4BFF5B48614DD9" box="[944,1088,224,248]" italics="true" pageId="8" pageNumber="229">P. liberiensis</emphasis>
</taxonomicName>
ranged from 0–4.42% (mean ± sd: 2.02 ± 1.35%, N= 351 comparisons). This high genetic variation may indicate further cryptic diversity, i.e. a further undescribed taxon from 
<collectingCountry id="F3365CACFFFFF23EFE01FE9D4E664C61" box="[506,583,295,320]" name="Ghana" pageId="8" pageNumber="229">Ghana</collectingCountry>
. The two included 
<taxonomicName id="4C2167BFFFFFF23EFCE7FE934F534C61" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[796,882,296,320]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tokba">
<emphasis id="B955C02EFFFFF23EFCE7FE934F534C61" box="[796,882,296,320]" italics="true" pageId="8" pageNumber="229">P. tokba</emphasis>
</taxonomicName>
sequences differed by 1.36%, while intraspecific variation in 
<taxonomicName id="4C2167BFFFFFF23EFEDEFEF74DE94C45" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[293,456,332,356]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFFFF23EFEDEFEF74DE94C45" box="[293,456,332,356]" italics="true" pageId="8" pageNumber="229">P. intermedius</emphasis>
</taxonomicName>
was zero. In 
<taxonomicName id="4C2167BFFFFFF23EFD90FEF74F214C45" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[619,768,333,356]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFFF23EFD90FEF74F214C45" box="[619,768,333,356]" italics="true" pageId="8" pageNumber="229">P. tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFFF23EFCF1FEF64F464C45" box="[778,871,332,357]" pageId="8" pageNumber="229" rank="species">
<emphasis id="B955C02EFFFFF23EFCF1FEF64F414C44" bold="true" box="[778,864,332,357]" pageId="8" pageNumber="229">sp. nov</emphasis>
.
</taxonomicNameLabel>
the intraspecific variation ranged from 0–0.19% (mean ± sd: 0.08 ± 0.10%, N= 10 comparisons).
</paragraph>
<paragraph id="8B9E1C3CFFFFF23EFF3CFE2E4F884F39" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="229">
<taxonomicName id="4C2167BFFFFFF23EFF3CFE2E4E264C8D" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[199,519,404,428]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFFF23EFF3CFE2E4E264C8D" box="[199,519,404,428]" italics="true" pageId="8" pageNumber="229">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFFF23EFDEBFE2E4E4B4C8C" box="[528,618,404,429]" pageId="8" pageNumber="229" rank="species">
<emphasis id="B955C02EFFFFF23EFDEBFE2E4E4B4C8C" bold="true" box="[528,618,404,429]" pageId="8" pageNumber="229">sp. nov.</emphasis>
</taxonomicNameLabel>
differed from 
<taxonomicName id="4C2167BFFFFFF23EFCEAFE2F4F904C8D" authorityName="Rodel, Boateng, Penner, &amp; Hillers" authorityYear="2009" box="[785,945,404,428]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B955C02EFFFFF23EFCEAFE2F4F904C8D" box="[785,945,404,428]" italics="true" pageId="8" pageNumber="229">P. intermedius</emphasis>
</taxonomicName>
by 6.39–6.70% (mean ± sd: 6.64 ± 0.13%, N= 10), and from 
<taxonomicName id="4C2167BFFFFFF23EFE92FE034DD64CF1" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[361,503,440,464]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFE92FE034DD64CF1" box="[361,503,440,464]" italics="true" pageId="8" pageNumber="229">P. liberiensis</emphasis>
</taxonomicName>
by 5.25–7.03% (mean ± sd: 5.95 ± 0.44%, N= 135), and from 
<taxonomicName id="4C2167BFFFFFF23EFB2FFE03490D4CF1" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[1236,1324,440,464]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tokba">
<emphasis id="B955C02EFFFFF23EFB2FFE03490D4CF1" box="[1236,1324,440,464]" italics="true" pageId="8" pageNumber="229">P. tokba</emphasis>
</taxonomicName>
by 6.60– 7.27% (mean ± sd: 6.97 ± 0.30%, N= 22). These differences are well within the range of what has been accepted as genetic distances between other species of the genus (see Discussion).
</paragraph>
<paragraph id="8B9E1C3CFFFFF23EFF3CFD9E48434E7C" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="229">
<emphasis id="B955C02EFFFFF23EFF3CFD9E4D0F4F1C" bold="true" box="[199,302,548,573]" pageId="8" pageNumber="229">Acoustic</emphasis>
. The recorded advertisement calls of two males of the new species comprise an amplitude modulated, short note, with few pulses that are difficult to tell apart (no full amplitude modulation = no complete silence between pulses; 
<figureCitation id="131A00B9FFFFF23EFEAEFDD64DE74FA4" box="[341,454,620,645]" captionStart="FIGURE 4" captionStartId="8.[151,250,1843,1865]" captionTargetBox="[180,1427,915,1796]" captionTargetId="figure@8.[151,1437,913,1822]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement calls of male Phrynobatrachus tanoeensis sp. nov. from Tanoé-Ehy forest, Dohouan village (A, ZMB 86954) and Nouamou (B, ZMB 86958), and P.liberiensis from Taï National Park (C; male not collected; bandpass filter between 1100‒4500 Hz) and Banco National Park (D, ZMB 86969; bandpass filter between 1100‒4500 Hz), all Ivory Coast (compare Table 3); spectrogram (above) and oscillogram (below)." httpUri="https://zenodo.org/record/1187961/files/figure.png" pageId="8" pageNumber="229">Fig. 4a, b</figureCitation>
). Call duration and frequency varied and showed some overlap to 
<taxonomicName id="4C2167BFFFFFF23EFB3DFDD749784FA5" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[1222,1369,620,644]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFB3DFDD749784FA5" box="[1222,1369,620,644]" italics="true" pageId="8" pageNumber="229">P. liberiensis</emphasis>
</taxonomicName>
calls, recorded in Banco and Taï National Parks, the latter calls as well showing some variation (
<figureCitation id="131A00B9FFFFF23EFB22FD3549024F89" box="[1241,1315,655,680]" captionStart="FIGURE 4" captionStartId="8.[151,250,1843,1865]" captionTargetBox="[180,1427,915,1796]" captionTargetId="figure@8.[151,1437,913,1822]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement calls of male Phrynobatrachus tanoeensis sp. nov. from Tanoé-Ehy forest, Dohouan village (A, ZMB 86954) and Nouamou (B, ZMB 86958), and P.liberiensis from Taï National Park (C; male not collected; bandpass filter between 1100‒4500 Hz) and Banco National Park (D, ZMB 86969; bandpass filter between 1100‒4500 Hz), all Ivory Coast (compare Table 3); spectrogram (above) and oscillogram (below)." httpUri="https://zenodo.org/record/1187961/files/figure.png" pageId="8" pageNumber="229">Fig. 4</figureCitation>
, 
<tableCitation id="C6A32987FFFFF23EFACFFD3549AF4F89" box="[1332,1422,655,680]" captionStart="TABLE 3" captionStartId="13.[151,239,290,313]" captionText="TABLE 3. Advertisement call values of Phrynobatrachus tanoeensis sp. nov. and P. liberiensis; given are the mean ± standard deviation, and range of calls per male (in brackets). Phrynobatrachus tanoeensis sp. nov. calls were both recorded on 3 October 2014, call 1 (ZMB 86954) in Dohouan (09h43 GMT); call 2 (ZMB86958) in Nouamou.The call of P. liberiensis from Banco National Park (BNP; ZMB 86969) was recorded on 6 August 2014 (15h03 GMT); the calls recorded in April1999 in Taï National Park (TNP) are from several males (minimum of 3) which have notbeen collected." pageId="8" pageNumber="229">Table 3</tableCitation>
). However, a comparison of the spectrogram and oscillogram between the calls of both taxa, shows distinct differences (
<figureCitation id="131A00B9FFFFF23EFEDFFD6D4D4D4FD1" box="[292,364,727,752]" captionStart="FIGURE 4" captionStartId="8.[151,250,1843,1865]" captionTargetBox="[180,1427,915,1796]" captionTargetId="figure@8.[151,1437,913,1822]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement calls of male Phrynobatrachus tanoeensis sp. nov. from Tanoé-Ehy forest, Dohouan village (A, ZMB 86954) and Nouamou (B, ZMB 86958), and P.liberiensis from Taï National Park (C; male not collected; bandpass filter between 1100‒4500 Hz) and Banco National Park (D, ZMB 86969; bandpass filter between 1100‒4500 Hz), all Ivory Coast (compare Table 3); spectrogram (above) and oscillogram (below)." httpUri="https://zenodo.org/record/1187961/files/figure.png" pageId="8" pageNumber="229">Fig. 4</figureCitation>
). The advertisement calls of 
<taxonomicName id="4C2167BFFFFFF23EFD47FD634F714FCE" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[700,848,728,752]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFFF23EFD47FD634F714FCE" box="[700,848,728,752]" italics="true" pageId="8" pageNumber="229">P. tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFFF23EFCA2FD6D4F944FCE" box="[857,949,727,752]" pageId="8" pageNumber="229" rank="species">
<emphasis id="B955C02EFFFFF23EFCA2FD6D4F8F4FD1" bold="true" box="[857,942,727,752]" pageId="8" pageNumber="229">sp. nov</emphasis>
.
</taxonomicNameLabel>
are much shorter and less pulsed than the calls of 
<emphasis id="B955C02EFFFFF23EFF0DFD474DB04E35" box="[246,401,764,788]" italics="true" pageId="8" pageNumber="229">
<taxonomicName id="4C2167BFFFFFF23EFF0DFD474DAC4E35" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[246,397,764,788]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">P. liberiensis</taxonomicName>
.
</emphasis>
These differences are also audible to the human ear. Whereas 
<taxonomicName id="4C2167BFFFFFF23EFB8FFD4749294E35" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[1140,1288,764,788]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFB8FFD4749294E35" box="[1140,1288,764,788]" italics="true" pageId="8" pageNumber="229">P. liberiensis</emphasis>
</taxonomicName>
calls can be described like a loud, hammer like “ackk,” the calls of the new species sound like a deep, short “tick.” Both species call predominantly during the day, in swamps or from the banks of small forest creeks.
</paragraph>
</subSubSection>
<caption id="DF5E4CB4FFFFF23EFF6CF8894F124A84" httpUri="https://zenodo.org/record/1187961/files/figure.png" pageId="8" pageNumber="229" startId="8.[151,250,1843,1865]" targetBox="[180,1427,915,1796]" targetPageId="8">
<paragraph id="8B9E1C3CFFFFF23EFF6CF8894F124A84" blockId="8.[151,1436,1843,1957]" pageId="8" pageNumber="229">
<emphasis id="B955C02EFFFFF23EFF6CF8894D344A69" bold="true" box="[151,277,1843,1865]" pageId="8" pageNumber="229">FIGURE 4.</emphasis>
Advertisement calls of male 
<taxonomicName id="4C2167BFFFFFF23EFDB3F8894F4D4A68" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[584,876,1843,1865]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFFF23EFDB3F8894F4D4A68" box="[584,876,1843,1865]" italics="true" pageId="8" pageNumber="229">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFFF23EFC8FF8894FE64A69" box="[884,967,1843,1864]" pageId="8" pageNumber="229" rank="species">
<emphasis id="B955C02EFFFFF23EFC8FF8894FE64A69" bold="true" box="[884,967,1843,1864]" pageId="8" pageNumber="229">sp. nov.</emphasis>
</taxonomicNameLabel>
from Tanoé-Ehy forest, Dohouan village (A, ZMB 86954) and Nouamou (B, ZMB 86958), and 
<taxonomicName id="4C2167BFFFFFF23EFD66F8E94F3E4A49" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[669,799,1874,1896]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="229" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFFF23EFD66F8E94F3E4A49" box="[669,799,1874,1896]" italics="true" pageId="8" pageNumber="229">P. liberiensis</emphasis>
</taxonomicName>
from Taï National Park (C; male not collected; bandpass filter between 1100‒4500 Hz) and Banco National Park (D, ZMB 86969; bandpass filter between 1100‒4500 Hz), all Ivory Coast (compare Table 3); spectrogram (above) and oscillogram (below).
</paragraph>
</caption>
<subSubSection id="C33B4FB7FFFEF23DFF3CFF2D48404AF5" lastPageId="11" lastPageNumber="232" pageId="9" pageNumber="230" type="distribution">
<paragraph id="8B9E1C3CFFFEF23FFF3CFF2D4D054DD6" blockId="9.[151,1437,151,464]" pageId="9" pageNumber="230">
<emphasis id="B955C02EFFFEF23FFF3CFF2D4D784D91" bold="true" box="[199,345,151,176]" pageId="9" pageNumber="230">Distribution</emphasis>
. The new species is so far only known from two swamp forests, the Tanoé-Ehy Swamp Forest region in south-eastern 
<collectingCountry id="F3365CACFFFEF23FFE66FF074E3E4DF5" box="[413,543,188,213]" name="Ivory Coast" pageId="9" pageNumber="230">Ivory Coast</collectingCountry>
and a forest near Ebonloa, south-western 
<collectingCountry id="F3365CACFFFEF23FFC09FF06481E4DF4" box="[1010,1087,188,213]" name="Ghana" pageId="9" pageNumber="230">Ghana</collectingCountry>
. These localities are only about 
<quantity id="4CD9B1D9FFFEF23FFF6CFF5A4CFF4DD9" box="[151,222,223,248]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="230" unit="km" value="40.0">40 km</quantity>
apart.
</paragraph>
<paragraph id="8B9E1C3CFFFEF23FFF3CFEBE4E4F4CF1" blockId="9.[151,1437,151,464]" pageId="9" pageNumber="230">
<emphasis id="B955C02EFFFEF23FFF3CFEBE4D5E4C3C" bold="true" box="[199,383,260,285]" pageId="9" pageNumber="230">Natural history</emphasis>
. Both known sites of the new species are predominantly swamp forests (
<figureCitation id="131A00B9FFFEF23FFB37FEBE49324C3C" box="[1228,1299,260,285]" captionStart="FIGURE5" captionStartId="9.[151,250,1940,1962]" captionTargetBox="[206,1358,517,1920]" captionTargetId="figure@9.[206,1358,517,1921]" captionTargetPageId="9" captionText="FIGURE5. Habitat of Phrynobatrachus tanoeensis sp. nov.; characterized by swampy area, and the presence of Raphia palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c)." httpUri="https://zenodo.org/record/1187963/files/figure.png" pageId="9" pageNumber="230">Fig. 5</figureCitation>
). The mean annual temperature of the Tanoé-Ehy forest is 26°C, the mean annual precipitation is about 
<quantity id="4CD9B1D9FFFEF23FFB5AFE9249314C1E" box="[1185,1296,296,320]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="230" unit="mm" value="2000.0">2000 mm</quantity>
. This region is characterized by a longer dry season (December to March), followed by the period with high precipitation (March to July). A minor rainy season extends from October to November (
<bibRefCitation id="EFB061CDFFFEF23FFC0FFED548F84CA9" box="[1012,1241,367,392]" pageId="9" pageNumber="230" refId="ref6747" refString="Avenard, J. - M., Eldin, M., Girard, G., Sircoulon, J., Touchebeuf, P., Guillaumet, J. - L. &amp; Adjanohoun, E. (1971) Le milieu naturel de la Cote d'Ivoire. ORSTOM, Paris, 391 pp." type="book">
Avenard 
<emphasis id="B955C02EFFFEF23FFBA7FECB48B74CA9" box="[1116,1174,368,392]" italics="true" pageId="9" pageNumber="230">et al.</emphasis>
1971
</bibRefCitation>
). The Tanoé-Ehy forest mainly consists of moist, partly primary forests on predominantly sandy soil, with vegetation typical for south-eastern 
<collectingCountry id="F3365CACFFFEF23FFEC8FE024D964CF1" box="[307,439,440,464]" name="Ivory Coast" pageId="9" pageNumber="230">Ivory Coast</collectingCountry>
(
<bibRefCitation id="EFB061CDFFFEF23FFE3EFE0D4E404CF1" box="[453,609,439,464]" pageId="9" pageNumber="230" refId="ref6805" refString="Beligne, V. (1994) Etude de l'etat du milieu naturel du Parc National du Banco: recommandation pour sa sauvegarde et son amenagement. Rapport du Fonds Mondial pour la Nature (WWF), Abidjan, 48 pp." type="book">Béligné 1994</bibRefCitation>
).
</paragraph>
<caption id="DF5E4CB4FFFEF23FFF6CF82E4E634AC9" httpUri="https://zenodo.org/record/1187963/files/figure.png" pageId="9" pageNumber="230" startId="9.[151,250,1940,1962]" targetBox="[206,1358,517,1920]" targetPageId="9">
<paragraph id="8B9E1C3CFFFEF23FFF6CF82E4E634AC9" blockId="9.[151,1436,1940,2024]" pageId="9" pageNumber="230">
<emphasis id="B955C02EFFFEF23FFF6CF82E4D334A8B" bold="true" box="[151,274,1940,1962]" pageId="9" pageNumber="230">FIGURE 5.</emphasis>
Habitat of 
<taxonomicName id="4C2167BFFFFEF23FFE7FF82E4E844A8B" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[388,677,1940,1962]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="230" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFEF23FFE7FF82E4E844A8B" box="[388,677,1940,1962]" italics="true" pageId="9" pageNumber="230">Phrynobatrachus tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFEF23FFD50F82F4EDA4A8B" box="[683,763,1941,1962]" pageId="9" pageNumber="230" rank="species">
<emphasis id="B955C02EFFFEF23FFD50F82F4EDA4A8B" bold="true" box="[683,763,1941,1962]" pageId="9" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
; characterized by swampy area, and the presence of 
<taxonomicName id="4C2167BFFFFEF23FFAEBF82E49784A8B" box="[1296,1369,1940,1962]" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B955C02EFFFEF23FFAEBF82E49784A8B" box="[1296,1369,1940,1962]" italics="true" pageId="9" pageNumber="230">Raphia</emphasis>
</taxonomicName>
palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c).
</paragraph>
</caption>
<caption id="DF5E4CB4FFFDF23CFF6CF8E44CDB4AF1" httpUri="https://zenodo.org/record/1187965/files/figure.png" pageId="10" pageNumber="231" startId="10.[151,250,1886,1908]" targetBox="[212,1375,193,1865]" targetPageId="10">
<paragraph id="8B9E1C3CFFFDF23CFF6CF8E44CDB4AF1" blockId="10.[151,1436,1886,2000]" pageId="10" pageNumber="231">
<emphasis id="B955C02EFFFDF23CFF6CF8E44D344A55" bold="true" box="[151,277,1886,1908]" pageId="10" pageNumber="231">FIGURE 6.</emphasis>
Dorsal and ventral views of adult 
<taxonomicName id="4C2167BFFFFDF23CFD7FF8E44F864A55" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[644,935,1886,1908]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="231" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFDF23CFD7FF8E44F864A55" box="[644,935,1886,1908]" italics="true" pageId="10" pageNumber="231">Phrynobatrachus liberiensis</emphasis>
</taxonomicName>
males from Banco National Park, south-eastern Ivory Coast (a, b &amp; g); Taï National Park, western Ivory Coast (d, e &amp; f), and Gola Rainforest National Park, eastern Sierra Leone (c); note differences in colour, skin structure and shape of scapular ridges. Specimens not collected or not assignable to vouchers.
</paragraph>
</caption>
<caption id="DF5E4CB4FFFCF23DFF6CFA0E4D8B4B07" httpUri="https://zenodo.org/record/1187967/files/figure.png" pageId="11" pageNumber="232" startId="11.[151,250,1460,1482]" targetBox="[188,1398,213,1423]" targetPageId="11">
<paragraph id="8B9E1C3CFFFCF23DFF6CFA0E4D8B4B07" blockId="11.[151,1436,1460,1574]" pageId="11" pageNumber="232">
<emphasis id="B955C02EFFFCF23DFF6CFA0E4D3548E8" bold="true" box="[151,276,1460,1482]" pageId="11" pageNumber="232">FIGURE 7.</emphasis>
Dorsal and ventral views of adult 
<taxonomicName id="4C2167BFFFFCF23DFD81FA0E4FBA48EB" authorityName="Barbour &amp; Loveridge" authorityYear="1927" box="[634,923,1460,1482]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="liberiensis">
<emphasis id="B955C02EFFFCF23DFD81FA0E4FBA48EB" box="[634,923,1460,1482]" italics="true" pageId="11" pageNumber="232">Phrynobatrachus liberiensis</emphasis>
</taxonomicName>
females from Banco National Park, south-eastern Ivory Coast (a, b, e‒g; f, g: ZMB 86981); Taï National Park, western Ivory Coast (d), and Yekeba, Liberia (c; photo: J. Penner); note differences in colour and shape of scapular ridges. If no accession number provided, the specimens were not collected or not assignable to vouchers.
</paragraph>
</caption>
<paragraph id="8B9E1C3CFFFCF23DFF3CF9E948F54AAD" blockId="11.[151,1436,1619,2004]" pageId="11" pageNumber="232">
Whereas all collected males from the Tanoé-Ehy forest were encountered well concealed in areas thickly covered with 
<taxonomicName id="4C2167BFFFFCF23DFEC9F9CD4DA24BAE" box="[306,387,1655,1679]" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="B955C02EFFFCF23DFEC9F9CD4DA24BAE" box="[306,387,1655,1679]" italics="true" pageId="11" pageNumber="232">Raphia</emphasis>
</taxonomicName>
palms (
<figureCitation id="131A00B9FFFCF23DFE25F9CC4E134BAE" box="[478,562,1654,1679]" captionStart="FIGURE5" captionStartId="9.[151,250,1940,1962]" captionTargetBox="[206,1358,517,1920]" captionTargetId="figure@9.[206,1358,517,1921]" captionTargetPageId="9" captionText="FIGURE5. Habitat of Phrynobatrachus tanoeensis sp. nov.; characterized by swampy area, and the presence of Raphia palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c)." httpUri="https://zenodo.org/record/1187963/files/figure.png" pageId="11" pageNumber="232">Fig. 5b</figureCitation>
), the females were seen on muddy ground in open patches of the forest. The canopy of the sites where 
<taxonomicName id="4C2167BFFFFCF23DFE46F9264E714B92" authority="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel, 2018" authorityName="Kpan &amp; Kouamé &amp; Barej &amp; Adeba &amp; Emmrich &amp; Ofori-Boateng &amp; Rödel" authorityYear="2018" box="[445,592,1692,1715]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="232" phylum="Chordata" rank="species" species="tanoeensis" status="sp. nov.">
<emphasis id="B955C02EFFFCF23DFE46F9264E714B92" box="[445,592,1692,1715]" italics="true" pageId="11" pageNumber="232">P. tanoeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2667D55FFFCF23DFDACF9214E934B95" box="[599,690,1691,1716]" pageId="11" pageNumber="232" rank="species">
<emphasis id="B955C02EFFFCF23DFDACF9214E934B95" bold="true" box="[599,690,1691,1716]" pageId="11" pageNumber="232">sp. nov.</emphasis>
</taxonomicNameLabel>
was found had gaps (
<figureCitation id="131A00B9FFFCF23DFC52F9214FDB4B95" box="[937,1018,1691,1716]" captionStart="FIGURE5" captionStartId="9.[151,250,1940,1962]" captionTargetBox="[206,1358,517,1920]" captionTargetId="figure@9.[206,1358,517,1921]" captionTargetPageId="9" captionText="FIGURE5. Habitat of Phrynobatrachus tanoeensis sp. nov.; characterized by swampy area, and the presence of Raphia palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c)." httpUri="https://zenodo.org/record/1187963/files/figure.png" pageId="11" pageNumber="232">Fig. 5a</figureCitation>
), and traces of recent human activity were visible. For instance, 
<taxonomicName id="4C2167BFFFFCF23DFE23F9054E084BF6" box="[472,553,1727,1751]" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="B955C02EFFFCF23DFE23F9054E084BF6" box="[472,553,1727,1751]" italics="true" pageId="11" pageNumber="232">Raphia</emphasis>
</taxonomicName>
palms were used by the local population for palm wine extraction and as construction material. An unpaved road, leading to Dohouan and Nouamou villages, crossed the area. Due to frequent traffic with motorbikes the Dohouan site was quite noisy. The road was only eight meters from the calling site of one male. This swamp forest was surrounded by heavily degraded forest and plantations for subsistence farming (i.e. cassava plantation associated with coconut trees). The Dohouan River, which crosses the Tanoé-Ehy forest, is polluted due to the activity of launderers and people washing their cars and motorbikes.
</paragraph>
<paragraph id="8B9E1C3CFFFCF23DFF3CF82248404AF5" blockId="11.[151,1436,1619,2004]" pageId="11" pageNumber="232">
In 
<collectingCountry id="F3365CACFFFCF23DFF12F82C4D144A8E" box="[233,309,1942,1967]" name="Ghana" pageId="11" pageNumber="232">Ghana</collectingCountry>
, the species was collected in small patches of land within an inundated landscape (
<figureCitation id="131A00B9FFFCF23DFAFFF82C49794A8E" box="[1284,1368,1942,1967]" captionStart="FIGURE5" captionStartId="9.[151,250,1940,1962]" captionTargetBox="[206,1358,517,1920]" captionTargetId="figure@9.[206,1358,517,1921]" captionTargetPageId="9" captionText="FIGURE5. Habitat of Phrynobatrachus tanoeensis sp. nov.; characterized by swampy area, and the presence of Raphia palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c)." httpUri="https://zenodo.org/record/1187963/files/figure.png" pageId="11" pageNumber="232">Fig. 5c</figureCitation>
). The dominant vegetation on these ‘islands’ are trees with stilt-like roots and 
<taxonomicName id="4C2167BFFFFCF23DFC44F80148314AF2" box="[959,1040,1979,2003]" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="B955C02EFFFCF23DFC44F80148314AF2" box="[959,1040,1979,2003]" italics="true" pageId="11" pageNumber="232">Raphia</emphasis>
</taxonomicName>
palms.
</paragraph>
</subSubSection>
<subSubSection id="C33B4FB7FFFBF23AFF3CFF2D4DD24DF5" pageId="12" pageNumber="233" type="etymology">
<paragraph id="8B9E1C3CFFFBF23AFF3CFF2D4DD24DF5" blockId="12.[151,1436,151,213]" pageId="12" pageNumber="233">
<emphasis id="B955C02EFFFBF23AFF3CFF2D4D6F4D91" bold="true" box="[199,334,151,176]" pageId="12" pageNumber="233">Etymology.</emphasis>
The specific name refers to the 
<typeStatus id="549AA29EFFFBF23AFD32FF224ED84D91" box="[713,761,152,176]" pageId="12" pageNumber="233">type</typeStatus>
locality, the Tanoé-Ehy Swamp Forest. We suggest Tanoé Puddle Frog as common name.
</paragraph>
</subSubSection>
</treatment>
</document>