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<document id="4EB52AD09FE77FD3A81225618A105425" ID-DOI="10.5281/zenodo.273398" ID-GBIF-Dataset="ae452b89-2798-4a2a-8bd9-963dead78ebc" ID-ISSN="1175­5326" ID-Zenodo-Dep="273398" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761708870" checkinUser="plazi" docAuthor="Howden, Henry F." docDate="2006" docId="0380EE1A1951FFE5FE94FEC03F860659" docLanguage="en" docName="zt01137p052.pdf" docOrigin="Zootaxa 1137" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Neoathyreus fallolobus Howden, 2006, new species" docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="FFB996621959FFEEFF9CFB0C3B2E025F" masterDocTitle="New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)" masterLastPageNumber="52" masterPageNumber="37" pageNumber="45" updateTime="1698215286212" updateUser="plazi">
<mods:mods id="165EECC7443FC68BF95D8DF005BDBA96" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2896EBFD6D6EB427E9F12415B8BA05C">
<mods:title id="8A4DBC35A3957FE02B8FDB2ABC9D32DA">New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)</mods:title>
</mods:titleInfo>
<mods:name id="2BBDD0F357314FC2AFACFA0A4F64D29B" type="personal">
<mods:role id="A4A075BEB2B4B7699796AF1DBCF8CC88">
<mods:roleTerm id="C1B689B6C0DBE9B8B5EE29907D4EAD2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1D8E3A4AB281B4A3CBF0FCFB4D5CEBA">Howden, Henry F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9B914ECC5FDA3ED555A1BAB87D4258F4">text</mods:typeOfResource>
<mods:relatedItem id="ED10471F752CB7B2D41AF2A8228C080D" type="host">
<mods:titleInfo id="CF7E00D4198D29F0D2E82998347AB632">
<mods:title id="D027F7BF6BB65990AA40A1B04C30D156">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="82F6BAF748CCDCD0B233AC160C9B9D63">
<mods:date id="BB3BE99A79F53FA6B1AA2FD133B860F9">2006</mods:date>
<mods:detail id="78BB5DCD2E0818CCAFC9F50E8A0D614D" type="volume">
<mods:number id="A0755336D7487B011A9BC5201CD3DFE3">1137</mods:number>
</mods:detail>
<mods:extent id="69B1EE94D8521F5CDB574BFE76982327" unit="page">
<mods:start id="212DEC3FE6A19FDAFDA2CC297979F125">37</mods:start>
<mods:end id="858F63A1AC601B84049D22301521C9EA">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="89637DFEFE64A7D2E5AB493E52299D3C">journal article</mods:classification>
<mods:identifier id="B85AB719EE820529E62D8E6D13A7ADC8" type="DOI">10.5281/zenodo.273398</mods:identifier>
<mods:identifier id="D686BE8F1DB1269A46ADE4C97EDDD1F0" type="GBIF-Dataset">ae452b89-2798-4a2a-8bd9-963dead78ebc</mods:identifier>
<mods:identifier id="91C158EEF4893F47BB395834307448C2" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="4695AFE6D42E31FE2DDD71A645B1E7E3" type="Zenodo-Dep">273398</mods:identifier>
</mods:mods>
<treatment id="0380EE1A1951FFE5FE94FEC03F860659" ID-DOI="http://doi.org/10.5281/zenodo.6260970" ID-GBIF-Taxon="119345042" ID-Zenodo-Dep="6260970" LSID="urn:lsid:plazi:treatment:0380EE1A1951FFE5FE94FEC03F860659" httpUri="http://treatment.plazi.org/id/0380EE1A1951FFE5FE94FEC03F860659" lastPageId="11" lastPageNumber="48" pageId="8" pageNumber="45">
<subSubSection id="C3330C871951FFE6FE94FEC0384007B9" box="[264,878,1484,1510]" pageId="8" pageNumber="45" type="nomenclature">
<paragraph id="8B965F0C1951FFE6FE94FEC0384007B9" blockId="8.[264,878,1484,1510]" box="[264,878,1484,1510]" pageId="8" pageNumber="45">
<heading id="D0DEE8601951FFE6FE94FEC0384007B9" bold="true" box="[264,878,1484,1510]" fontSize="11" level="1" pageId="8" pageNumber="45" reason="1">
<emphasis id="B95D831E1951FFE6FE94FEC0384007B9" bold="true" box="[264,878,1484,1510]" pageId="8" pageNumber="45">
<emphasis id="B95D831E1951FFE6FE94FEC0390707B9" bold="true" box="[264,553,1484,1510]" italics="true" pageId="8" pageNumber="45">
<taxonomicName id="4C29248F1951FFE6FE94FEC0390D07B9" box="[264,547,1484,1510]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="fallolobus" status="sp. nov.">Neoathyreus fallolobus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26E3E651951FFE6FDACFEC0399107B9" box="[560,703,1484,1510]" pageId="8" pageNumber="45" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="131243891951FFE6FD53FEC0387307B9" box="[719,861,1484,1510]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="8" pageNumber="45">Figs. 12–15</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3330C871951FFE7FE94FD1A3A9A04D4" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45" type="materials_examined">
<paragraph id="8B965F0C1951FFE7FE94FD1A3A5D0671" blockId="8.[264,1323,1558,1905]" lastBlockId="9.[264,1325,284,1875]" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45">
<emphasis id="B95D831E1951FFE6FE94FD1A3AAF046F" bold="true" box="[264,385,1558,1584]" pageId="8" pageNumber="45">
<typeStatus id="5492E1AE1951FFE6FE94FD1A3A52046F" box="[264,380,1558,1584]" pageId="8" pageNumber="45" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, length
<quantity id="4CD1F2E91951FFE6FDACFD1B398E046E" box="[560,672,1559,1585]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="8" pageNumber="45" unit="mm" value="14.9">14.9 mm</quantity>
, greatest width
<quantity id="4CD1F2E91951FFE6FCF8FD1B38EB046E" box="[868,965,1559,1585]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="8" pageNumber="45" unit="mm" value="8.7">8.7 mm</quantity>
. Dorsally black, surface with scattered, erect, pale setae. Clypeus (
<figureCitation id="131243891951FFE6FD45FD33381B0406" box="[729,821,1599,1625]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="8" pageNumber="45">Fig. 12</figureCitation>
) with longitudinal, median horn arising anteriorly and extending length of clypeus; lateral horn or tubercle on each side above antennal insertion almost same height as median horn; anterior carina on each side curved abruptly upward near median horn and extending up one­half anterior face of horn; posterior carina on each side arcuately depressed between horns; surface of clypeus on each side shallowly concave, contiguously punctate­rugose. Vertex punctate­rugose anteriorly near horns, closely granulate elsewhere, surface slightly convex between eyes. Gena with slightly reflexed outer margin, anterior lateral angle with small tubercle, surface of gena tuberculate. Pronotum (
<figureCitation id="131243891950FFE7FDE0FA1039FF0369" box="[636,721,284,310]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="9" pageNumber="46">Fig. 12</figureCitation>
) with small, shallow fossa on each side behind eye adjacent to margin, fossa slightly wider than width of marginal bead; anterior marginal bead gradually, arcuately raised between fossae, midline constricted by very small indentation, lacking tubercle or angulation. Pronotum with inner carina on each side arising about 1.0 mm posterior to fossa; carina gradually bowed inwardly and becoming wider to rounded end about
<quantity id="4CD1F2E91950FFE7FDCFFAE8398003A1" box="[595,686,484,510]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="46" unit="mm" value="0.3">0.3 mm</quantity>
from posterior midline; outer edge of carina elevated; between inner posterior edges of carina with very small tubercle on midline. Concavity (
<figureCitation id="131243891950FFE7FE8DF9383A420011" box="[273,364,564,590]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="9" pageNumber="46">Fig. 12</figureCitation>
) between inner carinae deep in anterior half, abrupt, posterior edge of deep depression broadly v­shaped, extending in posterior half as shallowly impressed midline, posterior half of concavity shallow, sloping evenly to midline. Outer carina on each side 1.0 mm long, very narrow, about
<quantity id="4CD1F2E91950FFE7FD3FF9A0382E0099" box="[675,768,684,710]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="46" unit="mm" value="0.1">0.1 mm</quantity>
wide or less, slightly sinuous, inconspicuous. Lateral pronotal marginal bead complete, not broken or obviously indented; entire surface of pronotum except for carinae closely, rather coarsely granulate. Elytron with several striae slightly indicated at base, surface closely punctate­rugose except for smooth surface of humeral umbone. Metasternum between middle coxae slightly convex, midline shallowly impressed; anterior end of metasternal midline sharply, almost acutely angled, angle becoming broadly obtuse; surface on either side of midline punctate, most punctures separated by about two diameters; some punctures each with long seta. Foretibia with five teeth on outer margin. Male genitalia as in
<figureCitation id="131243891950FFE7FC95F8E038B70659" box="[777,921,1004,1030]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="9" pageNumber="46">Figs. 14, 15</figureCitation>
, similar to those of male
<taxonomicName id="4C29248F1950FFE7FB54F8E03A5D0671" authority="Howden." authorityName="Howden." class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="lobus">
<emphasis id="B95D831E1950FFE7FB54F8E03E04065A" box="[1224,1322,1004,1029]" italics="true" pageId="9" pageNumber="46">N. lobus</emphasis>
Howden.
</taxonomicName>
</paragraph>
<paragraph id="8B965F0C1950FFE7FEA4FF3039EF07C9" blockId="9.[264,1325,284,1875]" pageId="9" pageNumber="46">
<emphasis id="B95D831E1950FFE7FEA4FF303A860609" bold="true" box="[312,424,1084,1110]" pageId="9" pageNumber="46">Allotype.</emphasis>
Female, length
<quantity id="4CD1F2E91950FFE7FDF6FF3039F90609" box="[618,727,1084,1110]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="9" pageNumber="46" unit="mm" value="13.6">13.6 mm</quantity>
, greatest width
<quantity id="4CD1F2E91950FFE7FC0EFF3038DC0609" box="[914,1010,1084,1110]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="46" unit="mm" value="8.1">8.1 mm</quantity>
. Similar to male except as follows: median clypeal horn (
<figureCitation id="131243891950FFE7FD15FF6839CA0621" box="[649,740,1124,1150]" captionStart="FIGURES 12 – 15" captionStartId="10.[264,383,1378,1402]" captionTargetBox="[294,1293,566,1354]" captionTargetId="figure@10.[294,1293,566,1354]" captionTargetPageId="10" captionText="FIGURES 12 – 15. Neoathyreus fallolobus, n. sp.: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="9" pageNumber="46">Fig. 13</figureCitation>
) reduced to large tubercle at anterior edge of clypeus, extending posteriorly as low ridge along midline to join slightly developed posterior carina; carina bowed downward between lateral horn or large tubercle on each side, horn 2–3 times height of median tubercle; posterior carina near midline only slightly developed. Pronotum with tubercle between posterior ends of inner carina reduced to low, rounded swelling no higher than inner edge of adjacent carina; outer carina almost obsolete, represented by narrow row of 4–5 fused tubercles. Pygidium more broadly rounded, briefly emarginate medially.
</paragraph>
<paragraph id="8B965F0C1950FFE7FEA4FEA93A9A04D4" blockId="9.[264,1325,284,1875]" pageId="9" pageNumber="46">
<emphasis id="B95D831E1950FFE7FEA4FEA93AFF07E0" bold="true" box="[312,465,1445,1471]" pageId="9" pageNumber="46">
<typeStatus id="5492E1AE1950FFE7FEA4FEA93A5B07E0" box="[312,373,1445,1471]" pageId="9" pageNumber="46">Type</typeStatus>
Series.
</emphasis>
<typeStatus id="5492E1AE1950FFE7FE47FEAA3965079F" box="[475,587,1446,1472]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
, male,
<collectingCountry id="F33E1F9C1950FFE7FD38FEAA39F3079F" box="[676,733,1446,1472]" name="Peru" pageId="9" pageNumber="46">Peru</collectingCountry>
, Cusco:
<collectingCountry id="F33E1F9C1950FFE7FCDBFEAA38EB079F" box="[839,965,1446,1472]" name="Saint Pierre and Miquelon" pageId="9" pageNumber="46">San Pedro</collectingCountry>
, 13 0 3'15&quot;
<geoCoordinate id="EE1D39CB1950FFE7FBC0FEAA3FE3079F" box="[1116,1229,1444,1472]" direction="south" orientation="latitude" pageId="9" pageNumber="46" precision="925" value="-71.53333">S 71 32'</geoCoordinate>
54&quot; W,
<quantity id="4CD1F2E91950FFE7FE94FEC33A4D07B6" box="[264,355,1487,1513]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="46" unit="m" value="1500.0">1500 m</quantity>
, 17,18.
<date id="FF9779CC1950FFE7FE25FEC3393207B6" box="[441,540,1487,1513]" pageId="9" pageNumber="46" value="1999-11">XI.1999</date>
, T. Larsen, Cloud forest, F.I.T. 1 (
<collectionCode id="ED38C7C91950FFE7FC34FEC33F2B07B6" box="[936,1029,1487,1513]" pageId="9" pageNumber="46">CMNC</collectionCode>
). Allotype, female,
<collectingCountry id="F33E1F9C1950FFE7FB71FEC33E0807B6" box="[1261,1318,1487,1513]" name="Peru" pageId="9" pageNumber="46">Peru</collectingCountry>
, Cusco: Iskaybamba coffee plantation,
<quantity id="4CD1F2E91950FFE7FD6CFEF43878044D" box="[752,854,1528,1554]" metricMagnitude="3" metricUnit="m" metricValue="1.835" pageId="9" pageNumber="46" unit="m" value="1835.0">1835 m</quantity>
, 13 30' 16.3&quot;
<geoCoordinate id="EE1D39CB1950FFE7FBB6FEF43F87044D" box="[1066,1193,1526,1554]" direction="south" orientation="latitude" pageId="9" pageNumber="46" precision="925" value="-70.88333">S 70 53'</geoCoordinate>
58.7&quot; W, 29,30.
<date id="FF9779CC1950FFE7FECDFD2D3A860464" box="[337,424,1569,1595]" pageId="9" pageNumber="46" value="2000-05">V.2000</date>
, T.Larsen, F.I.T. (
<collectionCode id="ED38C7C91950FFE7FD1AFD2D39CE0464" box="[646,736,1569,1595]" pageId="9" pageNumber="46">HAHC</collectionCode>
).
<typeStatus id="5492E1AE1950FFE7FD65FD2D38400464" box="[761,878,1569,1595]" pageId="9" pageNumber="46" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D2F94851950FFE7FC1CFD2D38CA0464" box="[896,996,1569,1595]" pageId="9" pageNumber="46" type="male">3 males</specimenCount>
: 1, same data as
<typeStatus id="5492E1AE1950FFE7FB58FD2D3E050464" box="[1220,1323,1569,1595]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
except 19,20.
<date id="FF9779CC1950FFE7FE3BFD453925043C" box="[423,523,1609,1635]" pageId="9" pageNumber="46" value="1999-11">XI.1999</date>
; 2, same data as
<typeStatus id="5492E1AE1950FFE7FD4AFD453813043C" box="[726,829,1609,1635]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
except 18,19.
<date id="FF9779CC1950FFE7FC7FFD453F69043C" box="[995,1095,1609,1635]" pageId="9" pageNumber="46" value="1999-11">XI.1999</date>
, F.I.T. 2.
<typeStatus id="5492E1AE1950FFE7FB25FD453E05043C" box="[1209,1323,1609,1635]" pageId="9" pageNumber="46" type="paratype">Paratypes</typeStatus>
are in:
<collectionCode id="ED38C7C91950FFE7FEC5FD7D3A9E04D4" box="[345,432,1649,1675]" pageId="9" pageNumber="46">HAHC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3330C871950FFE5FEA4FD953FB301D1" lastPageId="11" lastPageNumber="48" pageId="9" pageNumber="46" type="discussion">
<paragraph id="8B965F0C1950FFE4FEA4FD953F1703A1" blockId="9.[264,1325,284,1875]" lastBlockId="10.[264,1323,284,510]" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<emphasis id="B95D831E1950FFE7FEA4FD953A9F04EC" bold="true" box="[312,433,1689,1715]" pageId="9" pageNumber="46">Remarks.</emphasis>
Variation in the length of the three male
<typeStatus id="5492E1AE1950FFE7FC0DFD953F2F04EC" box="[913,1025,1689,1715]" pageId="9" pageNumber="46" type="paratype">paratypes</typeStatus>
ranges from 10.0 to
<quantity id="4CD1F2E91950FFE7FB69FD953A180484" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="46" unit="mm" value="12.5">12.5 mm</quantity>
and greatest width from
<quantity id="4CD1F2E91950FFE7FDF6FDCD38340484" box="[618,794,1729,1755]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.6" metricValueMin="6.5" pageId="9" pageNumber="46" unit="mm" value="7.05" valueMax="7.6" valueMin="6.5">6.5 to 7.6 mm</quantity>
. The smallest male is dark brown (possibly teneral) and has the clypeus shaped like the allotype; the lateral horns are reduced to tubercles, smaller than in the allotype. In two of the three males, both small, the anterior pronotal depression in the concavity, while still evident, is reduced in depth and the inner carinae are only slightly wider posteriorly. In the largest male
<typeStatus id="5492E1AE1953FFE4FB83FA103FAA0369" box="[1055,1156,284,310]" pageId="10" pageNumber="47" type="paratype">paratype</typeStatus>
the posterior pronotal tubercle between the ends of the inner carinae is only represented by a smooth granule; in the smaller males the tubercle is absent. The lateral carinae are absent in one male and only slightly indicated in the other two. Dorsal setae vary in density, perhaps because of immersion in the F I T fluid or because of abrasion. There are also slight differences in the positioning in the apical lobes of the male genitalia.
</paragraph>
<caption id="DF560F841953FFE4FE94FE6E399107E0" httpUri="https://zenodo.org/record/273401/files/figure.png" pageId="10" pageNumber="47" targetBox="[294,1293,566,1354]" targetPageId="10">
<paragraph id="8B965F0C1953FFE4FE94FE6E399107E0" blockId="10.[264,1323,1378,1471]" pageId="10" pageNumber="47">
<emphasis id="B95D831E1953FFE4FE94FE6E3AFF0725" bold="true" box="[264,465,1378,1402]" pageId="10" pageNumber="47">FIGURES 12–15.</emphasis>
<emphasis id="B95D831E1953FFE4FE46FE6F39F40725" box="[474,730,1379,1402]" italics="true" pageId="10" pageNumber="47">
<taxonomicName id="4C29248F1953FFE4FE46FE6F39F90725" box="[474,727,1379,1402]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="fallolobus" status="sp. nov.">Neoathyreus fallolobus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A26E3E651953FFE4FD7FFE6F380C0725" box="[739,802,1379,1402]" pageId="10" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
: 12, Male head and pronotum, dorsal view; 13, Female head and pronotum, lateral view; 14, Male genitalia, dorsal view; 15, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B965F0C1953FFE5FEA4FD163E2103D9" blockId="10.[264,1324,1562,1868]" lastBlockId="11.[264,1324,284,1030]" lastPageId="11" lastPageNumber="48" pageId="10" pageNumber="47">
Large males and possibly some large females will key, with some difficulty because of their uniform black color, to couplet
<quantity id="4CD1F2E91953FFE4FD4FFD4E383D0403" box="[723,787,1602,1628]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="10" pageNumber="47" unit="in" value="16.0">16 in</quantity>
my 1985 revision; this couplet includes
<taxonomicName id="4C29248F1953FFE4FA8CFD4F393A04DB" authority="Felsche" authorityName="Felsche" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="corniculatus">
<emphasis id="B95D831E1953FFE4FA8CFD4F3AB404DB" italics="true" pageId="10" pageNumber="47">N. corniculatus</emphasis>
(Felsche)
</taxonomicName>
from
<collectingCountry id="F33E1F9C1953FFE4FDF9FD6639F304DB" box="[613,733,1642,1668]" name="Venezuela" pageId="10" pageNumber="47">Venezuela</collectingCountry>
and
<taxonomicName id="4C29248F1953FFE4FCBCFD6738D804DB" authority="Howden" authorityName="Howden" box="[800,1014,1642,1668]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="lobus">
<emphasis id="B95D831E1953FFE4FCBCFD6738A904DB" box="[800,903,1643,1668]" italics="true" pageId="10" pageNumber="47">N. lobus</emphasis>
Howden
</taxonomicName>
from
<collectingCountry id="F33E1F9C1953FFE4FBDBFD663FEE04DB" box="[1095,1216,1642,1668]" name="Argentina" pageId="10" pageNumber="47">Argentina</collectingCountry>
. Smaller specimens that lack the posterior pronotal tubercle will not key beyond couplet 13, and specimens that also lack the outer pronotal carinae will not key beyond couplet 10. However,
<taxonomicName id="4C29248F1953FFE4FE1BFDEF390A04A3" box="[391,548,1763,1788]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="fallolobus">
<emphasis id="B95D831E1953FFE4FE1BFDEF390A04A3" box="[391,548,1763,1788]" italics="true" pageId="10" pageNumber="47">N. fallolobus</emphasis>
</taxonomicName>
can be identified by the following combination of characters: uniform dorsal color, unusual clypeus with longitudinal median ridge, lack of an anterior pronotal tubercle and with small or no posterior tubercle, deep depression in the anterior half of pronotal concavity, surface of pronotum, except carinae, closely granulate or punctate, lateral pronotal marginal bead unbroken, and with five teeth on outer edge of foretibia. No other described species of
<taxonomicName id="4C29248F1952FFE5FD40FA60384103DA" box="[732,879,364,389]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B95D831E1952FFE5FD40FA60384103DA" box="[732,879,364,389]" italics="true" pageId="11" pageNumber="48">Neoathyreus</emphasis>
</taxonomicName>
has this combination of characters.
</paragraph>
<paragraph id="8B965F0C1952FFE5FEA4FA983FB301D1" blockId="11.[264,1324,284,1030]" pageId="11" pageNumber="48">
The “
<taxonomicName id="4C29248F1952FFE5FEE6FA983A9003F1" box="[378,446,404,430]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="lobus">lobus</taxonomicName>
” complex of forms in southern South
<collectingCountry id="F33E1F9C1952FFE5FC13FA9838DA03F1" box="[911,1012,404,430]" name="United States of America" pageId="11" pageNumber="48">America</collectingCountry>
presents some interesting problems, possibly related to geographic variation. The
<typeStatus id="5492E1AE1952FFE5FC2FFAB038C80389" box="[947,998,444,470]" pageId="11" pageNumber="48">type</typeStatus>
locality of
<taxonomicName id="4C29248F1952FFE5FBEBFAB03FF3038A" box="[1143,1245,444,469]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="lobus">
<emphasis id="B95D831E1952FFE5FBEBFAB03FF3038A" box="[1143,1245,444,469]" italics="true" pageId="11" pageNumber="48">N. lobus</emphasis>
</taxonomicName>
is
<collectingCountry id="F33E1F9C1952FFE5FA9BFAB03A5303A1" name="Argentina" pageId="11" pageNumber="48">the Argentine</collectingCountry>
province of Catamarca; other typical specimens have been taken in Tucumán. All of these specimens have the anterior pronotal marginal bead medially thickened and with a median, posteriorly directed, obtuse angulation. In specimens from northern Salta and Jujuy,
<collectingCountry id="F33E1F9C1952FFE5FE17F950392D0029" box="[395,515,604,630]" name="Argentina" pageId="11" pageNumber="48">Argentina</collectingCountry>
, the anterior pronotal bead is only slightly thickened and lacks the obtuse angulation. All
<collectingCountry id="F33E1F9C1952FFE5FDBFF98839B600C1" box="[547,664,644,670]" name="Argentina" pageId="11" pageNumber="48">Argentine</collectingCountry>
specimens dorsally are either bicolored light to dark brown or entirely brown to dark brown. In Cochabamba,
<collectingCountry id="F33E1F9C1952FFE5FC2CF9A03F270099" box="[944,1033,684,710]" name="Bolivia" pageId="11" pageNumber="48">Bolivia</collectingCountry>
, on the eastern slopes of the Andes, specimens are dorsally uniformly black, the anterior pronotal bead is uniformly thin, and the posterior pronotal tubercle is pronounced but slightly more slender than in
<collectingCountry id="F33E1F9C1952FFE5FE94F8283A530161" box="[264,381,804,830]" name="Argentina" pageId="11" pageNumber="48">Argentine</collectingCountry>
specimens. All of these “forms” have similar genitalia, as does
<emphasis id="B95D831E1952FFE5FB15F8283E050162" box="[1161,1323,804,829]" italics="true" pageId="11" pageNumber="48">
<taxonomicName id="4C29248F1952FFE5FB15F8283E090162" box="[1161,1319,804,829]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="fallolobus">N. fallolobus</taxonomicName>
.
</emphasis>
With the material at hand, it is obvious that more material from different localities is needed to determine if the variation is clinal or if several species are involved.
</paragraph>
</subSubSection>
<subSubSection id="C3330C871952FFE5FEA4F8903F860659" pageId="11" pageNumber="48" type="etymology">
<paragraph id="8B965F0C1952FFE5FEA4F8903F860659" blockId="11.[264,1324,284,1030]" pageId="11" pageNumber="48">
<emphasis id="B95D831E1952FFE5FEA4F8903AEA01E9" bold="true" box="[312,452,924,950]" pageId="11" pageNumber="48">Etymology.</emphasis>
The combination of “fallo” meaning “false” and “
<taxonomicName id="4C29248F1952FFE5FB87F8903F4F01E9" box="[1051,1121,924,950]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="lobus">lobus</taxonomicName>
”, referring to the posterior pronotal tubercle in the described species, is intended to indicate the relationship between these two species based on a comparison of morphological characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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