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<mods:title id="47F9FB38CA6C7723D8E124B6D57701A0">Systematics, phylogeny and biogeography of the Australasian leaf-curling orb-weaving spiders (Araneae: Araneidae: Zygiellinae), with a comparative analysis of retreat evolution</mods:title>
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<subSubSection id="C38BBBEDFFAD3171F8E95A0AFCDBFE52" pageId="79" pageNumber="1134" type="nomenclature">
<paragraph id="8B2EE866FFAD3171F8E95A0AFCDBFE14" blockId="79.[163,780,344,1133]" pageId="79" pageNumber="1134">
<heading id="D0665F0AFFAD3171F8E95A0AFCDBFE14" centered="true" fontSize="9" level="2" pageId="79" pageNumber="1134" reason="2">
<taxonomicName id="4C9193E5FFAD3171F8E95A0AFCD4FEF6" authority="Barrion &amp; Litsinger, 1995" authorityName="Barrion &amp; Litsinger" authorityYear="1995" box="[163,773,344,366]" class="Arachnida" family="Araneidae" genus="Phonognatha" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="species" species="guanga">
<emphasis id="B9E53474FFAD3171F8E95A0AFE7FFEF6" box="[163,430,344,366]" italics="true" pageId="79" pageNumber="1134">Phonognatha guanga</emphasis>
<bibRefCitation id="EF009597FFAD3171F9F05A0AFCD4FEF6" author="Barrion AT &amp; Litsinger JA" box="[442,773,344,366]" pageId="79" pageNumber="1134" refId="ref59110" refString="Barrion AT, Litsinger JA. 1995. Riceland spiders of the South and Southeast Asia. Wallingford, UK: CAB International." type="book" year="1995">Barrion &amp; Litsinger, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9E53474FFAD3171F8F15A25FE9DFE14" box="[187,332,375,396]" italics="true" pageId="79" pageNumber="1134">
<collectingCountry id="F386A8F6FFAD3171F8F15A25FE94FE14" box="[187,325,375,396]" name="Philippines" pageId="79" pageNumber="1134">Philippines</collectingCountry>
:
</emphasis>
Luzon Island,
<collectingRegion id="49552684FFAD3171FA4F5A25FD02FE15" box="[517,723,375,397]" country="Philippines" name="Laguna" pageId="79" pageNumber="1134">Laguna Province</collectingRegion>
, Los
</heading>
</paragraph>
<paragraph id="8B2EE866FFAD3171F8F15AC4FCDBFE52" blockId="79.[163,780,344,1133]" pageId="79" pageNumber="1134">
<materialsCitation id="3BF9E23BFFAD3171F8F15AC4FCDBFE52" collectingDate="1984-04-04" collectionCode="IRRI, T" pageId="79" pageNumber="1134" specimenCount="1" typeStatus="holotype">
Baños,
<collectionCode id="ED8070A3FFAD3171F9585AC4FE9AFE33" box="[274,331,406,427]" country="Philippines" httpUri="http://grbio.org/cool/tp71-5wyw" name="International Rice Research Institute" pageId="79" pageNumber="1134">IRRI</collectionCode>
Farm,
<date id="FF2FCEA6FFAD3171F9EB5AC4FDDBFE34" box="[417,522,406,428]" pageId="79" pageNumber="1134" value="1984-04-04">
<collectingDate id="EF6B374EFFAD3171F9EB5AC4FDDBFE34" box="[417,522,406,428]" pageId="79" pageNumber="1134" value="1984-04-04">4.iv.1984</collectingDate>
</date>
, A.
<collectionCode id="ED8070A3FFAD3171FA7C5AC4FD96FE33" box="[566,583,406,427]" name="Tavera, Department of Geology and Geophysics" pageId="79" pageNumber="1134">T</collectionCode>
. Barrion, female
<typeStatus id="542A56C4FFAD3171F8F15AE6FEF2FE52" box="[187,291,436,458]" pageId="79" pageNumber="1134" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED8070A3FFAD3171F97C5AE6FEA5FE51" box="[310,372,436,457]" country="Philippines" httpUri="http://grbio.org/cool/tp71-5wyw" name="International Rice Research Institute" pageId="79" pageNumber="1134">IRRI</collectionCode>
, not examined); Liliw, Tuybaana
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C38BBBEDFFAD3171F8F15A81FBB6FEAF" pageId="79" pageNumber="1134" type="description">
<paragraph id="8B2EE866FFAD3171F8F15A81FDCEFD90" blockId="79.[163,780,344,1133]" pageId="79" pageNumber="1134">
<materialsCitation id="3BF9E23BFFAD3171F8F15A81FDCAFD90" collectingDate="1979-03-14" collectionCode="T, IRRI" collectorName="J. Perfect" pageId="79" pageNumber="1134" specimenCount="1" typeStatus="paratype">
Village,
<date id="FF2FCEA6FFAD3171F9555A81FE4FFE71" box="[287,414,467,489]" pageId="79" pageNumber="1134" value="1979-03-14">
<collectingDate id="EF6B374EFFAD3171F9555A81FE4FFE71" box="[287,414,467,489]" pageId="79" pageNumber="1134" value="1979-03-14">14.iii.1979</collectingDate>
</date>
,
<collectionCode id="ED8070A3FFAD3171F9E65A81FE6FFE70" box="[428,446,467,488]" name="Tavera, Department of Geology and Geophysics" pageId="79" pageNumber="1134">T</collectionCode>
.
<collectorName id="26648DB0FFAD3171F9835A81FDEAFE71" box="[457,571,467,489]" pageId="79" pageNumber="1134">J. Perfect</collectorName>
, subadult female
<typeStatus id="542A56C4FFAD3171F8F15AA0FEF0FD9F" box="[187,289,498,519]" pageId="79" pageNumber="1134" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED8070A3FFAD3171F97A5AA0FEBBFD9F" box="[304,362,498,519]" country="Philippines" httpUri="http://grbio.org/cool/tp71-5wyw" name="International Rice Research Institute" pageId="79" pageNumber="1134">IRRI</collectionCode>
, not examined)
</materialsCitation>
.
</paragraph>
<paragraph id="8B2EE866FFAD3171F8F15942FBB6FEAF" blockId="79.[163,780,344,1133]" lastBlockId="79.[827,1444,197,311]" pageId="79" pageNumber="1134">
This species from the
<collectingCountry id="F386A8F6FFAD3171F9815942FD84FDBE" box="[459,597,528,550]" name="Philippines" pageId="79" pageNumber="1134">Philippines</collectingCountry>
was originally described as a member of
<taxonomicName id="4C9193E5FFAD3171F99C597DFDA0FDDC" authorityName="Simon" authorityYear="1894" box="[470,625,559,580]" class="Arachnida" family="Araneidae" genus="Phonognatha" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="genus">
<emphasis id="B9E53474FFAD3171F99C597DFDA0FDDC" box="[470,625,559,580]" italics="true" pageId="79" pageNumber="1134">Phonognatha</emphasis>
</taxonomicName>
without providing an explicit justification of why the Philippine species should be classified in the genus. Unlike
<taxonomicName id="4C9193E5FFAD3171F8E959D9FEECFD38" authorityName="Simon" authorityYear="1894" box="[163,317,651,672]" class="Arachnida" family="Araneidae" genus="Phonognatha" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="genus">
<emphasis id="B9E53474FFAD3171F8E959D9FEECFD38" box="[163,317,651,672]" italics="true" pageId="79" pageNumber="1134">Phonognatha</emphasis>
</taxonomicName>
or closely related taxa, this species has a high clypeus, abdomen without dark-on-white markings, truncate abdomen with a posterior tubercle, V-shaped epigynum, lack of copulatory ducts inside a capsule and non-lobed spermathecae. Furthermore, there is no note about an orb web with leaf retreat; this specimen was collected under leaves. This species is a member of
<taxonomicName id="4C9193E5FFAD3171F9055830FE08FCE0" authorityName="BLACKWALL" authorityYear="1859" box="[335,473,866,888]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="family">Linyphiidae</taxonomicName>
as pointed out by
<bibRefCitation id="EF009597FFAD3171FAEC5830FF3AFC0D" author="Hormiga G" pageId="79" pageNumber="1134" pagination="125 - 132" refId="ref60964" refString="Hormiga G. 1998. The spider genus Napometa (Araneae, Araneoidea, Linyphiidae). Journal of Arachnology 26: 125-132." type="journal article" year="1998">Hormiga (1998)</bibRefCitation>
. The illustrations of the epigynum of the
<typeStatus id="542A56C4FFAD3171FA9858D2FF07FC2C" pageId="79" pageNumber="1134" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EF009597FFAD3171F8AC58CDFDC7FC2D" author="Barrion AT &amp; Litsinger JA" box="[230,534,927,949]" pageId="79" pageNumber="1134" refId="ref59110" refString="Barrion AT, Litsinger JA. 1995. Riceland spiders of the South and Southeast Asia. Wallingford, UK: CAB International." type="book" year="1995">Barrion &amp; Litsinger, 1995</bibRefCitation>
: fig. 321F, G) clearly suggest that
<taxonomicName id="4C9193E5FFAD3171F97F58ECFE5BFC4B" authorityName="Barrion &amp; Litsinger" authorityYear="1995" box="[309,394,958,979]" class="Arachnida" family="Araneidae" genus="Phonognatha" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="species" species="guanga">
<emphasis id="B9E53474FFAD3171F97F58ECFE5BFC4B" box="[309,394,958,979]" italics="true" pageId="79" pageNumber="1134">guanga</emphasis>
</taxonomicName>
is a member of the genus
<taxonomicName id="4C9193E5FFAD3171FAE458ECFCD4FC4B" authorityName="BLACKWALL" authorityYear="1833" box="[686,773,958,979]" class="Arachnida" family="Linyphiidae" genus="Neriene" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="genus">
<emphasis id="B9E53474FFAD3171FAE458ECFCD4FC4B" box="[686,773,958,979]" italics="true" pageId="79" pageNumber="1134">Neriene</emphasis>
</taxonomicName>
, as delimited and diagnosed in
<bibRefCitation id="EF009597FFAD3171F9B1588FFCDAFC6A" author="van Helsdingen PJ" box="[507,779,988,1010]" pageId="79" pageNumber="1134" pagination="1 - 303" refId="ref60797" refString="van Helsdingen PJ. 1969. A reclassification of the species of Linyphia Latreille based on the functioning of the genitalia (Araneae, Linpyhiidae), I. Zoologische Verhandelingen 105: 1-303." type="journal article" year="1969">van Helsdingens (1969)</bibRefCitation>
monograph, based on the presence of spirally coiled grooves in the atrium [e.g. see
<bibRefCitation id="EF009597FFAD3171F9B15F48FCDAFBB7" author="van Helsdingen PJ" box="[507,779,1049,1071]" pageId="79" pageNumber="1134" pagination="1 - 303" refId="ref60797" refString="van Helsdingen PJ. 1969. A reclassification of the species of Linyphia Latreille based on the functioning of the genitalia (Araneae, Linpyhiidae), I. Zoologische Verhandelingen 105: 1-303." type="journal article" year="1969">van Helsdingens (1969)</bibRefCitation>
figs 347 and 386 for two examples of the epigyna of Oriental species].
<bibRefCitation id="EF009597FFAD3171F9CE5F05FD01FBF5" author="Barrion AT &amp; Litsinger JA" box="[388,720,1111,1133]" pageId="79" pageNumber="1134" refId="ref59110" refString="Barrion AT, Litsinger JA. 1995. Riceland spiders of the South and Southeast Asia. Wallingford, UK: CAB International." type="book" year="1995">Barrion &amp; Litsinger (1995)</bibRefCitation>
also describe two species of
<taxonomicName id="4C9193E5FFAD3171FC285B94FB11FF43" authorityName="BLACKWALL" authorityYear="1833" box="[1122,1216,198,219]" class="Arachnida" family="Linyphiidae" genus="Neriene" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="genus">
<emphasis id="B9E53474FFAD3171FC285B94FB11FF43" box="[1122,1216,198,219]" italics="true" pageId="79" pageNumber="1134">Neriene</emphasis>
</taxonomicName>
under the tetragnathid genus
<taxonomicName id="4C9193E5FFAD3171FB925BB6FBC0FF61" box="[984,1041,228,249]" class="Arachnida" family="Tetragnathidae" genus="Meta" kingdom="Animalia" order="Araneae" pageId="79" pageNumber="1134" phylum="Arthropoda" rank="genus">
<emphasis id="B9E53474FFAD3171FB925BB6FBC0FF61" box="[984,1041,228,249]" italics="true" pageId="79" pageNumber="1134">Meta</emphasis>
</taxonomicName>
, as pointed out by
<bibRefCitation id="EF009597FFAD3171FCA05BB6FA4CFF61" author="Hormiga G" box="[1258,1437,228,250]" pageId="79" pageNumber="1134" pagination="125 - 132" refId="ref60964" refString="Hormiga G. 1998. The spider genus Napometa (Araneae, Araneoidea, Linyphiidae). Journal of Arachnology 26: 125-132." type="journal article" year="1998">Hormiga (1998)</bibRefCitation>
.
<bibRefCitation id="EF009597FFAD3171FB715A50FBECFE8F" author="Wunderlich J" box="[827,1085,258,280]" pageId="79" pageNumber="1134" pagination="44 - 139" refId="ref63907" refString="Wunderlich J. 2008. Descriptions of fossil spider (Araneae) taxa mainly in Baltic amber, as well as on certain related extant taxa. Beitrage zur Araneologie 5: 44-139." type="journal article" year="2008">Wunderlich (2008: 94)</bibRefCitation>
had also suggested that these two species are linyphiids.
</paragraph>
</subSubSection>
</treatment>
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