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<document id="536082BE09B0E495E240C6BE7DD015C6" ID-DOI="10.11646/zootaxa.5323.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8212306" ID-ZooBank="DAAF3E31-A245-4421-A41B-E9BE1538ABF9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691078968945" checkinUser="plazi" docAuthor="Noguera, Felipe A. &amp; Santos-Silva, Antonio" docDate="2023" docId="FE18879D9B1E831CFF2BFC1AFA4DFDF5" docLanguage="en" docName="zootaxa.5323.4.1.pdf" docOrigin="Zootaxa 5323 (4)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynidius Lacordaire 1869" docType="key" docVersion="2" lastPageNumber="475" masterDocId="0221FFE59B098304FFBCFFF4FFD1FFDF" masterDocTitle="New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)" masterLastPageNumber="476" masterPageNumber="451" pageNumber="474" updateTime="1691079457083" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B553A24A8BF61727C4FD1318E6C77602">New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
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<mods:namePart id="F81369CFF9AF5E4C0C8FA04BDF13F6C4">Noguera, Felipe A.</mods:namePart>
<mods:affiliation id="3D58139F683A29E6B69927C508B48BE5">Estacioìn de Biologiìa Chamela, Instituto de Biologiìa, Universidad Nacional Autoìnoma de Meìxico, Apartado Postal 21, San Patricio, Jalisco 48980, Mexico.</mods:affiliation>
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<mods:namePart id="D120C5A7AC21C2C85F457EDF96728853">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="7A87FD3031DC4C237D7B8031EFEB4490">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:date id="FF36D527665D0CDE479320DEF871C967">2023</mods:date>
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<mods:number id="B4708ED36582DA7C2460BDE0957F1DA7">2023-08-03</mods:number>
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<mods:identifier id="B380FE2FA54EEBF0AE8E6336BC8358D7" type="DOI">10.11646/zootaxa.5323.4.1</mods:identifier>
<mods:identifier id="0A7AD28090BC4C99668C26E8D4EF7101" type="ISSN">1175-5326</mods:identifier>
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<treatment id="FE18879D9B1E831CFF2BFC1AFA4DFDF5" LSID="urn:lsid:plazi:treatment:FE18879D9B1E831CFF2BFC1AFA4DFDF5" httpUri="http://treatment.plazi.org/id/FE18879D9B1E831CFF2BFC1AFA4DFDF5" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
<subSubSection id="3EAB65009B1E8313FF2BFC1AFE25FBD6" box="[151,500,1006,1033]" pageId="23" pageNumber="474" type="nomenclature">
<paragraph id="760E368B9B1E8313FF2BFC1AFE25FBD6" blockId="23.[151,500,1006,1033]" box="[151,500,1006,1033]" pageId="23" pageNumber="474">
<heading id="2D4681E79B1E8313FF2BFC1AFE25FBD6" bold="true" box="[151,500,1006,1033]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
<emphasis id="44C5EA999B1E8313FF2BFC1AFE25FBD6" bold="true" box="[151,500,1006,1033]" pageId="23" pageNumber="474">
Key to species of
<taxonomicName id="B1B14D089B1E8313FED2FC1AFE25FBD6" ID-CoL="63MKQ" authorityName="Lacordaire" authorityYear="1869" box="[366,500,1006,1033]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="44C5EA999B1E8313FED2FC1AFE25FBD6" bold="true" box="[366,500,1006,1033]" italics="true" pageId="23" pageNumber="474">Phrynidius</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B1E831CFF2BFBC7FA4DFDF5" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474" type="key">
<keyStep id="CD452E2E9B1E8313FF2BFBC7FA4DFBB9" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFBC7FA4DFB97" blockId="23.[151,1437,1075,2036]" box="[151,1436,1075,1097]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFBC7FA4DFB97" box="[151,1436,1075,1097]" pageId="23" pageNumber="474">
1. Elytra with distinct, short, erect spatulate setae (
<emphasis id="44C5EA999B1E8313FD74FBC0FD3AFB96" box="[712,747,1076,1097]" italics="true" pageId="23" pageNumber="474">e.g.</emphasis>
<figureCitation id="EE8A2A0E9B1E8313FD4CFBC7FCE4FB97" box="[752,821,1075,1097]" captionStart="FIGURES 10–13" captionStartId="5.[152,270,1322,1346]" captionTargetBox="[151,1436,238,1298]" captionTargetId="figure-35@5.[151,1436,237,1298]" captionTargetPageId="5" captionText="FIGURES 10–13. Phrynidius poriferus sp. nov., holotype female: 10) Dorsal habitus; 11) Ventral habitus; 12) Lateral habitus; 13) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212316" httpUri="https://zenodo.org/record/8212316/files/figure.png" pageId="23" pageNumber="474">Fig. 12</figureCitation>
)................................................. 2
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFBA5FA4DFBB9" blockId="23.[151,1437,1075,2036]" box="[151,1436,1104,1126]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFBA5FA4DFBB9" box="[151,1436,1104,1126]" pageId="23" pageNumber="474">
- Elytra without erect spatulate setae (
<emphasis id="44C5EA999B1E8313FDE4FBA5FDAAFBB9" box="[600,635,1105,1126]" italics="true" pageId="23" pageNumber="474">e.g.</emphasis>
<figureCitation id="EE8A2A0E9B1E8313FD3CFBA4FD16FBB9" box="[640,711,1104,1126]" captionStart="FIGURES 46–51" captionStartId="21.[152,270,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURES 46–51. Phrynidius echinus Bates, 1880. 46–49) Male: 46) Dorsal habitus; 47) Ventral habitus; 48) Lateral habitus; 49) Head, frontal view. 50–51) Female: 50) Dorsal habitus; 51) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212338" httpUri="https://zenodo.org/record/8212338/files/figure.png" pageId="23" pageNumber="474">Fig. 48</figureCitation>
), or with erect setae almost absent (
<emphasis id="44C5EA999B1E8313FBA3FBA5FB93FBB9" box="[1055,1090,1105,1126]" italics="true" pageId="23" pageNumber="474">e.g.</emphasis>
<figureCitation id="EE8A2A0E9B1E8313FBF4FBA4FB5CFBB9" box="[1096,1165,1104,1126]" captionStart="FIGURES 22–28" captionStartId="11.[152,270,1833,1857]" captionTargetBox="[151,1436,237,1809]" captionTargetId="figure-23@11.[151,1436,237,1809]" captionTargetPageId="11" captionText="FIGURES 22–28. Phrynidius wibmeri sp. nov. 22–25) Holotype male: 22) Dorsal habitus; 23) Ventral habitus; 24) Lateral habitus; 25) Head, frontal view. 26) Paratype male, dorsal habitus. 27–28) Paratype female: 27) Dorsal habitus; 28) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212326" httpUri="https://zenodo.org/record/8212326/files/figure.png" pageId="23" pageNumber="474">Fig. 24</figureCitation>
).................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFB9AFA4DFB7E" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFB9AFA4DFB5C" blockId="23.[151,1437,1075,2036]" box="[151,1436,1134,1156]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFB9AFA4DFB5C" box="[151,1436,1134,1156]" pageId="23" pageNumber="474">
2(1). Antennal tubercles contiguous basally (e.g.
<figureCitation id="EE8A2A0E9B1E8313FD22FB9AFD32FB5C" box="[670,739,1134,1156]" captionStart="FIGURES 14–17" captionStartId="7.[152,270,1519,1543]" captionTargetBox="[152,1435,382,1495]" captionTargetId="figure-55@7.[152,1436,381,1495]" captionTargetPageId="7" captionText="FIGURES 14–17. Phrynidius skillmani sp. nov., holotype female: 14) Dorsal habitus; 15) Ventral habitus; 16) Lateral habitus; 17) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212320" httpUri="https://zenodo.org/record/8212320/files/figure.png" pageId="23" pageNumber="474">Fig. 14</figureCitation>
)........................................................ 3
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFB78FA4DFB7E" blockId="23.[151,1437,1075,2036]" box="[151,1436,1163,1185]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFB78FA4DFB7E" box="[151,1436,1163,1185]" pageId="23" pageNumber="474">
- Antennal tubercles not contiguous basally (e.g.
<figureCitation id="EE8A2A0E9B1E8313FD7FFB7FFCD9FB7F" box="[707,776,1163,1185]" captionStart="FIGURES 22–28" captionStartId="11.[152,270,1833,1857]" captionTargetBox="[151,1436,237,1809]" captionTargetId="figure-23@11.[151,1436,237,1809]" captionTargetPageId="11" captionText="FIGURES 22–28. Phrynidius wibmeri sp. nov. 22–25) Holotype male: 22) Dorsal habitus; 23) Ventral habitus; 24) Lateral habitus; 25) Head, frontal view. 26) Paratype male, dorsal habitus. 27–28) Paratype female: 27) Dorsal habitus; 28) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212326" httpUri="https://zenodo.org/record/8212326/files/figure.png" pageId="23" pageNumber="474">Fig. 22</figureCitation>
)..................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFB5DFA4DFB04" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFB5DFA4DFB61" blockId="23.[151,1437,1075,2036]" box="[151,1436,1192,1214]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFB5DFA4DFB61" box="[151,1436,1192,1214]" pageId="23" pageNumber="474">
3(2). Erect setae on elytra present only apically.
<collectingCountry id="0EA6761B9B1E8313FD29FB5CFD2CFB61" box="[661,765,1192,1214]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
...................................
<taxonomicName id="B1B14D089B1E8313FB15FB5DFA98FB62" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1193,1353,1192,1214]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="guatemalensis" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB15FB5DFA98FB62" box="[1193,1353,1192,1214]" italics="true" pageId="23" pageNumber="474">P. guatemalensis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECFB5DFA4DFB61" bold="true" box="[1360,1436,1193,1214]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECFB5DFA4DFB61" box="[1360,1436,1193,1214]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFB32FA4DFB04" blockId="23.[151,1437,1075,2036]" box="[151,1436,1222,1244]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFB32FA4DFB04" box="[151,1436,1222,1244]" pageId="23" pageNumber="474">- Erect setae on elytra abundant throughout................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFB17FA4DFAC9" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFB17FA4DFB27" blockId="23.[151,1437,1075,2036]" box="[151,1436,1251,1273]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFB17FA4DFB27" box="[151,1436,1251,1273]" pageId="23" pageNumber="474">
4(3). Antennomere III 0.6 times longer than IV; sides of prothorax distinctly rounded (
<figureCitation id="EE8A2A0E9B1E8313FBB8FB17FB9DFB27" box="[1028,1100,1251,1273]" captionStart="FIGURES 10–13" captionStartId="5.[152,270,1322,1346]" captionTargetBox="[151,1436,238,1298]" captionTargetId="figure-35@5.[151,1436,237,1298]" captionTargetPageId="5" captionText="FIGURES 10–13. Phrynidius poriferus sp. nov., holotype female: 10) Dorsal habitus; 11) Ventral habitus; 12) Lateral habitus; 13) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212316" httpUri="https://zenodo.org/record/8212316/files/figure.png" pageId="23" pageNumber="474">Fig. 10</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FBE1FB17FB6BFB26" box="[1117,1210,1251,1273]" name="Honduras" pageId="23" pageNumber="474">Honduras</collectingCountry>
..
<taxonomicName id="B1B14D089B1E8313FB66FB10FA9BFB27" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1242,1354,1251,1273]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="poriferus" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB66FB10FA9BFB27" box="[1242,1354,1251,1273]" italics="true" pageId="23" pageNumber="474">P. poriferus</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECFB17FA4DFB27" bold="true" box="[1360,1436,1251,1272]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECFB17FA4DFB27" box="[1360,1436,1251,1272]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFAF5FA4DFAC9" blockId="23.[151,1437,1075,2036]" box="[151,1436,1280,1302]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFAF5FA4DFAC9" box="[151,1436,1280,1302]" pageId="23" pageNumber="474">
- Antennomere III twice length of IV; sides of prothorax slightly rounded (
<figureCitation id="EE8A2A0E9B1E8313FC04FAF4FBD1FAC9" box="[952,1024,1280,1302]" captionStart="FIGURES 14–17" captionStartId="7.[152,270,1519,1543]" captionTargetBox="[152,1435,382,1495]" captionTargetId="figure-55@7.[152,1436,381,1495]" captionTargetPageId="7" captionText="FIGURES 14–17. Phrynidius skillmani sp. nov., holotype female: 14) Dorsal habitus; 15) Ventral habitus; 16) Lateral habitus; 17) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212320" httpUri="https://zenodo.org/record/8212320/files/figure.png" pageId="23" pageNumber="474">Fig. 14</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FBADFAF4FBA8FAC9" box="[1041,1145,1280,1302]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
........
<taxonomicName id="B1B14D089B1E8313FB66FAF5FA9BFACA" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1242,1354,1280,1302]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="skillmani" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB66FAF5FA9BFACA" box="[1242,1354,1280,1302]" italics="true" pageId="23" pageNumber="474">P. skillmani</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECFAF5FA4DFAC9" bold="true" box="[1360,1436,1281,1302]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECFAF5FA4DFAC9" box="[1360,1436,1281,1302]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFAEAFA4DFA8F" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFAEAFA4DFAEC" blockId="23.[151,1437,1075,2036]" box="[151,1436,1310,1332]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFAEAFA4DFAEC" box="[151,1436,1310,1332]" pageId="23" pageNumber="474">
5(2). Antennomere IV about as long as V and VI together (
<figureCitation id="EE8A2A0E9B1E8313FD47FAEAFC93FAEC" box="[763,834,1310,1332]" captionStart="FIGURES 18–21" captionStartId="9.[152,270,1432,1456]" captionTargetBox="[151,1436,346,1409]" captionTargetId="figure-52@9.[151,1436,345,1409]" captionTargetPageId="9" captionText="FIGURES 18–21. Phrynidius obrieni sp. nov., holotype female: 18) Dorsal habitus; 19) Ventral habitus; 20) Lateral habitus; 21) Head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.8212322" httpUri="https://zenodo.org/record/8212322/files/figure.png" pageId="23" pageNumber="474">Fig. 18</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FCE8FAEAFC60FAEB" box="[852,945,1310,1332]" name="Honduras" pageId="23" pageNumber="474">Honduras</collectingCountry>
.........................
<taxonomicName id="B1B14D089B1E8313FB51FAEAFA98FAEC" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1261,1353,1310,1331]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="obrieni" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB51FAEAFA98FAEC" box="[1261,1353,1310,1331]" italics="true" pageId="23" pageNumber="474">P. obrieni</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECFAEAFA4DFAEC" bold="true" box="[1360,1436,1310,1331]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECFAEAFA4DFAEC" box="[1360,1436,1310,1331]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFAC8FA4DFA8F" blockId="23.[151,1437,1075,2036]" box="[151,1436,1339,1361]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFAC8FA4DFA8F" box="[151,1436,1339,1361]" pageId="23" pageNumber="474">
- Antennomere IV distinctly longer than V and VI together (
<figureCitation id="EE8A2A0E9B1E8313FC91FACFFC4DFA8F" box="[813,924,1339,1361]" captionStart="FIGURES 22–28" captionStartId="11.[152,270,1833,1857]" captionTargetBox="[151,1436,237,1809]" captionTargetId="figure-23@11.[151,1436,237,1809]" captionTargetPageId="11" captionText="FIGURES 22–28. Phrynidius wibmeri sp. nov. 22–25) Holotype male: 22) Dorsal habitus; 23) Ventral habitus; 24) Lateral habitus; 25) Head, frontal view. 26) Paratype male, dorsal habitus. 27–28) Paratype female: 27) Dorsal habitus; 28) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212326" httpUri="https://zenodo.org/record/8212326/files/figure.png" pageId="23" pageNumber="474">Figs 22, 27</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FC11FACFFC27FA8E" box="[941,1014,1339,1361]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FBBFFACFFB8DFA8F" box="[1027,1116,1339,1360]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="474">Veracruz</collectingRegion>
)...........
<taxonomicName id="B1B14D089B1E8313FB5FFAC8FA9BFA8F" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1251,1354,1339,1361]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="wibmeri" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB5FFAC8FA9BFA8F" box="[1251,1354,1339,1361]" italics="true" pageId="23" pageNumber="474">P. wibmeri</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECFACFFA4DFA8F" bold="true" box="[1360,1436,1339,1360]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECFACFFA4DFA8F" box="[1360,1436,1339,1360]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFAACFA4DFA54" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFAACFA4DFAB1" blockId="23.[151,1437,1075,2036]" box="[151,1436,1368,1390]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFAACFA4DFAB1" box="[151,1436,1368,1390]" pageId="23" pageNumber="474">6(1). Antennomere III at most as long as scape................................................................ 7</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFA82FA4DFA54" blockId="23.[151,1437,1075,2036]" box="[151,1436,1398,1420]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFA82FA4DFA54" box="[151,1436,1398,1420]" pageId="23" pageNumber="474">- Antennomere III longer than scape.................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFA67FA4DFA19" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFA67FA4DFA77" blockId="23.[151,1437,1075,2036]" box="[151,1436,1427,1449]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFA67FA4DFA77" box="[151,1436,1427,1449]" pageId="23" pageNumber="474">
7(6). Pronotum with distinct elevated anterocentral gibbosity (
<emphasis id="44C5EA999B1E8313FCA0FA60FCEEFA76" box="[796,831,1428,1449]" italics="true" pageId="23" pageNumber="474">e.g.</emphasis>
<figureCitation id="EE8A2A0E9B1E8313FCF9FA67FC5BFA77" box="[837,906,1427,1449]" captionStart="FIGURES 29–35" captionStartId="15.[152,270,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-17@15.[151,1436,181,1823]" captionTargetPageId="15" captionText="FIGURES 29–35. Phrynidius pallifemoralis sp. nov. 29–33) Holotype male: 29) Dorsal habitus; 30) Ventral habitus; 31) Lateral habitus; 32) Head, frontal view; 33) Prothorax, oblique view. 34–35) Female paratype: 34) Dorsal habitus; 35) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212328" httpUri="https://zenodo.org/record/8212328/files/figure.png" pageId="23" pageNumber="474">Fig. 33</figureCitation>
).......................................... 8
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BFA45FA4DFA19" blockId="23.[151,1437,1075,2036]" box="[151,1436,1456,1478]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFA45FA4DFA19" box="[151,1436,1456,1478]" pageId="23" pageNumber="474">
- Pronotum without anterocentral gibbosity or anterocentral gibbosity almost indistinct (
<emphasis id="44C5EA999B1E8313FB86FA45FB8CFA19" box="[1082,1117,1457,1478]" italics="true" pageId="23" pageNumber="474">e.g.</emphasis>
<figureCitation id="EE8A2A0E9B1E8313FBDFFA44FB79FA19" box="[1123,1192,1456,1478]" captionStart="FIGURES 52–56. 52–54" captionStartId="22.[152,270,1506,1530]" captionTargetBox="[161,1436,186,1476]" captionTargetId="figure-26@22.[151,1436,181,1481]" captionTargetPageId="22" captionText="FIGURES 52–56. 52–54) Phrynidius echinus Bates, 1880, holotype: 52) Dorsal habitus; 53) Lateral habitus; 54) Labels. 55–56) Phrynidius armatus Linsley, 1933, holotype male: 55) Dorsal habitus; 56) Lateral habitus. Figures 52–54 by Keita Matsumoto (BMNH); 55–56 by Ranchel Diaz-Bastin (CASC)." figureDoi="http://doi.org/10.5281/zenodo.8212340" httpUri="https://zenodo.org/record/8212340/files/figure.png" pageId="23" pageNumber="474">Fig. 55</figureCitation>
).................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BFA3AFA4DF9E4" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BFA3AFA4DF9DF" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BFA3AFA4DF9DF" pageId="23" pageNumber="474">
<materialsCitation id="C6D93CD69B1E8313FF2BFA3AFA49F9DF" country="Mexico" location="Oaxaca" pageId="23" pageNumber="474" specimenCount="1" stateProvince="Oaxaca">
8(7). Body stout (
<figureCitation id="EE8A2A0E9B1E8313FED2FA3AFE0CFA3C" box="[366,477,1486,1508]" captionStart="FIGURES 29–35" captionStartId="15.[152,270,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-17@15.[151,1436,181,1823]" captionTargetPageId="15" captionText="FIGURES 29–35. Phrynidius pallifemoralis sp. nov. 29–33) Holotype male: 29) Dorsal habitus; 30) Ventral habitus; 31) Lateral habitus; 32) Head, frontal view; 33) Prothorax, oblique view. 34–35) Female paratype: 34) Dorsal habitus; 35) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212328" httpUri="https://zenodo.org/record/8212328/files/figure.png" pageId="23" pageNumber="474">Figs 29, 34</figureCitation>
); anterocentral gibbosity on the pronotum uniformly convex dorsally, not sinuous laterally (
<figureCitation id="EE8A2A0E9B1E8313FAF6FA3AFA40FA3C" box="[1354,1425,1486,1508]" captionStart="FIGURES 29–35" captionStartId="15.[152,270,1847,1871]" captionTargetBox="[151,1436,181,1823]" captionTargetId="figure-17@15.[151,1436,181,1823]" captionTargetPageId="15" captionText="FIGURES 29–35. Phrynidius pallifemoralis sp. nov. 29–33) Holotype male: 29) Dorsal habitus; 30) Ventral habitus; 31) Lateral habitus; 32) Head, frontal view; 33) Prothorax, oblique view. 34–35) Female paratype: 34) Dorsal habitus; 35) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212328" httpUri="https://zenodo.org/record/8212328/files/figure.png" pageId="23" pageNumber="474">Fig. 33</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FF46FA1FFE92F9DE" box="[250,323,1515,1537]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FEECFA1FFE4DF9DF" box="[336,412,1515,1536]" country="Mexico" name="Oaxaca" pageId="23" pageNumber="474">Oaxaca</collectingRegion>
,
<collectingRegion id="B475F8699B1E8313FE19FA1FFE25F9DE" box="[421,500,1515,1537]" country="Mexico" name="Hidalgo" pageId="23" pageNumber="474">Hidalgo</collectingRegion>
).........................................................
<taxonomicName id="B1B14D089B1E8313FB17FA18FA9BF9DF" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1195,1354,1515,1537]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="pallifemoralis" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB17FA18FA9BF9DF" box="[1195,1354,1515,1537]" italics="true" pageId="23" pageNumber="474">P. pallifemoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FF657E29B1E8313FAECFA1FFA49F9DF" box="[1360,1432,1515,1536]" pageId="23" pageNumber="474" rank="species">
<emphasis id="44C5EA999B1E8313FAECFA1FFA49F9DF" bold="true" box="[1360,1432,1515,1536]" pageId="23" pageNumber="474">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="44C5EA999B1E8313FA24FA1FFA4DF9DF" bold="true" box="[1432,1436,1515,1536]" pageId="23" pageNumber="474">.</emphasis>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF9FDFA4DF9E4" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF9FDFA4DF9E4" pageId="23" pageNumber="474">
- Body slender (
<figureCitation id="EE8A2A0E9B1E8313FE34F9FCFE2BF9C1" box="[392,506,1544,1566]" captionStart="FIGURES 36–41" captionStartId="17.[152,270,1966,1990]" captionTargetBox="[216,1371,309,1943]" captionTargetId="figure-88@17.[216,1371,309,1943]" captionTargetPageId="17" captionText="FIGURES 36–41. Phrynidius oaxacanensis sp. nov. 36–39) Holotype male: 36) Dorsal habitus; 37) Ventral habitus; 38) Lateral habitus; 39) Head, frontal view. 40–41) Paratype female: 40) Dorsal habitus; 41) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="23" pageNumber="474">Figs 36, 40</figureCitation>
); anterocentral gibbosity on the pronotum irregular dorsally, sinuous laterally (
<figureCitation id="EE8A2A0E9B1E8313FB44F9FCFA90F9C1" box="[1272,1345,1544,1566]" captionStart="FIGURES 36–41" captionStartId="17.[152,270,1966,1990]" captionTargetBox="[216,1371,309,1943]" captionTargetId="figure-88@17.[216,1371,309,1943]" captionTargetPageId="17" captionText="FIGURES 36–41. Phrynidius oaxacanensis sp. nov. 36–39) Holotype male: 36) Dorsal habitus; 37) Ventral habitus; 38) Lateral habitus; 39) Head, frontal view. 40–41) Paratype female: 40) Dorsal habitus; 41) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212332" httpUri="https://zenodo.org/record/8212332/files/figure.png" pageId="23" pageNumber="474">Fig. 38</figureCitation>
).
<collectingCountry id="0EA6761B9B1E8313FAEFF9FCFA4DF9C1" box="[1363,1436,1544,1566]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FEBCF9D2FE9AF9E4" box="[256,331,1574,1595]" country="Mexico" name="Oaxaca" pageId="23" pageNumber="474">Oaxaca</collectingRegion>
).......................................................................
<taxonomicName id="B1B14D089B1E8313FB0DF9D2FA98F9E4" authority="Noguera &amp; Santos-Silva, 2023" authorityName="Noguera &amp; Santos-Silva" authorityYear="2023" box="[1201,1353,1574,1595]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="oaxacanensis" status="sp. nov.">
<emphasis id="44C5EA999B1E8313FB0DF9D2FA98F9E4" box="[1201,1353,1574,1595]" italics="true" pageId="23" pageNumber="474">P. oaxacanensis</emphasis>
</taxonomicName>
<emphasis id="44C5EA999B1E8313FAECF9D2FA4DF9E4" bold="true" box="[1360,1436,1574,1595]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="5FF657E29B1E8313FAECF9D2FA4DF9E4" box="[1360,1436,1574,1595]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BF9B7FA4DF9A9" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF9B7FA4DF987" blockId="23.[151,1437,1075,2036]" box="[151,1436,1603,1625]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF9B7FA4DF987" box="[151,1436,1603,1625]" pageId="23" pageNumber="474">9(7). Scape longer than pedicel and antennomere III together.................................................... 10</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF995FA4DF9A9" blockId="23.[151,1437,1075,2036]" box="[151,1436,1632,1654]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF995FA4DF9A9" box="[151,1436,1632,1654]" pageId="23" pageNumber="474">- Scape shorter or as long as pedicel and antennomere III together............................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BF98AFA4DF911" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF98AFA4DF96E" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF98AFA4DF96E" pageId="23" pageNumber="474">
10(9). Elytra elongate, pubescent.
<collectingCountry id="0EA6761B9B1E8313FDB4F98AFD80F94B" box="[520,593,1662,1684]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FDE3F98AFD60F94B" box="[607,689,1662,1684]" country="Mexico" name="Hidalgo" pageId="23" pageNumber="474">Hidalgo</collectingRegion>
;
<collectingRegion id="B475F8699B1E8313FD07F98AFCC2F94C" box="[699,787,1662,1683]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="474">Veracruz</collectingRegion>
, see also Volcán de Orizaba (
<collectingRegion id="B475F8699B1E8313FB8EF98AFB58F94C" box="[1074,1161,1662,1683]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="23" pageNumber="474">Veracruz</collectingRegion>
and/or
<collectingRegion id="B475F8699B1E8313FB69F98AFAC6F94B" box="[1237,1303,1662,1684]" country="Mexico" name="Puebla" pageId="23" pageNumber="474">Puebla</collectingRegion>
—
<bibRefCitation id="12204B7A9B1E8313FA93F98AFEBEF96E" author="Selander, B. S. &amp; Vaurie, P." pageId="23" pageNumber="474" pagination="1 - 70" refId="ref17591" refString="Selander, B. S. &amp; Vaurie, P. (1962) A gazetteer to accompany the &quot; Insecta &quot; volumes of the &quot; Biologia Centrali-Americana. &quot; American Museum Novitates, 2099, 1 - 70." type="journal article" year="1962">Selander &amp; Vaurie 1962</bibRefCitation>
)....................................................................
<taxonomicName id="B1B14D089B1E8313FB10F968FA4DF96E" authority="(Say, 1835)" baseAuthorityName="Say" baseAuthorityYear="1835" box="[1196,1436,1691,1713]" class="Insecta" family="Cerambycidae" genus="Moneilema" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="44C5EA999B1E8313FB10F968FAF9F96F" box="[1196,1320,1691,1713]" italics="true" pageId="23" pageNumber="474">P. inaequalis</emphasis>
(
<bibRefCitation id="12204B7A9B1E8313FA89F96FFA44F96E" author="Say, T." box="[1333,1429,1691,1713]" pageId="23" pageNumber="474" pagination="151 - 203" refId="ref17554" refString="Say, T. (1835) Descriptions of New North American coleopterous insects, and observations on some already described. Boston Journal of Natural History, 1 (2), 151 - 203." type="journal article" year="1835">Say, 1835</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF94DFA4DF911" blockId="23.[151,1437,1075,2036]" box="[151,1436,1720,1742]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF94DFA4DF911" box="[151,1436,1720,1742]" pageId="23" pageNumber="474">
- Elytra ovoid, subglabrous.
<collectingCountry id="0EA6761B9B1E8313FE41F94CFD97F911" box="[509,582,1720,1742]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FDEFF94CFD72F911" box="[595,675,1720,1742]" country="Mexico" name="Chiapas" pageId="23" pageNumber="474">Chiapas</collectingRegion>
).......................
<taxonomicName id="B1B14D089B1E8313FC76F94DFA4DF911" authority="Gutierrez, Toledo &amp; Noguera, 2020" authorityName="Gutierrez, Toledo &amp; Noguera" authorityYear="2020" box="[970,1436,1720,1742]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="diminutus">
<emphasis id="44C5EA999B1E8313FC76F94DFB90F912" box="[970,1089,1720,1742]" italics="true" pageId="23" pageNumber="474">P. diminutus</emphasis>
Gutiérrez, Toledo &amp; Noguera, 2020
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BF922FA4DF8D7" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF922FA4DF933" blockId="23.[151,1437,1075,2036]" box="[151,1436,1750,1772]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF922FA4DF933" box="[151,1436,1750,1772]" pageId="23" pageNumber="474">
11(9). Antennomere IV as long as V–VI together.
<collectingCountry id="0EA6761B9B1E8313FD20F922FCD7F933" box="[668,774,1750,1772]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
,
<collectingCountry id="0EA6761B9B1E8313FCB3F922FCBEF933" box="[783,879,1750,1772]" name="Honduras" pageId="23" pageNumber="474">Honduras</collectingCountry>
,
<collectingCountry id="0EA6761B9B1E8313FCC5F922FC0DF933" box="[889,988,1750,1772]" name="Nicaragua" pageId="23" pageNumber="474">Nicaragua</collectingCountry>
.....................
<taxonomicName id="B1B14D089B1E8313FB64F922FA4DF933" authority="Bates, 1885" authorityName="Bates" authorityYear="1885" box="[1240,1436,1750,1772]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="44C5EA999B1E8313FB64F922FAF4F934" box="[1240,1317,1750,1771]" italics="true" pageId="23" pageNumber="474">P. asper</emphasis>
<bibRefCitation id="12204B7A9B1E8313FA97F922FA4DF933" author="Bates, H. W." box="[1323,1436,1750,1772]" pageId="23" pageNumber="474" refId="ref16431" refString="Bates, H. W. (1885) Supplement to Longicornia. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 249 - 436." type="book" year="1885">Bates, 1885</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF900FA4DF8D7" blockId="23.[151,1437,1075,2036]" box="[151,1436,1779,1801]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF900FA4DF8D7" box="[151,1436,1779,1801]" pageId="23" pageNumber="474">- Antennomere IV shorter than V–VI together............................................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BF8E5FA4DF89C" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF8E5FA4DF8F9" blockId="23.[151,1437,1075,2036]" box="[151,1436,1808,1830]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF8E5FA4DF8F9" box="[151,1436,1808,1830]" pageId="23" pageNumber="474">
12(11). Scutellum more than twice as wide as long.
<collectingCountry id="0EA6761B9B1E8313FD1DF8E4FD3CF8F9" box="[673,749,1808,1830]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
,
<collectingCountry id="0EA6761B9B1E8313FD4AF8E4FC8FF8F9" box="[758,862,1808,1830]" name="Guatemala" pageId="23" pageNumber="474">Guatemala</collectingCountry>
...........................
<taxonomicName id="B1B14D089B1E8313FB12F8E5FA4DF8F9" authority="Linsley, 1933" authorityName="Linsley" authorityYear="1933" box="[1198,1436,1808,1830]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B1E8313FB12F8E5FAC5F8F9" box="[1198,1300,1809,1830]" italics="true" pageId="23" pageNumber="474">P. armatus</emphasis>
<bibRefCitation id="12204B7A9B1E8313FAA6F8E4FA4DF8F9" author="Linsley, E. G." box="[1306,1436,1808,1830]" pageId="23" pageNumber="474" pagination="131 - 132" refId="ref17088" refString="Linsley, E. G. (1933) A new longicorn beetle from Central America. The Pan-Pacific Entomologist, 9 (3), 131 - 132." type="journal article" year="1933">Linsley, 1933</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF8DAFA4DF89C" blockId="23.[151,1437,1075,2036]" box="[151,1436,1838,1860]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF8DAFA4DF89C" box="[151,1436,1838,1860]" pageId="23" pageNumber="474">- Scutellum very small, almost as wide as long............................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E8313FF2BF8BFFA4DF866" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF8BFFA4DF8A1" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF8BFFA4DF8A1" pageId="23" pageNumber="474">
13(12). Antennal tubercles very close to each other basally, with the area between them V-shaped; prothorax longer than wide.
<collectingCountry id="0EA6761B9B1E8313FA3BF8BFFE9EF8A1" name="El Salvador" pageId="23" pageNumber="474">El Salvador</collectingCountry>
.........................................................
<taxonomicName id="B1B14D089B1E8313FBBCF89DFA4DF8A1" authority="Franz, 1954" authorityName="Franz" authorityYear="1954" box="[1024,1436,1896,1918]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="subSpecies" species="salvadorensis" subSpecies="salvadorensis">
<emphasis id="44C5EA999B1E8313FBBCF89DFAF2F8A2" box="[1024,1315,1896,1918]" italics="true" pageId="23" pageNumber="474">P. salvadorensis salvadorensis</emphasis>
Franz, 1954
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B1E8313FF2BF872FA4DF866" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF872FA4DF866" pageId="23" pageNumber="474">
- Antennal tubercles not very close to each other basally, with the area between them U-shaped; prothorax as wide as long.
<collectingCountry id="0EA6761B9B1E8313FA3BF872FE80F866" name="El Salvador" pageId="23" pageNumber="474">El Salvador</collectingCountry>
,
<collectingCountry id="0EA6761B9B1E8313FEE6F857FE66F866" box="[346,439,1955,1977]" name="Honduras" pageId="23" pageNumber="474">Honduras</collectingCountry>
..............................................
<taxonomicName id="B1B14D089B1E8313FC52F850FA4DF866" authority="Franz, 1954" authorityName="Franz" authorityYear="1954" box="[1006,1436,1955,1977]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="subSpecies" species="salvadorensis" subSpecies="montecristensis">
<emphasis id="44C5EA999B1E8313FC52F850FAF2F866" box="[1006,1315,1955,1977]" italics="true" pageId="23" pageNumber="474">P. salvadorensis montecristensis</emphasis>
Franz, 1954
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B1E831CFF2BF834FA4DFF72" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
<paragraph id="760E368B9B1E8313FF2BF834FA4DF82C" blockId="23.[151,1437,1075,2036]" pageId="23" pageNumber="474">
<keyLead id="CD4095BE9B1E8313FF2BF834FA4DF82C" pageId="23" pageNumber="474">
14(6). Elytra with large tubercle laterally after middle, with its base wider than apical width of metafemora.
<collectingCountry id="0EA6761B9B1E8313FB74F834FAC0F809" box="[1224,1297,1984,2006]" name="Mexico" pageId="23" pageNumber="474">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B1E8313FAA2F834FABFF809" box="[1310,1390,1984,2006]" country="Mexico" name="Chiapas" pageId="23" pageNumber="474">Chiapas</collectingRegion>
)............................................................
<taxonomicName id="B1B14D089B1E8313FC0EF82AFA4DF82C" authority="Gutierrez, Toledo &amp; Noguera, 2020" authorityName="Gutierrez, Toledo &amp; Noguera" authorityYear="2020" box="[946,1436,2014,2036]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="44C5EA999B1E8313FC0EF82AFB91F82C" box="[946,1088,2014,2035]" italics="true" pageId="23" pageNumber="474">P. tuberculatus</emphasis>
Gutiérrez, Toledo &amp; Noguera, 2020
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFF6CFA4DFF72" blockId="24.[151,1436,151,555]" box="[151,1436,151,173]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFF6CFA4DFF72" box="[151,1436,151,173]" pageId="24" pageNumber="475">- None of the elytral tubercles as wide at the base as the apex of the metafemora................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFF41FA4DFF38" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFF41FA4DFF14" blockId="24.[151,1436,151,555]" box="[151,1436,181,203]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFF41FA4DFF14" box="[151,1436,181,203]" pageId="24" pageNumber="475">15(14). Apex of elytral tubercles glabrous..................................................................... 16</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFF27FA4DFF38" blockId="24.[151,1436,151,555]" box="[151,1436,210,232]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFF27FA4DFF38" box="[151,1436,210,232]" pageId="24" pageNumber="475">- Apex of elytral tubercles pubescent.................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFF1BFA4DFEFD" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFF1BFA4DFEDA" blockId="24.[151,1436,151,555]" box="[151,1436,239,261]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFF1BFA4DFEDA" box="[151,1436,239,261]" pageId="24" pageNumber="475">
16(15). Central tubercle of pronotum distinctly conical.
<collectingCountry id="0EA6761B9B11831CFD74FF1BFCC0FEDA" box="[712,785,239,261]" name="Mexico" pageId="24" pageNumber="475">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B11831CFCA2FF1BFCBFFEDA" box="[798,878,239,261]" country="Mexico" name="Chiapas" pageId="24" pageNumber="475">Chiapas</collectingRegion>
).........
<taxonomicName id="B1B14D089B11831CFC4CFF04FA4DFEDA" authority="Gutierrez, Toledo &amp; Noguera, 2020" authorityName="Gutierrez, Toledo &amp; Noguera" authorityYear="2020" box="[1008,1436,239,261]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="jonesi">
<emphasis id="44C5EA999B11831CFC4CFF04FB91FEDA" box="[1008,1088,240,261]" italics="true" pageId="24" pageNumber="475">P. jonesi</emphasis>
Gutiérrez, Toledo &amp; Noguera, 2020
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFEF9FA4DFEFD" blockId="24.[151,1436,151,555]" box="[151,1436,269,291]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFEF9FA4DFEFD" box="[151,1436,269,291]" pageId="24" pageNumber="475">
- Central tubercle of pronotum uniformly convex.
<collectingCountry id="0EA6761B9B11831CFD72FEF9FCE9FEFC" box="[718,824,269,291]" name="Guatemala" pageId="24" pageNumber="475">Guatemala</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFCFEFEF9FC4EFEFC" box="[834,927,269,291]" name="Honduras" pageId="24" pageNumber="475">Honduras</collectingCountry>
.
<taxonomicName id="B1B14D089B11831CFC12FEF9FA4DFEFD" authority="Santos-Silva, Van Roie &amp; Jocque, 2021" authorityName="Santos-Silva, Van Roie &amp; Jocque" authorityYear="2021" box="[942,1436,269,291]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="guifarroi">
<emphasis id="44C5EA999B11831CFC12FEF9FBCDFEFD" box="[942,1052,269,290]" italics="true" pageId="24" pageNumber="475">P. guifarroi</emphasis>
<bibRefCitation id="12204B7A9B11831CFB9DFEF9FA4DFEFD" author="Santos-Silva, A. &amp; Van Roie, M. &amp; Jocque, M." box="[1057,1436,269,291]" pageId="24" pageNumber="475" pagination="37 - 61" refId="ref17489" refString="Santos-Silva, A., Van Roie, M. &amp; Jocque, M. (2021) Longhorned woodboring beetles (Coleoptera, Cerambycidae) from Cusuco National Park, Honduras: new species, new records, and revalidation. European Journal of Taxonomy, 764, 37 - 61. https: // doi. org / 10.5852 / ejt. 2021.764.1469" type="journal article" year="2021">Santos-Silva, Van Roie &amp; Jocqué, 2021</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFEDEFA4DFE82" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFEDEFA4DFEE0" blockId="24.[151,1436,151,555]" box="[151,1436,298,320]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFEDEFA4DFEE0" box="[151,1436,298,320]" pageId="24" pageNumber="475">17(15). Antennal tubercles contiguous basally, with the area between them V-shaped................................... 18</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFEBCFA4DFE82" blockId="24.[151,1436,151,555]" box="[151,1436,327,349]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFEBCFA4DFE82" box="[151,1436,327,349]" pageId="24" pageNumber="475">- Antennal tubercles not contiguous basally, with the area between them U-shaped................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFE91FA4DFE48" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFE91FA4DFEA5" blockId="24.[151,1436,151,555]" box="[151,1436,357,379]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE91FA4DFEA5" box="[151,1436,357,379]" pageId="24" pageNumber="475">
18(17). Antennomere III as long as scape.
<collectingCountry id="0EA6761B9B11831CFDFAFE91FD5EFEA4" box="[582,655,357,379]" name="Mexico" pageId="24" pageNumber="475">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B11831CFD20FE91FD3DFEA4" box="[668,748,357,379]" country="Mexico" name="Chiapas" pageId="24" pageNumber="475">Chiapas</collectingRegion>
)..................
<taxonomicName id="B1B14D089B11831CFC69FE91FA4DFEA5" authority="Gutierrez, Toledo &amp; Noguera, 2020" authorityName="Gutierrez, Toledo &amp; Noguera" authorityYear="2020" box="[981,1436,357,379]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="cristinae">
<emphasis id="44C5EA999B11831CFC69FE91FB91FEA5" box="[981,1088,357,378]" italics="true" pageId="24" pageNumber="475">P. cristinae</emphasis>
Gutiérrez, Toledo &amp; Noguera, 2020
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFE77FA4DFE48" blockId="24.[151,1436,151,555]" box="[151,1436,386,408]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE77FA4DFE48" box="[151,1436,386,408]" pageId="24" pageNumber="475">
- Antennomere III longer than scape.
<collectingCountry id="0EA6761B9B11831CFDE8FE76FD6DFE47" box="[596,700,386,408]" name="Guatemala" pageId="24" pageNumber="475">Guatemala</collectingCountry>
......................................
<taxonomicName id="B1B14D089B11831CFB3FFE77FA4DFE48" authority="Breuning, 1940" authorityName="Breuning" authorityYear="1940" box="[1155,1436,386,408]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="echinoides">
<emphasis id="44C5EA999B11831CFB3FFE77FAD0FE48" box="[1155,1281,386,408]" italics="true" pageId="24" pageNumber="475">P. echinoides</emphasis>
Breuning, 1940
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFE54FA4DFE0D" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFE54FA4DFE6A" blockId="24.[151,1436,151,555]" box="[151,1436,415,437]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE54FA4DFE6A" box="[151,1436,415,437]" pageId="24" pageNumber="475">
19(17). Pronotum without strongly elevated central tubercle.
<collectingCountry id="0EA6761B9B11831CFD45FE6BFCB2FE6A" box="[761,867,415,437]" name="Guatemala" pageId="24" pageNumber="475">Guatemala</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFCD1FE6BFC1CFE6A" box="[877,973,415,437]" name="Honduras" pageId="24" pageNumber="475">Honduras</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFC6AFE54FB90FE6A" box="[982,1089,415,437]" name="Costa Rica" pageId="24" pageNumber="475">Costa Rica</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFBF7FE54FB46FE6A" box="[1099,1175,416,437]" name="Panama" pageId="24" pageNumber="475">Panama</collectingCountry>
...
<taxonomicName id="B1B14D089B11831CFB78FE54FA4DFE6A" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[1220,1436,415,437]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B11831CFB78FE54FAF4FE6B" box="[1220,1317,415,437]" italics="true" pageId="24" pageNumber="475">P. echinus</emphasis>
<bibRefCitation id="12204B7A9B11831CFA97FE54FA4DFE6A" author="Bates, H. W." box="[1323,1436,416,437]" pageId="24" pageNumber="475" refId="ref16372" refString="Bates, H. W. (1880) Longicornia, Fam. Cerambycidae [continuation]. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 17 - 152." type="book" year="1880">Bates, 1880</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFE49FA4DFE0D" blockId="24.[151,1436,151,555]" box="[151,1436,445,467]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE49FA4DFE0D" box="[151,1436,445,467]" pageId="24" pageNumber="475">- Pronotum with strongly elevated central tubercle with convex apex........................................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="CD452E2E9B11831CFF2BFE2EFA4DFDF5" pageId="24" pageNumber="475">
<paragraph id="760E368B9B11831CFF2BFE2EFA4DFE30" blockId="24.[151,1436,151,555]" box="[151,1436,474,496]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE2EFA4DFE30" box="[151,1436,474,496]" pageId="24" pageNumber="475">
20(19). Width of lower eye lobes equal to basal width of scape.
<collectingCountry id="0EA6761B9B11831CFCAFFE2EFC8EFE2F" box="[787,863,474,496]" name="Mexico" pageId="24" pageNumber="475">Mexico</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFCD5FE2EFC02FE2F" box="[873,979,474,496]" name="Guatemala" pageId="24" pageNumber="475">Guatemala</collectingCountry>
,
<collectingCountry id="0EA6761B9B11831CFC61FE2EFBEBFE2F" box="[989,1082,474,496]" name="Honduras" pageId="24" pageNumber="475">Honduras</collectingCountry>
.........
<taxonomicName id="B1B14D089B11831CFB11FE2FFA4DFE30" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[1197,1436,474,496]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="44C5EA999B11831CFB11FE2FFAF4FE30" box="[1197,1317,474,496]" italics="true" pageId="24" pageNumber="475">P. singularis</emphasis>
<bibRefCitation id="12204B7A9B11831CFA97FE2FFA4DFE30" author="Bates, H. W." box="[1323,1436,474,496]" pageId="24" pageNumber="475" refId="ref16372" refString="Bates, H. W. (1880) Longicornia, Fam. Cerambycidae [continuation]. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 17 - 152." type="book" year="1880">Bates, 1880</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="760E368B9B11831CFF2BFE0CFA4DFDF5" blockId="24.[151,1436,151,555]" pageId="24" pageNumber="475">
<keyLead id="CD4095BE9B11831CFF2BFE0CFA4DFDF5" pageId="24" pageNumber="475">
- Width of lower eye lobes greater than basal width of scape.
<collectingCountry id="0EA6761B9B11831CFC85FE03FC53FDD2" box="[825,898,503,525]" name="Mexico" pageId="24" pageNumber="475">Mexico</collectingCountry>
(
<collectingRegion id="B475F8699B11831CFC33FE03FC0DFDD2" box="[911,988,503,525]" country="Mexico" name="Nayarit" pageId="24" pageNumber="475">Nayarit</collectingRegion>
),
<collectingCountry id="0EA6761B9B11831CFC50FE03FB98FDD2" box="[1004,1097,503,525]" name="Honduras" pageId="24" pageNumber="475">Honduras</collectingCountry>
...............................................................................
<taxonomicName id="B1B14D089B11831CFCD1FDE1FA4DFDF5" authority="Heffern, Nascimento &amp; Santos-Silva, 2018" authorityName="Heffern, Nascimento &amp; Santos-Silva" authorityYear="2018" box="[877,1436,533,555]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="nayaritensis">
<emphasis id="44C5EA999B11831CFCD1FDE1FC28FDF5" box="[877,1017,533,554]" italics="true" pageId="24" pageNumber="475">P. nayaritensis</emphasis>
<bibRefCitation id="12204B7A9B11831CFC43FDE1FA4DFDF5" author="Heffern, D. &amp; Nascimento, F. E. L. &amp; Santos-Silva, A." box="[1023,1436,533,555]" pageId="24" pageNumber="475" pagination="59 - 80" refId="ref16886" refString="Heffern, D., Nascimento, F. E. L. &amp; Santos-Silva, A. (2018) Descriptions, redescription, notes, and new ranks in American Cerambycidae (Coleoptera). Zootaxa, 4531 (1), 59 - 80. https: // doi. org / 10.11646 / zootaxa. 4531.1.2" type="journal article" year="2018">Heffern, Nascimento &amp; Santos-Silva, 2018</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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